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1.
The reproductive biology of the seabream Diplodus argenteus, a dominant coastal fishery species, was investigated over two consecutive seasons (2001–2002) at Cabo Frio, Brazil, a low-latitude upwelling system. The sex ratio was dominated by females (1.4:1.0) and the length-at-50% sexual maturity (females) was 203mm total length. Females were multiple spawners and the reproductive pattern appeared to be digynous protandrous hermaphroditism. Monthly variations in gonadosomatic index and proportion of ripe females indicated that reproductive activity occurred from late winter to summer (August–February), following coastal upwelling.  相似文献   

2.
The Cabo Frio region in the state of Rio de Janeiro, southeast coast of Brazil, is characterized by a local coastal upwelling system and converging littoral sediment transport systems that are deflected offshore at Cabo Frio, as a consequence of which a thick cross-shelf sediment deposit has developed over time. To investigate the evolution of this muddy deposit, geophysical, sedimentological and geochemical data from four sediment cores (3.8–4.1 m in length) recovered in water depths between 88 and 141 m were analyzed. The high-resolution seismic data show variable sediment thicknesses ranging from 1 to 20 m, comprising two sedimentary units separated by a high-impedance layer at a depth of about 10 m below the seafloor at the coring sites. According to the available age datings, the upper sedimentary unit is late Pleistocene to Holocene in age, whereas the lower unit (not dated) must, by implication, be entirely Pleistocene in age. The boomer-seismic reflection signal can be divided into three echo-types, namely transparent (inner shelf), stratified (middle shelf) and reflective (outer shelf), each type seemingly related to the local sediment composition. The upper 4 m of the upper sedimentary unit is dominated by silty sediment on the middle shelf, and by upward-fining sediments (silty sand to sandy silt) on the inner and outer shelf. The downcore trends of P-wave velocity, gamma-ray density and acoustic impedance are largely similar, but generally reversed to those of water and organic carbon contents. Total organic carbon contents increase with decreasing mean grain size, periodic fluctuations suggesting temporal changes in the regional hydrodynamics and primary productivity fuelled by the local upwelling system. The reconstruction of sedimentation rates in the course of the Holocene is based on 35 AMS age datings of organic material recovered from variable downcore depths. These range from a maximum of 13.3 cm/decade near the base of the inner shelf core (7.73–7.70 ka BP) to generally very low values (<0.11 cm/century) over the last thousand years in all cores. Over the last 6 ka there appear to have been three distinct sedimentation peaks, one between 6 and 5 ka BP, another between 4 and 3 ka PB, and one around 1 ka BP. Due to different time intervals between dates, not every peak is equally well resolved in all four cores. Based on the similar sedimentology of the inner and outer shelf cores, an essentially identical sedimentation model is proposed to have been active in both cases, albeit at different times. Thus, already during the last glacial maximum, alongshore sediment transport was deflected offshore by a change in shoreline orientation caused by the Cabo Frio structural high. The source of terrigenous material was probably a barrier-island complex that was subsequently displaced landward in the course of sea-level rise until it stabilized some 6.5 ka BP along the modern coast.  相似文献   

3.
The mackerel scad Decapterus macarellus is of great social and economic importance to Cabo Verde citizens; however, in recent years catches have shown an unexplained decline. The harvest is regulated by way of a minimum legal size and a seasonal closure, with both measures based on the results of reproductive studies that did not consider the microscopic characteristics of the gonads. This study aimed to analyse the gonads microscopically in order to classify the sexual development stage of individuals and to estimate the size at first maturity and the reproductive season. This information was then used to analyse the match between the reproductive parameters and the management measures. The samples comprised 284 females (20–36 cm fork length [FL]) and 85 males (21–35 cm FL). The annual variation in gonadosomatic index and the monthly frequencies of mature phases suggest that the reproductive season occurs from March to October, with two spawning peaks, one in March and another from July to October. The current seasonal closure (from July to September) covers the main reproductive period. However, the species has a long spawning period and the timing of peaks can vary between years; hence, a reassessment of this management measure is recommended. The sizes at first maturity were estimated at 24.1 cm FL for females, and 26.6 cm FL for males. These values are higher than the minimum legal size (20 cm FL) currently established for management of the species at Cabo Verde; thus, we strongly recommend an increase in the minimum legal size in an attempt to reduce catches of immature individuals.  相似文献   

4.
Size at age, growth rates, seasonality of spawning, and size at maturity were described for the silver sweep Scorpis lineolatus off the coast of New South Wales, Australia. Estimates of age were made by counting annual zones in sections of otoliths. The ageing technique was validated using young‐of‐the‐year fish, staining fish with tetracy‐cline, and by marginal increment analysis. Silver sweep exhibited extreme longevity with an observed maximum age of 54 years and more than 50% of the fishery being greater than 15 years old. Growth was rapid during the first few years before reaching sexual maturity at 2–3 years and at a length of c. 17 cm fork length, after which growth slowed dramatically. Silver sweep displayed a winter spawning period. The fishery is unusual in that despite heavy fishing pressure and rapidly declining catches the age structure of the catch suggests a population that has been subjected to minimal fishing mortality. It is proposed that the large declines in commercial landings may be the result of serial depletion on local reefs and that current landings are being taken from schools of fish that have" receiveddate="little fishing pressure historically.  相似文献   

5.
One specimen ofPteraclis aesticola was collected off San José del Cabo, Baja California Sur, México (22°54′N, 109°45′W), in March 2007. Present record is the first reported occurrence of the species in the Tropical Eastern Pacific biogeographic region (Gulf of California to southern Ecuadorian waters). Its large fan-like anal and dorsal fins and its counts of fin rays and vertebrae can distinguish the Pacific fanfish from the other species in the genus. This fish may have not yet been recorded in the region because its presence has been overlooked in the past because of rarity and lack of commercial value.  相似文献   

6.
This study evaluated the effect of environmental stimuli and selective pressures in different geographical areas along a latitudinal gradient, on the juvenile recruitment, population structure, and sex ratio of the speckled swimming crab Arenaeus cribrarius. Samples were collected monthly during 1 year in three locations along the Brazilian coast: Macaé, state of Rio de Janeiro (MAC, 22°47′ S, 41°45′ W); Ubatuba, São Paulo (UBA, 23°27′ S, 44°58′ W); and São Francisco do Sul, Santa Catarina (SFS, 26°08′ S, 48°34′ W). The specimens of A. cribrarius were identified, counted, sexed, and measured for maximum carapace width (CW). The largest juvenile found was in UBA (47.7 ± 1.36 mm); and the largest adult females and males in MAC (74.26 ± 0.93 and 77.04 ± 0.79 mm, respectively). Recruitment in MAC was continuous, whereas in UBA and SFS, recruitment showed seasonal characteristics. The sex ratio was skewed toward females only in UBA; in MAC and SFS, males and females were present in equal proportions. These results indicate that geographical variations can cause differences in the recruitment and population structure of A. cribrarius. These regional differences call attention to the necessity for improved management plans and control of shrimp fishing, which can affect population patterns such as juvenile recruitment, population structure and life history of the target species and species that are caught in bycatch from shrimping, such as the swimming crab A. cribrarius.  相似文献   

7.
This study aimed to compare the spatio-temporal distribution of demographic groups of Hepatus pudibundus in the Ubatuba Bay, São Paulo, Brazil, over a 20-year interval. Two 1-year samplings were conducted: the 1st period (P1) ran from September 1995 to August 1996 and the 2nd period (P2) ran from September 2016 to August 2017. Samples were taken from five stations that differed in depth (5, 10, and 15 m) and location (exposed area and sheltered area), aboard a shrimp fishing boat. Over the 20-year period, the percentage of fine sediment increased in all stations. In total, we captured 865 H. pudibundus individuals in the P1 and 4,222 in the P2. The higher abundance seen in the P2 may have been caused by the effects of an intense ENSO event in Ubatuba, as well as co-occurring protective actions against fisheries activities in the region. The spatial distribution also changed after 20 years: In the P1, H. pudibundus were more concentrated in the 15 m station and exposed area, but they shifted to the sheltered area in the P2. This change may be related to the increase of fine sediments in this area, and to the fact that it is naturally protected from fisheries. According to the correspondence analysis, the abundance of males and juveniles was correlated with spring and winter, respectively. In the P1, females were more abundant in spring, whereas in the P2 they were more abundant in summer, which may be explained by a higher nutrient availability to the larvae in the summer of the P2. Our comparisons over the 20-year interval revealed that the spatio-temporal distribution of H. pudibundus has changed and that the population is well established in Ubatuba Bay.  相似文献   

8.
The deep-water giant red shrimp, Aristaeomorpha foliacea (Risso, 1827) (Decapoda, Aristeidae), represents a highly valuable resource for bottom trawling in the Mediterranean Sea. Recent assessments have described both a worsening of the status of the traditionally exploited stocks and low levels of annual-based instantaneous natural mortality (M 0.4–0.7 year−1) in the unexploited stocks. The mortality (M) figures, however, are in contrast with the longevity (Tmax) of 3–4-year males and 4–5-year females, estimated from classical length frequency distribution analysis. Reduced growth (after the onset of sexual maturity) and pile-up of older individuals in the larger size classes have been considered to be reasonable explanations for the contrast between M and Tmax. We propose that a discontinuity of the growth models could address this contrast. Because a clear discontinuity in sexual maturity is evident only in males, length frequency distribution data for different sets of males collected from the South of Sicily and the Maltese Islands deep bottoms (400–800 m) were fitted with both the classic (c) and double-phased (d) von Bertalanffy growth formula (VBGF). According to the dVBGF, adult males sharply reduce their growth rate after reaching an age between 1.2 and 1.5 year, which corresponds to the estimates of age at sexual maturity (tm between 1 and 1.5 years). The reduction in growth determines a higher Tmax (7.3–9.5 year) and lower M (0.4–0.6 year−1) than previously derived on the basis of cVBGF estimates (Tmax 3–4 years and M 1.2 year−1). The dVBGF results suggest that sexual dimorphism and length–sex segregation in giant red shrimps reflect an alternative life history strategy that is based mainly on growth reduction in adult males and it might be adopted for implementing more conservative assessments.  相似文献   

9.
Frontal upwelling is an important phenomenon in summer in the Yellow Sea (YS) and plays an essential role in the distribution of nutrients and biological species. In this paper, a three-dimensional hydrodynamic model is applied to investigate the characteristics and influencing factors of frontal upwelling in the YS. The results show that the strength and distribution of frontal upwelling are largely dependent on the topography and bottom temperature fronts. The frontal upwelling in the YS is stronger and narrower near the eastern coast than near the western coast due to the steeper shelf slope. Moreover, external forcings, such as the meridional wind speed and air temperature in summer and the air temperature in the preceding winter and spring, have certain influences on the strength of frontal upwelling. An increase in air temperature in the previous winter and spring weakens the frontal upwelling in summer; in contrast, an increase in air temperature in summer strengthens the frontal upwelling. When the southerly wind in summer increases, the upwelling intensifies in the western YS and weakens in the eastern YS. The air temperature influences the strength of upwelling by changing the baroclinicity in the frontal region. Furthermore, the meridional wind speed in summer affects frontal upwelling via Ekman pumping.  相似文献   

10.
The sparid Dentex macrophthalmus is a widespread, important fishery species along most of the West African coast from southern Namibia to the Mediterranean. In southern Angola it is an important artisanal species targeted predominantly by handline fishers. A biological and fisheries study was conducted on this species in southern Angola between June 2008 and July 2009. It was the dominant species in the artisanal fishery, accounting for 99% of the sparids captured and 67% (by mass) of the total catch. The life history of D. macrophthalmus was characterised by slow growth (females: Lt = 309[1 ? e?0.06(t + 5.43)], males: Lt = 248[1 ? e?0.16(t ? 1.77)]), advanced age at maturity (females: 7.4 years, males: 6.0 years) and high longevity (females: 36 years, males: 38 years). The sex ratio was 1:1 male:female. The length- and age-frequency distributions and macroscopic observations suggested that the species is a late gonochorist. Males and females reached 50% maturity at 151 and 166 mm fork length respectively. Although individuals with ripe gonads were found during most of the year, the peak spawning period appeared to be in December and January. Despite a life history that renders D. macrophthal-mus vulnerable to overexploitation, only 38% of artisanal fishers noticed a decline in the catches of this species. Potential reasons for this include: technology creep; limited pressure on the juvenile portion of the stock as a result of late recruitment (above the length of 50% maturity); the ‘basin affect’, whereby depleted areas are reseeded by areas (e.g. deeper water) inaccessible to the linefishery; and a deep-water reserve of large individuals. It is recommended that precautionary management strategies be implemented until the age estimates are revised by the countries in which this species forms significant fisheries.  相似文献   

11.
We have studied the relation between the hydrography, the composition and horizontal structure of the larval fish community, and the horizontal distribution patterns of larval fish abundances in an area characterised by strong mesoscale oceanographic activity, located between the Canary Islands and the African coast (the Canaries Coastal Transition Zone), during August 1999. Upwelling, upwelling filaments, cyclonic and anticyclonic eddies and island wakes are typical mesoscale features of the northwest African coast in summer. A single upwelling filament off Cabo Juby was joined in mid-August by a second that originated about 100 km to the north. The two filaments flowed together and merged 100 km offshore. The merged filament was partially entrained around a cyclonic eddy, trapped between the Canary Islands and the African coast, and interacted with cyclonic and anticyclonic eddies shed from Gran Canaria. Mesoscale oceanographic features strongly influenced the horizontal distributions of fish larvae. Eddies acted as a mechanism of concentration, while upwelling filaments were dispersive, transporting larvae from the African neritic zone into oceanic areas and towards the Canary archipelago. This transport was the major cause of the predominance of neritic larvae in the composition of the larval fish community of the area. The results also suggest: (1) that anchovy larvae are good indicators of the offshore displacement of upwelled water; (2) that the alternation between anchovy and sardine as species dominant in the larval fish community of the area during summer depends upon the water temperature in the African upwelling region, anchovy dominating at higher temperature; (3) that a coupling of anchovy and sardine spawning with the mesoscale oceanographic structure formed by the upwelling filaments and trapped eddy overcomes the negative effect that Ekman transport has on their populations.  相似文献   

12.
Previously published estimates of the area of the Peru upwelling ecosystem vary by more than an order of magnitude. In an effort to improve this situation, we used a 24-month sequence of SeaWiFS satellite images of chlorophyll in the surface water off Peru from 5°S to 18.5°S during September 1997–August 1999 to estimate the size of the nutrient enhanced productive habitat associated with the upwelling. The first 12-month period was marked by El Niño conditions, the second by strong upwelling. Using a chlorophyll threshold of >1.0 mg m−3 to define the limit of the productive habitat resulted in maximum area estimates of 120×103 km2 during September 1997–August 1998, and 220×103 km2 during September 1998–August 1999. The latter result is consistent with an area estimate we calculated using total fishery landings and a regression relating fishery yields per unit area to annual primary production per unit area. Although year-to-year variation in the annual mean size of the upwelling ecosystem must be significant, even discounting El Niño events, our analysis has shown that at least five of the extreme earlier values are not good estimates of the size of the productive habitat. We may now be close to knowing the average size of the ecosystem to within a factor of about two.  相似文献   

13.
The consequences of two upwelling events in mid- (MW) and late (LW) winter on biogeochemical and phytoplankton patterns were studied in the Pontevedra Ria and compared with the patterns measured under typical winter conditions and under a summer upwelling event. Thermohaline patterns measured during the mid-winter upwelling event (MW-up) revealed the intrusion of saltier seawater (35.9) into the ria associated with the Iberian Poleward Current (IPC). During the late-winter upwelling event (LW-up), the seawater which had welled up into the ria showed characteristics of the Eastern North Atlantic Central Water mass (ENACW). In both cases the measured water residence time (4 days during MW-up and 10 days during LW-up) was related to both meteorological and fluvial forcing. This residence time contrasts with that of summer upwelling (7 days) and with that estimated under unfavorable upwelling atmospheric conditions (2–4 weeks). During MW-up, the ria became poor in nutrients due to continental freshwater dilution, associated with the shorter residence time of the water, and the intrusion of IPC, which is a water body poor in nutrient salts: 2.9 μM of nitrate, 0.1 μM of phosphate and 1.5 μM of silicate. During this event, the ria exported 3.4 molDIN s−1, compared with 6.9 molDIN s−1 in non-upwelling conditions. Phytoplankton showed a uniform distribution throughout the ria, as during unfavorable upwelling conditions, and was characterized by the dominance of diatoms, mainly Nitzschia longissima and Skeletonema costatum. During LW-up, a nutrient depletion in the photic layer also occurred, but as a result of a phytoplankton spring bloom developing at this time. The ria was a nutrient trap where 4.1 molDIN s−1 were processed by photosynthesis. This budget is three times higher than the one under non-upwelling conditions. In contrast with the MW-up, which had no effect on primary production, during LW-up the ria became more productive, although not as productive as during a summer upwelling event (9.9 molDIN s−1). The taxonomic composition of the phytoplankton community did not change noticeably during LW-up and the summer upwelling, with the same species present and changing only in relative proportions. Diatoms were always the dominant microphytoplankton community, with Pseudonitzschia pungens, Thalassionema nitzschioides and several species of Chaetoceros as characteristic taxons.  相似文献   

14.
Main features of the zooplankton distribution and the ecological characteristics of the dominant species in the northern Benguela during different phases of upwelling are discussed. The composition of the zooplankton between 17 and 27°S was similar each year. Among the 20°30 most abundant species, 3°4 copepods dominated, influencing the distribution of total zooplankton biomass. During quiescent upwelling, zooplankton abundance was low and there were no significant differences in the inshore-offshore distribution of zooplankton biomass, the maximum occurring over the slope. During active upwelling, zooplankton biomass increased significantly, the maximum over the shelf being constituted almost entirely of developmental stages of herbivorous copepods. Over the inner shelf, all stages of the copepod Calanoides carinatus were feeding actively, removing up to 5 per cent per day of the standing stock of phytoplankton. Comparison of daily ration, respiration rate and biochemical composition of C. carinatus revealed active storage of energy inshore. Offshore populations of C. carinatus, found deeper than 200 m, comprised mainly copepodite stage V, which were not feeding and were characterized by decreased mobility and respiration and a high lipid content. It is estimated that the energy stored during active upwelling enables copepods to survive up to six months without any additional source of energy.  相似文献   

15.
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16.
The cold-water belt (CWB) is frequently formed off the Soya Warm Current (SWC) during summer and autumn. The detailed distribution of the flow and temperature fields observed by the R/V Sinyo-maru in the summer of 2001 captured the structures of the SWC and the CWB. The temperature and density distributions showed that the vertical distribution of the CWB is associated with the upwelling formed off the SWC. Numerical experiments using a two-layer model with realistic bottom topography have been performed to understand the formation mechanism of CWB and the upwelling structure off the current. In the experiment, the sea level difference between the Japan Sea and the Okhotsk Sea, and baroclinic flow assuming the Tsushima Warm Current were given along the open boundary. The numerical model well reproduces the current system of the SWC and upwelling region off it. The upwelling region is formed at the Soya Strait first, and then it spreads on the offshore side along the SWC as a developing current system. Analysis of the model data indicated that the geostrophic balance mainly dominates in the current system, while convergence of the bottom Ekman transport due to the SWC forms the upwelling region as the secondary circulation. In addition, the advection effect due to the SWC is found to strengthen the upwelling.  相似文献   

17.
New in situ time-series data were acquired by two ADCP moorings placed on the shelf off Richards Bay on the east coast of South Africa at depths of 25 m and 582 m between October 2009 and August 2010. The 11-month inshore bottom-temperature record revealed five substantial upwelling events lasting 5–10 days each where temperatures decreased by about 7 °C to 17–18 °C. Satellite sea surface temperature data showed these events to coincide with cold-water plumes occupying the northern wedge of the KwaZulu-Natal (KZN) Bight. Numerous shorter duration (1–2 days) upwelling events with less vivid surface expressions were also observed throughout the entire record where bottom temperature dropped by 2–3 °C. The last four months of the record were characterised by a protracted cool period lacking a seasonal trend but punctuated with oscillations of warm and cooler bottom water. In contrast to earlier studies that suggested upwelling was topographically and dynamically driven by the juxtaposition of the Cape St Lucia offset and the Agulhas Current (a solitary mechanism), our analysis showed almost all major and minor cold-water intrusions to coincide with upwelling-favourable north-easterly winds that simultaneously force a south-westerly coastal current. Ekman veering in the bottom boundary layer of the Agulhas Current, and the concomitant movement of cold water up the slope, was found to coexist at times with coastal upwelling, but its absence did not impede inshore cold-water intrusions, calling into question its role as a primary driver of upwelling. Both major and minor upwelling events were observed to promote phytoplankton blooms in the northern KZN Bight which commonly extended to the Thukela River. Wind-driven upwelling was also observed in the inner bight between Richards Bay and Port Durnford, explaining the ribbon of coastal chlorophyll continuously observed on ocean colour images between Cape St Lucia and the Thukela River. Similarities in upwelling character and mechanisms are observed between the northern KZN Bight and the Florida Current shelf systems.  相似文献   

18.
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19.
Elevated levels of phytoplankton were observed at the Northern California coastal upwelling ecosystem studied as part of the CoOP-WEST project during monthly summer surveys in 2000, 2001 and 2002. The high concentrations of chlorophyll were made up mostly of larger cells; the phytoplankton communities that resulted were dominated by centric diatoms. The highest chlorophyll a concentrations were observed when large diatoms or small colony-forming species dominated the assemblages. In contrast, when smaller nano-flagellates and picoplankton were dominant, total chlorophyll a concentrations were over four or five-fold lower than when diatoms were prevalent, illustrating the importance of large diatoms to total phytoplankton biomass. Each year, when chlorophyll concentrations were greatest, Chaetoceros species created a Chaetoceros-dominated system. A few other genera contributed to the upwelling diatom community, mostly the centric diatoms Asterionella, Thalassiosira, Rhizosolenia (including Guinardia and Proboscia), and the pennate Nitzschia. These diatoms have been described in other coastal upwelling ecosystems, and at this study site form a functional group that are apparently well adapted to the high-nutrient, turbulent conditions that are typical of these coastal regions.  相似文献   

20.
Observations are made of the maturation status of 81 male and 88 female sixgill sharks Hexanchus griseus from southern African waters. Males mature at about 310 cm total length (TL) with the calcification of the terminal cartilage elements of the claspers. Determination of maturity for females was problematic, but most were fully mature by at least 420 cm TL. Newborns were captured at the same location over three consecutive summer seasons, indicating the location of a possible pupping ground.  相似文献   

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