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1.
In October 2004, the North Pacific Marine Science Organization (PICES) sponsored a symposium to consider “Mechanisms that regulate North Pacific ecosystems: Bottom up, top down, or something else?” It sought to examine how marine populations, particularly the upper-trophic-level species, are regulated and to understand how energy flows through marine ecosystems. This introductory essay examines aspects of control mechanisms in pelagic marine ecosystems and some of the issues discussed during the symposium and in the 11 papers that were selected for this special issue. At global scales, the greatest biomass of fishes, seabirds and marine mammals tends to occur in regions of the world ocean with high primary production, e.g., the sub-arctic seas and up-welling regions of continental shelves. These large-scale animal distribution patterns are driven by food availability, not the absence of predators. At regional scales however, it is likely that current predation or past predation events have shaped local distributions, at least in marine birds and pinnipeds. Wasp-waist control occurs when one of the intermediate trophic levels is dominated by a single species, as occurs with small pelagic fishes of the world’s up-welling zones. Processes in these ecosystems may have features that result in a switch from bottom-up to top-down control.  相似文献   

2.
The “Wind Events and Shelf Transport” (WEST) program was an interdisciplinary study of coastal upwelling off northern California in 2000–03. WEST was comprised of modeling and field observations. The primary goal of WEST was to better describe and understand the competing influences of wind forcing on planktonic productivity in coastal waters. While increased upwelling-favorable winds lead to increased nutrient supply, they also result in reduced light exposure due to deeper surface mixed layers and increased advective loss of plankton from coastal waters. The key to understanding high levels of productivity, amidst these competing responses to wind forcing, is the temporal and spatial structure of upwelling. Temporal fluctuations and spatial patterns allow strong upwelling that favors nutrient delivery to be juxtaposed with less energetic conditions that favor stratification and plankton blooms. Observations of winds, ocean circulation, nutrients, phytoplankton and zooplankton off Bodega Bay and Point Reyes (38°N) were combined with model studies of winds, circulation and productivity. This overview of the WEST program provides an introduction to the WEST special issue of Deep-Sea Research, including the motivation for WEST, a summary of study components, an integrative synthesis of major research results to-date, and background on conditions during field studies in May–June 2001 (the upwelling period on which this special issue is focused).  相似文献   

3.
We introduce the Northern Oscillation Index (NOI), a new index of climate variability based on the difference in sea level pressure (SLP) anomalies at the North Pacific High (NPH) in the northeast Pacific (NEP) and near Darwin, Australia, in a climatologically low SLP region. These two locations are centers of action for the north Pacific Hadley–Walker atmospheric circulation. SLPs at these sites have a strong negative correlation that reflects their roles in this circulation. Global atmospheric circulation anomaly patterns indicate that the NEP is linked to the western tropical Pacific and southeast Asia via atmospheric wave trains associated with fluctuations in this circulation. Thus the NOI represents a wide range of tropical and extratropical climate events impacting the north Pacific on intraseasonal, interannual, and decadal scales. The NOI is roughly the north Pacific equivalent of the Southern Oscillation Index (SOI), but extends between the tropics and extratropics. Because the NOI is partially based in the NEP, it provides a more direct indication of the mechanisms by which global-scale climate events affect the north Pacific and North America.The NOI is dominated by interannual variations associated with El Niño and La Niña (EN/LN) events. Large positive (negative) index values are usually associated with LN (EN) and negative (positive) upper ocean temperature anomalies in the NEP, particularly along the North American west coast. The NOI and SOI are highly correlated, but are clearly different in several respects. EN/LN variations tend to be represented by larger swings in the NOI. Forty percent of the interannual moderate and strong interannual NOI events are seen by the SOI as events that are either weak or opposite in sign. The NOI appears to be a better index of environmental variability in the NEP than the SOI, and NPH SLP alone, suggesting the NOI is more effective at incorporating the influences of regional and remotely teleconnected climate processes.The NOI contains alternating decadal-scale periods dominated by positive and negative values, suggesting substantial climate shifts on a roughly 14-year ‘cycle’. The NOI was predominantly positive prior to 1965, during 1970–1976 and 1984–1991, and since 1998. Negative values predominated in 1965–1970, 1977–1983, and 1991–1998. In the NEP, interannual and decadal-scale negative NOI periods (e.g. EN events) are generally associated with weaker trade winds, weaker coastal upwelling-favorable winds, warmer upper ocean temperatures, lower Pacific Northwest salmon catch, higher Alaska salmon catch, and generally decreased macrozooplankton biomass off southern California. The opposite physical and biological patterns generally occur when the index is positive. Simultaneous correlations of the NOI with north Pacific upper ocean temperature anomalies are greatest during the boreal winter and spring. Lagged correlations of the winter and spring NOI with subsequent upper ocean temperatures are high for several seasons. The relationships between the NOI and atmospheric and physical and biological oceanic anomalies in the NEP indicate this index is a useful diagnostic of climate change in the NEP, and suggest mechanisms linking variations in the physical environment to marine resources on interannual to decadal climate scales. The NOI time series is available online at: http://www.pfeg.noaa.gov.  相似文献   

4.
The world’s largest mono-specific fishery, the Peruvian anchovy or anchoveta (Engraulis ringens) fishery, has been the subject of many studies since the 1960s. Details of its relationship with other species have mainly focused on alternations with sardine, Sardinops sagax, and little effort has so far been paid to interactions with other species sharing the same ecosystem. This is the case for Pleuroncodes monodon, the crustacean squat lobster or ’munida’, which has become highly abundant along the Peruvian coast since the mid-1990s. Munida is now an important prey for seabirds, mammals and coastal predatory fish. Knowledge of patterns of distribution and ecological niche of munida is scarce however off Peru. Here we describe and compare spatial patterns of distribution of anchoveta and munida and their ecological niches based on data from 26 acoustic surveys performed along the Peruvian coast between 1998 and 2006. The results indicate that munida and anchoveta share ecological niches but that munida is restricted to the coldest part of the productive cold coastal waters whereas anchoveta do not present any temperature preference over a large range (14–23 °C). The recent increase in munida abundance off Peru is concomitant with colder conditions; with their onset munida extended its range from central Chile northwards. Off Peru the very shallow oxycline keeps munida from its usual bottom habitat and has forced it to adopt pelagic behaviour.  相似文献   

5.
In the coastal waters off northern California, seasonal wind-driven upwelling supplies abundant nutrients to be processed by phytoplankton productivity. As part of the Coastal Ocean Processes: Wind Events and Shelf Transport (CoOP WEST) study, nutrients, CO2, size-fractionated chlorophyll, and phytoplankton community structure were measured in the upwelling region off Bodega Bay, CA, during May–June 2000, 2001 and 2002. The ability of this ecosystem to assimilate nitrate (NO3) and silicic acid/silicate (Si(OH)4) and accumulate particulate material (i.e. phytoplankton) was realized in all 3 years, following short events of upwelling-favorable winds, followed by periods of relaxed winds. This was observed as phytoplankton blooms, dominated by chlorophyll in cells greater than 5 μm in diameter, that reduced the ambient nutrients to zero. These communities were located over the near-shore shelf (<100 m depth) and were dominated by diatoms. An optimal window of 3–7 days of relaxed winds, following an upwelling pulse, was required for chlorophyll accumulation. The large-celled phytoplankton that result are likely important players in coastal new production and carbon cycling.  相似文献   

6.
Numerical simulations using a physiologically-based model of marine ecosystem size spectrum are conducted to study the influence of primary production and temperature on energy flux through marine ecosystems. In stable environmental conditions, the model converges toward a stationary linear log–log size-spectrum. In very productive ecosystems, the model predicts that small size classes are depleted by predation, leading to a curved size-spectrum.It is shown that the absolute level of primary production does not affect the slope of the stationary size-spectrum but has a nonlinear effect on its intercept and hence on the total biomass of consumer organisms (the carrying capacity). Three domains are distinguished: at low primary production, total biomass is independent from production changes because loss processes dominate dissipative processes (biological work); at high production, ecosystem biomass is proportional to primary production because dissipation dominates losses; an intermediate transition domain characterizes mid-production ecosystems. Our results enlighten the paradox of the very high ecosystem biomass/primary production ratios which are observed in poor oceanic regions. Thus, maximal dissipation (least action and low ecosystem biomass/primary production ratios) is reached at high primary production levels when the ecosystem is efficient in transferring energy from small sizes to large sizes. Conversely, least dissipation (most action and high ecosystem biomass/primary production ratios) characterizes the simulated ecosystem at low primary production levels when it is not efficient in dissipating energy.Increasing temperature causes enhanced predation mortality and decreases the intercept of the stationary size spectrum, i.e., the total ecosystem biomass. Total biomass varies as the inverse of the Arrhenius coefficient in the loss domain. This approximation is no longer true in the dissipation domain where nonlinear dissipation processes dominate over linear loss processes. Our results suggest that in a global warming context, at constant primary production, a 2–4 °C warming would lead to a 20–43% decrease of ecosystem biomass in oligotrophic regions and to a 15–32% decrease of biomass in eutrophic regions.Oscillations of primary production or temperature induce waves which propagate along the size-spectrum and which amplify until a “resonant range” which depends on the period of the environmental oscillations. Small organisms oscillate in phase with producers and are bottom-up controlled by primary production oscillations. In the “resonant range”, prey and predators oscillate out of phase with alternating periods of top-down and bottom-up controls. Large organisms are not influenced by bottom-up effects of high frequency phytoplankton variability or by oscillations of temperature.  相似文献   

7.
Many marine ecosystems exhibit a characteristic “wasp-waist” structure, where a single species, or at most several species, of small planktivorous fishes entirely dominate their trophic level. These species have complex life histories that result in radical variability that may propagate to both higher and lower trophic levels of the ecosystem. In addition, these populations have two key attributes: (1) they represent the lowest trophic level that is mobile, so they are capable of relocating their area of operation according to their own internal dynamics; (2) they may prey upon the early life stages of their predators, forming an unstable feedback loop in the trophic system that may, for example, precipitate abrupt regime shifts. Experience with the typical “boom-bust” dynamics of this type of population, and with populations that interact trophically with them, suggests a “predator pit” type of dynamics. This features a refuge from predation when abundance is very low, very destructive predation between an abundance level sufficient to attract interest from predators and an abundance level sufficient to satiate available predators, and, as abundance increases beyond this satiation point, decreasing specific predation mortality and population breakout. A simple formalism is developed to describe these dynamics. Examples of its application include (a) a hypothetical mechanism for progressive geographical habitat expansion at high biomass, (b) an explanation for the out-of-phase alternations of abundances of anchovies and sardines in many regional systems that appear to occur without substantial adverse interactions between the two species groups, and (c) an account of an interaction of environmental processes and fishery exploitation that caused a regime shift. The last is the example of the Baltic Sea, where the cod resource collapsed in concert with establishment of dominance of that ecosystem by the cod’s ‘wasp-waist” prey, herring and sprat.  相似文献   

8.
Mesozooplankton (>200 μm) grazing impact (% phytoplankton standing crop consumed d−1) was determined by the gut fluorescence method during three springs (2000, 2001 and 2002) and two winters (2002 and 2003) in a coastal upwelling region off northern California. Wind events, in terms of both magnitude and duration, varied inter-annually and seasonally and included both upwelling-favorable and relaxation events. Grazing impact of mesozooplankton also varied inter-annually and seasonally, and was highest during June 2000 (mean=129% of standing crop d−1), a prolonged period of wind “relaxation” and phytoplankton bloom. In contrast, mean grazing impact was lower during periods of stronger, more persistent winds, more active upwelling, greater cross-shelf transport, and lower chlorophyll concentration (25% and 38% in May–June 2001 and 2002, respectively). Wintertime conditions (January 2002 and 2003) were characterized by weakly upwelling or downwelling-favorable winds, low chlorophyll concentration, and lower mean mesozooplankton grazing impact (13% and 12%, respectively). The larger (>500 μm) size class contributed proportionally more to total mesozooplankton (>200 μm) grazing impact than the smaller (200–500 μm) size class during all sampling periods except spring 2002. These results suggest that mesozooplankton grazing impact is higher in spring than in winter, and that during the spring upwelling season, grazing is higher during periods of wind relaxation (weak upwelling) than during periods of stronger upwelling. Further, these results suggest an important role of mesozooplankton grazers on phytoplankton dynamics in the upwelling region off northern California.  相似文献   

9.
Community-based fisheries management is being widely promoted as an alternative to centralized systems based on the familiar bioeconomic models that have manifestly failed to prevent a near catastrophic overexploitation of fish stocks worldwide. The Pacific Island Region probably contains the world’s greatest concentration of still-functioning traditional community-based systems for managing coastal-marine fisheries and other resources.It has been frequently asserted that many such traditional systems provide both a firm foundation for future coastal fisheries management in the Pacific Islands Region, as well as a conceptual framework for managing fisheries elsewhere. Although now seemingly self-evident to fisheries development “experts”, such assertions remain largely unverified.Whereas it is a relatively straightforward task to distil basic “design principles” from a sample of systems, it is far more complex to analyze the multi-sectoral national environment in which they function, especially when their history is taken into account. In other words, it is far less widely appreciated that many contemporary community-based fisheries management systems are the end products of a long process of change and adaptation to external pressures and constraints.In this article I address some of the broader contextual issues that should be appreciated in policymaking with respect to a potential modern role for traditional management systems in general, and in the analysis of a future role for any given system. First, the principal external factors that have caused change in systems are described and exemplified. The recognition of the potential role of existing community-based fisheries systems, and attempts to act on it, is summarized for some Pacific Island nations, with a focus on the complex problem of reconciling customary and statutory legal systems. In the final section I examine three principal national policy alternatives regarding the potential role of existing local fisheries management systems, together with three main criteria for determining whether or not a system can be adapted to fulfill modern requirements.  相似文献   

10.
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