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1.
The famous Rhaetian bone bed (Late Triassic, 205 Ma) is well known because it marks a major switch in depositional environment from terrestrial red beds to fully marine conditions throughout the UK and much of Europe. The bone bed is generally cemented and less than 10 cm thick. However, we report here an unusual case from Saltford, near Bath, S.W. England where the bone bed is unconsolidated and up to nearly 1 m thick. The exposure of the basal beds of the Westbury Formation, Penarth Group includes a bone bed containing a diverse Rhaetian marine microvertebrate fauna dominated by sharks, actinopterygian fishes and reptiles. Despite the unusual sedimentary character of the bone bed, we find similar proportions of taxa as in other basal Rhaetian bone beds (55–59 % Lissodus teeth, 13–16 % Rhomphaiodon teeth, 12–14 % Severnichthys teeth, 6–9% Gyrolepis teeth, 3–4% undetermined sharks’ teeth, 1–3% undetermined bony fish teeth, and < 1% of each of Hybodus, Parascylloides, and Sargodon), the only differences being in the proportions of Rhomphaiodon teeth, which can represent 30–40 % of specimens elsewhere. This suggests that taphonomic bias of varying Rhaetian bone beds may be comparable despite different sedimentary settings, and that the proportions of taxa say something about their original proportions in the ecosystem.  相似文献   

2.
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3.
The Rhaetian (latest Triassic) succession of Doniford Bay, North Somerset has been noted as a site of fossils for over 200 years, and yet has never been described in detail despite its importance for palaeontology, for knowledge of a classic Triassic-to-Jurassic transition sequence, for structural geology, and as a venue for field trips. There are two bone beds, which differ substantially in sedimentary and palaeontological characteristics. Fossils include the usual teeth, denticles, and scales of small hybodont sharks, bony fishes, and marine reptiles. The lower (basal) bone bed is in many ways like those from other localities around Bristol and in South Wales, whereas the upper bone bed shows rich organic matter and an absence of calcite, suggesting a deeper location of deposition. Further, the lower bone bed contains abundant abraded silica grains, suggesting transport of sediment and bone debris from a beach or river. The two bone beds differ in faunal composition, and the upper bone bed lacks the locally derived clasts, larger silica grains, and calcite seen in the lower bone bed. Bones and teeth are equally abraded in both bone beds, confirming long-distance transport of fish and reptile fossils and that the upper bone bed cannot be interpreted as having derived from the lower.  相似文献   

4.
Westbury Garden Cliff has been a noted site for Rhaetian bone beds for over a century. It is known especially as a source of excellently preserved bones of the small marine reptile Pachystropheus as well as other reptiles, and fishes. Further it is the type locality of the Westbury Formation, the lower half of the British Rhaetian (Penarth Group). It was also featured in a debate over lateral equivalence of the basal Rhaetian bone bed, with supposedly 5–6 m of pre-basal bone bed deposition. However, the bone beds at different localities are unlikely to be of exactly the same age, and the succession at Westbury Garden Cliff lacks the erosive base of the Westbury Formation seen elsewhere and so presumably started later, perhaps reflecting progressive inundation of the Welsh High, the nearest land. However, the main bone bed occurs in lenses up to 20 cm wide, and may represent hummocky cross stratification, evidence of storm bed deposition. Trace fossils in several sandstones include Selenichnites and Crescentichnus, evidence of shallow-water limulids ploughing the sediment for food, Lockeia, the living burrows of bivalves, and Chondrites burrow systems, suggesting subsequent stability of the sandstone beds.  相似文献   

5.
The Late Triassic Rhaetian stage is perhaps best known in south-west Britain for the bone beds of the Westbury Formation, but there are other fossil-rich horizons within this and the underlying Blue Anchor Formation. Samples from a borehole drilled at the Filton West Chord, and collected from exposures near Bristol Parkway railway station, have yielded significant fossil material from both of these formations. The assemblage recovered from the Blue Anchor Formation is similar to those from the lower Westbury Formation, yielding roughly equal proportions of chondrichthyans and osteichthyans. Assemblages recovered from the Westbury Formation are typical of those from the upper Westbury Formation, in being dominated by osteichthyans. The borehole samples have produced the first recorded evidence of crinoids in the British Triassic, and the first evidence of coleoid cephalopods, in the form of grasping hooklets, from the Rhaetian, and indeed the first from the British Triassic.  相似文献   

6.
Since the discovery of the basal sauropodomorph dinosaur Thecodontosaurus in the 1830s, the associated fauna from the Triassic fissures at Durdham Down (Bristol, UK) has not been investigated, largely because the quarries are built over. Other fissure sites around the Bristol Channel show that dinosaurs represented a minor part of the fauna of the Late Triassic archipelago. Here we present data on microvertebrates from the original Durdham Down fissure rocks, which considerably expand the taxonomic diversity of the island fauna, revealing that it was dominated by the sphenodontian Diphydontosaurus, and that archosauromorphs, including sphenosuchian crocodylomorphs, coelophysoid theropods, and the basal sauropodomorph Thecodontosaurus, were diverse. Importantly, a few fish teeth provide new information about the debated age of the fissure deposit, which is identified as lower Rhaetian. Thecodontosaurus had been assigned an age range over 20–25 Myr of the Late Triassic, so this narrower age determination (209.5–204 Myr) is important for studies of early dinosaurian evolution.  相似文献   

7.
A new basal non-pterodactyloid pterosaur, Raeticodactylus filisurensis gen. et sp. nov., is reported. It has been discovered in shallow marine sediments from the Upper Triassic of the lowest Kössen beds (late Norian/early Rhaetian boundary) in the central Austroalpine of Canton Grisons (Switzerland). The disarticulated specimen is comprised of an almost complete skull and a partial postcranial skeleton. A high and thin bony, sagittal cranial crest characterizes the anterodorsal region of the skull. The large mandible, with an additional keel-like expansion at the front, partly matches the enlarged sagittal cranial crest. A direct and close relationship to Austriadactylus cristatus, the only known Triassic pterosaur with a bony cranial crest so far, cannot be established. The teeth of the premaxilla are monocuspid and exhibit very strongly bowed enamel wrinkles on the lingual side whereas the enamel is smooth on the labial side. These monocuspid teeth are large and fang-like. The numerous smaller teeth of the maxilla show three, four and five cusps. These are very similar to the teeth of the Triassic pterosaur Eudimorphodon ranzii. The humerus shows a thinner construction than that seen in other Triassic pterosaurs. The femur is quite unusual with a caput femoris perpendicular to the shaft. The bones of the extremities are almost twice as long as the ones from the largest Triassic specimen E. ranzii (MCSNB 2888). The newly described pterosaur is an adult, with a wingspan of approximately 135 cm. A morphofunctional analysis suggests that R. filisurensis was a highly specialized piscivore and possibly a skim-feeder.  相似文献   

8.
In the Late Triassic the landscape NE of present-day Bristol, SW England was dominated by Carboniferous Limestone ridges and cuestas that became progressively buried by continental Mercia Mudstones and finally inundated during the Rhaetian marine transgression. Mussini et al. (2020) adopt the assertions of earlier collaborators back to Whiteside and Marshall (2008) that terrestrial vertebrate assemblages from sediments contained within karstic fissure systems in the former limestone ridges at Cromhall, Tytherington and elsewhere are restricted to the Rhaetian. We review and reject the sedimentological, stratigraphic, geomorphological and topographic arguments for this and reassert a long pre-Rhaetian (Norian) history for the vertebrate-bearing fissure systems at both Tytherington and Cromhall. We also reject the contemporaneous Rhaetian freshwater-seawater mixing zone dissolution model for the fissure systems adopted by Mussini et al. (2020) and reaffirm that the Tytherington and Cromhall fissures developed as conduit caves with a long Triassic history. Applying a new regional study of the Rhaetian transgressive surface, we also show that whilst the fissures at Cromhall remained sealed after the Norian, those at nearby Tytherington were re-exposed in the Late Rhaetian. Already partially filled with Norian sediments, the Tytherington fissures were subject to reworking on the seabed. Internal collapses, probably triggered by well documented repeated regional seismicity, led to the chaotic state of the Tytherington fills when downward moving Rhaetian marine components came to lie amongst and mix with earlier Norian terrestrial sediments. The vertebrate associations in the Tytherington fissures therefore contain a substantial Rhaetian input whilst those at Cromhall do not.  相似文献   

9.
Foraminifers representing species Kaeveria fluegeli have been found in the Zorkaradjilga Formation (Sagenites quinquepunctatus Zone) of the upper Norian (or lower Rhaetian) in the central structural-facies zone of the South-East Pamirs. Their occurrence here is an additional criterion substantiating age and correlation of host deposits and an evidence in favor of fauna migration from the southern Tethys during the respective time span not only in northwestern areas of that ocean (Northern Calcareous Alps), but also in its central part (the South-East Pamirs).  相似文献   

10.
浙江长兴煤山地区晚二叠世末、早三叠世初的火山活动   总被引：14，自引：2，他引：14  
<正> 近年来,不少学者分别从生物地层(赵金科等,1981;盛金章等,1983;王义刚,1984;Sheng Jinzhang et al.,1984),事件地层(孙亦因等,1984;何锦文,1985),岩相、生物相(杨万容等,1981;芮琳等,1984),粘土矿物(何锦文,1981),元素地球化学特征(柴之芳等,1986),构造岩浆活动(周瑶琪,1986)以及碳同位素(陈锦石等,1984)等方面对浙江  相似文献   

11.
The Late Triassic fissure fills from the region of Bristol, SW England and S Wales, preserve unique assemblages of small vertebrates derived from an archipelago of palaeo-islands that document aspects of a critical transition in the history of terrestrial ecosystems. Tytherington Quarry, in south Gloucestershire, is the site of several fossiliferous fissures, all dated as Rhaetian (terminal Triassic), and source of abundant remains of the ‘Bristol dinosaur’, Thecodontosaurus antiquus. In addition, the fissure sediments have yielded previously unreported microvertebrate assemblages, including over 400 jaw remains from three genera of sphenodontians and 100 archosaur teeth assigned to 15 morphotypes. The land fauna is dominated by sphenodontians, with Diphydontosaurus by far the most common form, followed by Clevosaurus, then the sauropodomorph dinosaur Thecodontosaurus, and then the sphenodontian Planocephalosaurus. There are, in addition, rare remains of contemporaneous bony fishes, as well as fossils apparently reworked from the Carboniferous limestones, namely conodonts, holocephalian (chimaeroid) teeth, and a shark tooth. Many typical latest Triassic animals, such as temnospondyls, phytosaurs, aetosaurs, rauisuchians, plateosaurids and dicynodonts are not represented at Tytherington, perhaps because these generally larger animals did not live on the palaeo-island, or because their carcasses could not fit into the fissures. The absence of tritylodonts and early mammals is, however, less easy to explain on the basis of size, although it is known that these forms were abundant here by the Early Jurassic.  相似文献   

12.
In this paper the short snout Cenomanian enchodontids from the El Chango quarry, Chiapas, southeastern Mexico, are reviewed. Unicachichthys multidentata nov. gen. and sp. is named and identified as a new member of the Enchodontidae because it has the predorsal scute series that is the distinctive character of such family. This new genus resembles other short snout enchodontids, as Enchodus and Eurypholis; however, Unicachichthys differs from these and other members of the family because it shows characters ever observed, including a multitoothed dermopalatine, the presence of a basal sclerotic bone, and the serrations in the posterior edge of the preopercle vertical limb. The inclusion of Unicachichthys in two phylogenetic analyses, based on studies previously performed, suggests that this is a primitive representative of the family Enchodontidae. Additionally specimens of two different species of Enchodus from the El Chango quarry are also described; these specimens are so scarce and fragmentary that it is not possible to determine their specific taxonomic identity. The record of these Mexican fossils enrich the knowledge about the diversity of Cenomanian enchodontids in the North American domain of the Tethys Sea, which now contains new taxa that should be considered in future efforts to recognize the phylogenetic and biogeographic processes experienced by this fish group.  相似文献   

13.
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14.
The Early Cretaceous (latest Hauterivian) Faraoni Oceanic Anoxic Event (F-OAE) in the Río Argos section (Caravaca region, southern Spain), the candidate for the Global Boundary Stratotype Section and Point (GSSP) of the Hauterivian–Barremian transition, has been studied. Its ichnological bed-by-bed analysis allows for interpretation of oxygenation changes through the Faraoni Level interval and improvement upon previous characterization of oxygen conditions prior, during and after the F-OAE. The trace fossil assemblage belongs to the Zoophycos ichnofacies and it includes Chondrites intricatus, Chondrites targionii, Halimedides isp., Palaeophycus isp., ?Patagonichnus isp., Planolites isp., Rhizocorallium isp., Taenidium isp., Thalassinoides isp., Trichichnus linearis, Zavitokichnus fusiformis, and Zoophycos isp. Their diversity in particular beds fluctuates. Beds with four to six ichnotaxa reflect a multi-tiered macrobenthic trace maker assemblage living in good oxic and trophic conditions. In one bed below and one bed above the Faraoni Level (both marls without primary lamination), there are only two or three, mostly opportunistic ichnotaxa (Trichichnus, Chondrites, Planolites). They record dysoxic conditions. At the base of the Faraoni Level, one thicker and two thinner beds of marly mudstones (21.2 and 3.5 cm thick, respectively) are characterized by primary lamination. At the top and in the basal part of the thicker bed and in the thinner beds some trace fossils are present. These beds were deposited under anoxic conditions and later colonized by trace makers either from overlying beds deposited under oxic conditions or from the level of a greenish lamina in the lower part of the thicker bed, recording short episodes of dysoxic conditions. The thinner anoxic beds are separated by marls deposited under dysoxic conditions.  相似文献   

15.
《Gondwana Research》2006,9(4):579-584
A first report of discovery of spherules, glassy balls, highly magnetic fine dust and microbracciated matrix in the Fatehgarh Formation of Barmer Basin, Rajasthan, India is being presented in this paper. The Fatehgarh Formation is a mixed siliciclastic, carbonate and phosphorite formation of Cretaceous age in the Barmer Basin that comprises sediments of Middle Jurassic to Lower Eocene age. The phosphorite zone in the Fatehgarh Formation is ∼8 metre-thick zone that comprises phosphatic sandstone, bone bed, bedded phosphorite and phosphatic and non phosphatic gastropod beds. The spherules occur in a thin phosphatic-clay mud and silt band of bone bed, which also yielded a very rich and diverse microvertebrate assemblage with a dominant Late Cretaceous (Maastrichtian) form of Igdabatis along with forms comprising of Semionodontid, Lapisosteum and Enchodontid. The end Cretaceous is marked for a mass extinction of numerous species including dinosaurs. An extraterrestrial impact is interpreted as the reason for this mass extinction. Whether these spherules are related to the volcanic source or K/T Boundary impact ejecta found at Caribbean and Gulf of Mexico region needs detailed chemical and age characterization for which study is in progress.  相似文献   

16.
Alpine inversion in the Bristol Channel Basin includes reverse-reactivated normal faults with hanging wall buttress anticlines. At Lilstock Beach, joint sets in Lower Jurassic limestone beds cluster about the trend of the hinge of the Lilstock buttress anticline. In horizontal and gently north-dipping beds, J3 joints ( 295–285° strike) are rare, while other joint sets indicate an anticlockwise sequence of development. In the steeper south-dipping beds, J3 joints are the most frequent in the vicinity of the reverse-reactivated normal fault responsible for the anticline. The J3 joints strike parallel to the fold hinge, and their poles tilt to the south when bedding is restored to horizontal. This southward tilt aims at the direction of σ1 for Alpine inversion.Finite-element analysis is used to explain the southward tilt of J3 joints that propagate under a local σ3 in the direction of σ1 for Alpine inversion. Tilted principal stresses are characteristic of limestone–shale sequences that are sheared during parallel (flexural-flow) folding. Shear tractions on the dipping beds generate a tensile stress in the stiffer limestone beds even when remote principal stresses are compressive. This situation favors the paradoxical opening of joints in the direction of the regional maximum horizontal stress. We conclude that J3 joints propagated during the Alpine compression caused the growth of the Lilstock buttress anticline.  相似文献   

17.
New brachiopods (Dimerelloidea, Rhynchonellida) from Lower Jurassic (?lower Hettangian) hemipelagic sediments of the Swiss National Park in south-eastern Engadine are described: Sulcirostra doesseggeri sp. nov. and Carapezzia engadinensis sp. nov. Sulcirostra doesseggeri is externally similar to S. fuggeri (Frauscher 1883), a dubious species, that could not be included in a comparative study, because relevant samples no longer exist. A single specimen was tentatively assigned to Sulcirostra ?zitteli (Böse 1894) by comparison of its external morphology with S. zitteli from the type locality. The partly silicified brachiopods are associated with sponge spicules, radiolarians and crinoid ossicles. Macrofossils are rare: dictyid sponges, gastropods, bivalves, crustaceans, shark teeth and scales of an actinopterygian fish. The Lower Jurassic sediments (Alpisella beds, a basal member of the Allgäu Formation) preserving the brachiopods belong to the Ortler nappe (Upper Austroalpine nappes). The exact age of the Alpisella beds is not known, as index fossils are lacking. Their stratigraphic position above the Rhaetian Kössen Formation and below the ammonite-dated Trupchun beds suggests a very Early Jurassic, probably early Hettangian age for the new brachiopod fauna. The new species of Sulcirostra and Carapezzia are confined to a very small geographic area, a peculiarity also observed in other Early Jurassic dimerelloid brachiopods. These brachiopods presumably adapted to current-dominated submarine highs, where their shells could not accumulate, except when trapped in submarine cavities or re-deposited in submarine fans. Transport by turbidity currents is suggested for the Early Jurassic dimerelloids from the Engadine. Problems with the generic definition of Sulcirostra and the higher rank classification of Carapezzia are discussed.  相似文献   

18.
贵州剑河八郎乌溜—曾家崖寒武系第5阶底界潜在的全球层型剖面及点位上下的动物群(三叶虫和无铰纲腕足类)有显著的变化;研究的剖面厚约4.5m,穿过首现点位Oryctocephalus indicus。经逐层化石的采集和深入的研究,证实在寒武系第4阶或寒武系第2统顶部Ovatoryctocara granulata-Bathynotus holopygus带与寒武系第5阶或寒武系第3统底部Oryctocephalus indicus带之间大约有20cm的贫化石间隔区。这一动物群的突变是在均一的岩相中发生的,表明该剖面是建立寒武系第5阶(或第3统)底界的全球层型剖面和点位(GSSP)理想剖面。  相似文献   

19.
Distribution of faunal associations in the Silurian sequences of the Precordilleran and Northwestern Basins of Argentina is integrated with sedimentologic and taphonomic data, demonstrating that marked temporal and spatial changes in the Malvinokaffric brachiopod assemblages are related mainly to the type and intensity of the hydraulic regime of the shelf environment. The vertical arrangement of associations within each basin is controlled by local fluctuations in water depth, whereas the spatial variations are linked to minor environmental changes (substrate type and oxygen content). The basal strata of the Middle-Upper Silurian Los Espejos Formation of the Precordilleran Basin are mudstones of an open-shelf environment. These pass upward into facies of a storm-dominated shelf and shoreface. Four brachiopod assemblages have been recognized in the muddy facies: the Harringtonina Low-Diversity and Australina-Dominated associations characterize distal storm beds; the Harringtonina High-Diversity and Coelospira-Dominated communities occur in the fine-grained rocks of proximal tempestites. Storm shell beds, which dominate the middle-upper part of the sequence, are represented by allochthonous and parautochthonous coquinas. The Clarkeia Association flourished in shallow-water, high-energy, nearshore environments. Silurian rocks in the Northwestern Basin were deposited in an offshore, low-energy, restricted environment, and two main faunal assemblages have been recognized: a more distal Heterorthella Association and a relatively more proximal Ancillotoechia Association.  相似文献   

20.
We describe here new Late Triassic haramiyidan mammaliaform and reptile fossils from near the classic ‘Microlestes’ Quarry’ at Holwell, Somerset, U.K., where Charles Moore discovered a huge collection of microvertebrates in the 1850s. Moore’s discoveries included the haramiyid Thomasia (formerly ‘Microlestes’ and Microcleptes) for which he achieved worldwide fame. Subsequently, despite much fossicking by researchers at Holwell, few new identifiable specimens of mammaliamorphs and lepidosaurs have been recorded and these were by Kühne in 1939. The new finds described here from a bedded sequence, not from a fissure, add significantly to our knowledge of the Holwell tetrapods and to the Rhaetian terrestrial faunas of the SW U.K. Our discovery of haramiyidan teeth includes a previously unknown type of Theroteinus, a genus not previously recorded from outside of the St-Nicholas-de-Port locality in France. An archosaur tooth, possibly from a phytosaur, is also recorded. The new lepidosaur specimens add significant detail to the recently described ‘basal’ rhynchocephalian Penegephyrosaurus curtiscoppi as well as demonstrating that the global diversity of Lepidosauria in the Late Triassic remains incompletely known.  相似文献   

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