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1.
现代海底低温热液弥散流区微生物生态的发育情况已经成为当前热液系统研究关注的热点之一。大量的分析表明在低温热液弥散流区赋存着丰富的化能自养微生物,以硫、铁等元素的氧化还原反应获取新陈代谢能量,这些微生物的分布与低温热液流体的物理化学条件有着密切的联系。这些发现极大地丰富了我们对低温弥散流区微生物生态、关键地球化学过程与微生物新陈代谢耦合关系的认识。此外,低温热液流体是研究洋壳深部生物圈的窗口,通过这个窗口可以了解地壳内部生命的新陈代谢方式,进而理解地球内部微生物与洋壳内部流体、岩石之间的相互作用机制。  相似文献   

2.
The Saldanha hydrothermal field is hosted atop a mafic–ultramafic seamount, located at a non-transform offset on the Mid-Atlantic Ridge. Previous observations revealed a field where transparent low-temperature fluids discharge through centimeter-sized vents without the formation of chimney structures. We present geochemical and stable isotope (O and C) analyses from sediment samples collected at this field, both at and far from the vent area. Most sediments, including some directly adjacent to orifice vents, are pelagic oozes with only a weak hydrothermal overprinting. Hydrothermal precipitates are characterized by Fe–Mn oxyhydroxides and a minor amount of Cu–Zn sulphide minerals. However, one of the cores (SCD7) collected at the vent area shows a much stronger hydrothermal signature. This core is composed of a matrix of serpentine + talc ± chlorite with high porosity, where calcite + chalcopyrite + sphalerite/wurtzite ± pyrite–pyrrhotite were precipitated. In this core, metal enrichments, REE patterns, and the oxygen and carbon isotope composition of calcites indicate that mineralization must have occurred in the subsurface by high-temperature fluids, with minor mixing with seawater and with a significant magmatic contribution. Thus, while most samples confirm previous findings indicating that Saldanha hydrothermal fluid discharge is mainly diffuse and of low temperature, data from core SCD7 suggest that areas of high-temperature hydrothermal activity also occur, where temperatures of the fluids could reach > 260 °C and maximum temperatures of 330 °C. We suggest that fluids can flow through faults at the top of the mount and discharge in a more focused way through vent orifices, producing intense hydrothermal alteration of the sediments. At these locations complex hydrothermal processes occur, including reactions of the hydrothermal fluids with mafic and ultramafic rocks and magma degassing, as suggested by the carbon isotope composition of hydrothermal calcites. The high temperature of the fluid inferred from the geochemistry of the hydrothermal minerals requires a significant heat input to the system, suggesting an additional magmatic heat source to the already proposed exothermic serpentinization reactions.  相似文献   

3.
A low-temperature diffuse flow site associated with abundant vent fauna was found by submersible observations on the southern East Pacific Rise at 17°25′ S in 1997. This site was characterized by thin sediment covered pillow and sheet lavas with collapsed pits up to ∼15 m in diameter. There were three warm water vents (temperature: 6.5 to 10.5 °C) within the site above which the vented fluids rise as plumes. To estimate heat flux of the warm water vents, a temperature logger array was deployed and the vertical temperature distribution in the water column up to 38 m above the seafloor was monitored. A stationary deep seafloor observatory system was also deployed to monitor hydrothermal activity in this site. The temperature logger array measured temperature anomalies, while the plumes from the vents passed through the array. Because the temperature anomalies were measured in only specific current directions, we identified one of the vents as the source. Heat flux from the vent was estimated by applying a plume model in crossflow in a density-stratified environment. The average heat flux from September 13 to October 18, 1997 was 39 MW. This heat flux is as same order as those of high-temperature black smokers, indicating that a large volume flux was discharged from the vent (1.9 m3/s). Previous observations found many similar warm water flow vents along the spreading axis between 17°20′ S–30′ S. The total heat flux was estimated to be at least a few hundred mega-watts. This venting style would contribute to form effluent hydrothermal plumes extended above the spreading axis.  相似文献   

4.
The vestimentiferan tubeworm Ridgeia piscesae is an ecosystem-structuring organism in the hydrothermal vent environments of the Northeast Pacific. During this study, a single representative aggregation of the long-skinny morphotype of R. piscesae from the main endeavor segment was monitored for 3 yr before being collected in its entirety with a hydraulically actuated collection device manipulated in situ by a research vehicle. Vestimentiferan growth rates in this aggregation were determined by staining the exterior of the tubes and measuring newly deposited tube sections. The average growth rate of R. piscesae in this aggregation was very low in both years of the growth study (3.2 mm yr−1). Although the incidence of plume damage from partial predation was very high (>95%), mortality was very low (<4% yr−1). The distribution and the very tight clustering of recently recruited individuals indicated gregarious settlement behavior that is hypothesized to be partly due to biotic cues from settled larvae. Coupled measurements of vent fluid sulfide concentration and temperature were used to calculate the exposure of the vestimentiferans to sulfide from short- and long-term temperature monitoring. Plume-level temperature records indicate that most of the time individuals in this aggregation were exposed to extremely low levels of vent fluid, and therefore sulfide (<0.1 μM), while their posterior sections were consistently exposed to sulfide concentrations in the 100 μM range. A rootball-like structure formed the common base of the aggregation. In contrast to the anterior sections of the tubeworm tubes, the portions of the tubes within the “rootball” were freely permeable to sulfide. The results of this study show that R. piscesae, unlike vestimentiferans from the East Pacific Rise, can survive and grow in areas of low diffuse vent flow with very low plume-level exposure to sulfide. We propose that this morphotype of R. piscesae has the ability to acquire sulfide from sources near their posterior ends, similar to some species of cold seep vestimentiferans from the Gulf of Mexico. The ability of this single species of vestimentiferan to survive low exposure to vent flow with low mortality coupled with sulfide uptake across posterior tube sections may help explain the occurrence of a single vent vestimentiferan species in a wide variety of habitat conditions at hydrothermal vent sites in the Northeast Pacific.  相似文献   

5.
A new gas-tight isobaric sampler for the collection of hydrothermal fluids venting at the seafloor has been designed, constructed, and tested at a ridge-crest vent site. The new device is constructed of chemically inert titanium, is gas-tight to 450 bar and can be used to sample fluids with temperatures up to 400°C. Compressed gas is used to maintain the sample at seafloor pressure before and during sample withdrawal onboard ship, allowing subsampling without degassing the fluid remaining in the sampler. This feature eliminates the need to collect separate gas-tight and major element samples since a single fluid sample can be analyzed quantitatively for major, trace, semi-volatile, and volatile components. The sampler fill rate is regulated to minimize entrainment of ambient seawater during collection of fluids from environments characterized by low fluid flow such as diffuse hydrothermal vents. In addition to deployment at the ridge-crest, the samplers can be used to collect gas-tight samples from other subseasurface environments such as hydrocarbon seeps, areas of methane-gas hydrate formation, cold seeps associated with serpentinites, regions of groundwater egress to the oceans, and the water column.  相似文献   

6.
The Qiangtang Basin is a significant prospective area for hydrocarbon and gas hydrate resources in the Tibetan Plateau, China. However, relatively little work has been performed to characterise heat flow in this basin, which has restricted petroleum and gas hydrate exploration. In this study, we compare present and palaeo-heat flow in the Qiangtang Basin to provide information on geothermal regime, hydrocarbon generation and permafrost that is necessary for further petroleum and gas hydrate exploration. We base our study on temperature data from a thermometer well, thermal conductivity tests, vitrinite reflectance data, homogenisation temperature data from fluid inclusions, stratigraphic information and a time-independent modelling approach. Our results indicate that in the central Qiangtang Basin, the present thermal gradient is approximately 15.5 °C/km, and heat flow is approximately 46.69 mW/m2. Heat flow in the Qiangtang Basin is not relatively stable since the Early Jurassic, as previous research has suggested, and it is generally decreasing with time. Additionally, there is a clear difference between the hottest thermal regime of the southern and northern Qiangtang Depressions during Cretaceous to Pleistocene time. In the southern Qiangtang Depression, the palaeogeothermal gradient is approximately 32.0 °C/km, and palaeo-heat flow is approximately 70 mW/m2. However, in the northern Qiangtang Depression, the palaeogeothermal gradient exceeds 81.8 °C/km, and palaeo-heat flow is greater than 172.09 mW/m2. The high thermal regime in the northern Qiangtang Depression is driven mainly by hydrothermal convection. Gas reservoirs are possible targets for hydrocarbon exploration in this depression. Currently, the northwestern part of the northern Qiangtang Depression is the most favourable area for gas hydrate exploration in the Qiangtang Basin.  相似文献   

7.
Habitat created or modified by the physical architecture of large or spatially dominant species plays an important role in structuring communities in a variety of terrestrial, aquatic, and marine habitats. At hydrothermal vents, the giant tubeworm Riftia pachyptila forms large and dense aggregations in a spatially and temporally variable environment. The density and diversity of smaller invertebrates is higher in association with aggregations of R. pachyptila than on the surrounding basalt rock seafloor. Artificial substrata designed to mimic R. pachyptila aggregations were deployed along a gradient of productivity to test the hypothesis that high local species diversity is maintained by the provision of complex physical structure in areas of diffuse hydrothermal flow. After 1 year, species assemblages were compared among artificial aggregations in low‐, intermediate‐, and high‐productivity zones and compared to natural aggregations of R. pachyptila from the same site. Hydrothermal vent fauna colonized every artificial aggregation, and both epifaunal density and species richness were highest in areas of high chemosynthetic primary production. The species richness was also similar between natural aggregations of R. pachyptila and artificial aggregations in intermediate‐ and high‐productivity zones, suggesting that complex physical structure alone can support local species diversity in areas of chemosynthetic primary production. Differences in the community composition between natural and artificial aggregations reflect the variability in microhabitat conditions and biological interactions associated with hydrothermal fluid flux at low‐temperature hydrothermal vents. Moreover, these local ecological factors may further contribute to the maintenance of regional species diversity in hydrothermal vent communities on the East Pacific Rise.  相似文献   

8.
The saddle dolomites occur more intensely in cores closely to fault than that in cores far away from the fault in Upper Cambrian carbonate of western Tarim basin, suggesting that formation of the saddle dolomites is likely related to fault-controlled fluid flow. They partially fill in fractures and vugs of replacement dolomite. The saddle dolomites exhibit complex internal textures, commonly consisting of core and cortex. In comparison with the matrix dolomites, the saddle dolomites show lower Sr-content and 87Sr/86Sr ratios, higher Fe- and Mn-content, and more negative δ18O values. Combined with high Th (100–130 °C) of primary fluid inclusions, it is suggested that the saddle dolomites precipitated from hydrothermal fluid derived from the deep evaporite-bearing Middle Cambrian strata, and the magnesium source may be due to dissolution of host dolomite during hydrothermal fluid migration. Fault activity resulted in petrographic and geochemical difference of the core and cortex of the saddle dolomites. The cores precipitated from the formation water mixed by deep brines at the early stage of fault activity, and the cortexes precipitated from the deep fluid with higher temperatures through the Middle Cambrian later. In summary, the formation of the saddle dolomites implies a hydrothermal fluid event related to fault activity, which also resulted in high porosity in Upper Cambrian carbonate in western Tarim Basin.  相似文献   

9.
Significant quantities of fluids and dissolved geochemical components are expelled through the sediment surface in ocean margin and sedimented ridge environments. Recently, significant interest has been generated in constraining hydrological processes in these environments, but direct measurement of fluid flow in the marine environment has proven to be difficult and many aspects of marine hydrogeology remain poorly understood. To address the need for a means to make a significant number of direct measurements in a wide range of low to moderate flow environments, we have developed a new type of benthic aqueous flux meter that is capable of measuring diffuse fluid flow through the sediment surface on the order of 0.1 mm yr−1–15 m yr−1 when the flow is through sediments with permeabilities of less than 10−8 cm2 (typical seafloor sediments). The instrument measures fluid flow by determining the degree of dilution of a chemical tracer that is injected by an osmotic pump at a known rate into the fluids venting into or out of a collection chamber situated on the sea bed. The pump also withdraws a subsample of this tracer/fluid mix into sample coils allowing a serial record of the flow rates to be determined. Both upward and downward flow can be measured and, when flux rates are high enough to effectively flush the collecting chamber, the instruments also act as geochemical samplers. Three years of laboratory testing and field use have constrained the effects of (1) temperature, pressure, and deployment duration on osmotic pump performance, (2) dispersion/diffusion in the sample coils, and (3) deflection of flow under a range of sediment permeabilities. Recent deployments on the Kodiak and Cascadia accretionary prisms document the range and capabilities of the instrument in the field.  相似文献   

10.
A phase of ferroan burial calcite from the Middle Jurassic Lincolnshire Limestone exhibits a systematic spatial arrangement of oxygen isotopic characteristics. Mean δ18O values of the ferroan calcites from each of 15 core and outcrop localities over a study area 25 × 25 km were obtained. These values show a marked depletion from west to east across the study area of approximately 3‰, such that the oxygen isotopic composition of the ferroan calcites can be contoured. The systematic change in oxygen isotopic composition across the study area is believed to have recorded the thermal gradient in the limestone during ferroan calcite precipitation. This thermal gradient can be partially attributed to approximately 200 m of differential burial of the Lincolnshire Limestone across the study area during the Chalk deposition, with a maximum burial of 550 m to the east of the area at this time. A component of up-dip fluid flow (from east to west) through the formation is required to generate the temperature enhancements above those predicted for conduction alone by simple differential burial. Using a finite-difference step computer program, rates of fluid flow during ferroan calcite precipitation are calculated to be approximately 25 m/year. This rate of fluid flow is considerably greater than rates usually predicted for buried sedimentary basins. The causes of such rapid, probably relatively short-lived flow-rates may be the sudden dewatering of adjacent shales, the release of overpressure within the formation of interest, seismic pumping, or fluid circulation round a supracrustal convective loop.  相似文献   

11.
《Marine Geology》2006,225(1-4):191-208
The pore water chemistry of mud volcanoes from the Olimpi Mud Volcano Field and the Anaximander Mountains in the eastern Mediterranean Sea have been studied for three major purposes: (1) modes and velocities of fluid transport were derived to assess the role of (upward) advection, and bioirrigation for benthic fluxes. (2) Differences in the fluid chemistry at sites of Milano mud volcano (Olimpi area) were compiled in a map to illustrate the spatial heterogeneity reflecting differences in fluid origin and transport in discrete conduits in near proximity. (3) Formation water temperatures of seeping fluids were calculated from theoretical geothermometers to predict the depth of fluid origin and geochemical reactions in the deeper subsurface.No indications for downward advection as required for convection cells have been found. Instead, measured pore water profiles have been simulated successfully by accounting for upward advection and bioirrigation. Advective flow velocities are found to be generally moderate (3–50 cm y− 1) compared to other cold seep areas. Depth-integrated rates of bioirrigation are 1–2 orders of magnitude higher than advective flow velocities documenting the importance of bioirrigation for flux considerations in surface sediments. Calculated formation water temperatures from the Anaximander Mountains are in the range of 80 to 145 °C suggesting a fluid origin from a depth zone associated with the seismic decollement. It is proposed that at that depth clay mineral dehydration leads to the formation and advection of fluids reduced in salinity relative to sea water. This explains the ubiquitous pore water freshening observed in surface sediments of the Anaximander Mountain area. Multiple fluid sources and formation water temperatures of 55 to 80 °C were derived for expelled fluids of the Olimpi area.  相似文献   

12.
An In Situ Ultraviolet Spectrophotometer (ISUS) was coupled to a benthic chamber to characterize the bisulfide flux emanating from a warm spring at the Extrovert Cliff locality within Monterey Bay, California. The chamber was periodically flushed with bottom seawater to reset chemical concentrations, which enabled deployments over multiple days. Data from several deployments, each lasting at least 10 days, were used to calculate flow rates, fluid concentrations, and fluxes over time. The bisulfide concentration of the fluid entering the chamber varied from 75 to 4500 μmol l?1. Positive temperature anomalies up to 3.5° were associated with these elevated concentrations. Linear flow rates ranged from 2 to >17 m day?1, while the bisulfide fluxes varied from 0.2 to 80 mol m?2 day?1. The bisulfide originated at depth and was not produced in the surface sediments via an anaerobic oxidation of methane coupled to sulfate reduction. Tides modulated the flow as well as the composition of the fluid entering the chamber. It appeared that a deep sourced fluid, which supplied the bisulfide, was mixed with a second, ambient seawater-like fluid before entering the flux chamber. At low tides, flow rates were at their highest and the contribution of the deep sourced fluid to the fluid entering the chamber was at a maximum.  相似文献   

13.
Fluid migration rates are important parameters for understanding the structural characteristics and evolution of the crustal tectonics and hydrocarbon exploration. However, they are difficult to measure on the seafloor. Densely sampled temperature measurements might shed light on our study of the fluid migration rates. In this study, we first use reflection seismic data to derive geothermal gradient patterns at different sub-seafloor depths, then calculated 1D vertical fluid flow models by analyzing the Péclet numbers in the offshore regions of SW Taiwan. We found Péclet numbers ranging from 2 to 5, implying that vertical fluid flow velocities are between 6 and 44 cm/year. Assumed a homogenous permeable layer and no lateral variation of the temperature fields, we analyzed possible errors caused by depth-dependent thermal conductivity and velocity-depth function. They probably cause less than 2 cm/year of errors in the estimated vertical fluid flow rate. In terms of regional fluid flow patterns, we found higher fluid flow rates near the toe of the trench, and a dramatic increase in fluid flow rate when the continental slope of the Chinese passive margin enters into the trench, and reduced fluid flow rates in the hinterland. We propose that this is a new and potentially useful method to derive regional fluid flow rate models for studying geochemical and biological processes in shallow seafloor sediments and the regional hydrological budget.  相似文献   

14.
Faulting in Middle Jurassic reservoirs occurred at shallow depth during regional extension. Clean sandstones (<15% clay) deformed without significant grain fracturing and permeability reduction. Faults in impure sandstones (15–40% clay) experienced significant syn-deformation compaction and permeability reduction. Enhanced compaction during deeper burial reduced their permeabilities further from an average of 0.05 mD at <2.5 km to 0.001 mD at >4 km. Clay-rich sediments (>40% clay) deformed to produce clay smears with very low permeabilities (<0.001 mD). Faulting in the Rotliegendes occurred at greater depth during both basin extension and inversion. Extensional faulting produced cataclasites with permeability reductions of <10–>106; their permeabilities range from 0.2 to 0.0001 mD and are inversely related to their maximum burial depth. Faults formed or reactivated during inversion experienced permeability increase. These results can be extrapolated to other hydrocarbon reservoirs if differences in stress and temperature history are taken into account.The permeability of most (>80%) faults is not sufficiently low, compared to their wallrock, to retard single-phase fluid flow on a km-scale. Nevertheless, most faults could retard the flow of a non-wetting phase if present at low saturations. It may be necessary to incorporate the two-phase fluid flow properties of fault rocks into reservoir simulators using upscaling or pseudoisation techniques. Fault property data should be calibrated against production data before it can be used confidently.  相似文献   

15.
Drilling/coring activities onboard JOIDES Resolution for hydrate resource estimation have confirmed gas hydrate in the continental slope of Krishna-Godavari (KG) basin, Bay of Bengal and the expedition recovered fracture filled gas hydrate at the site NGHP-01-10. In this paper we analyze high resolution multi-channel seismic (MCS), high resolution sparker (HRS), bathymetry, and sub-bottom profiler data in the vicinity of site NGHP-01-10 to understand the fault system and thermal regime. We interpreted the large-scale fault system (>5 km) predominantly oriented in NNW-SSE direction near NGHP-01-10 site, which plays an important role in gas hydrate formation and its distribution. The increase in interval velocity from the baseline velocity of 1600 m/s to 1750–1800 m/s within the gas hydrate stability zone (GHSZ) is considered as a proxy for the gas hydrate occurrence, whereas the drop in interval velocity to 1400 m/s suggest the presence of free gas below the GHSZ. The analysis of interval velocity suggests that the high concentration of gas hydrate occurs close to the large-scale fault system. We conclude that the gas hydrate concentration near site NGHP-01-10, and likely in the entire KG Basin, is controlled primarily by the faults and therefore has high spatial variability.We also estimated the heat flow and geothermal gradient (GTG) in the vicinity of NGHP-01-10 site using depth and temperature of the seafloor and the BSR. We observed an abnormal GTG increase from 38 °C/km to 45 °C/km at the top of the mound, which remarkably agrees with the measured temperature gradient at the mound (NGHP-01-10) and away from the mound (NGHP-01-03). We analyze various geological scenarios such as topography, salinity, thermal non-equilibrium of BSR and fluid/gas advection along the fault system to explain the observed increase in GTG. The geophysical data along with the coring results suggest that the fluid advection along the fault system is the primary mechanism that explains the increase in GTG. The approximate advective fluid flux estimated based on the thermal measurement is of the order of few tenths of mm/yr (0.37–0.6 mm/yr).  相似文献   

16.
Whilst the fauna inhabiting hydrothermal vent structures in the Atlantic Ocean is reasonably well known, less is understood about the spatial distributions of the fauna in relation to abiotic and biotic factors. In this study, a major active hydrothermal edifice (Eiffel Tower, at 1690 m depth) on the Lucky Strike vent field (Mid-Atlantic Ridge (MAR)) was investigated. Video transects were carried out by ROV Victor 6000 and complete image coverage was acquired. Four distinct assemblages, ranging from dense larger-sized Bathymodiolus mussel beds to smaller-sized mussel clumps and alvinocaridid shrimps, and two types of substrata were defined based on high definition photographs and video imagery. To evaluate spatial variation, faunal distribution was mapped in three dimensions. A high degree of patchiness characterizes this 11 m high sulfide structure. The differences observed in assemblage and substratum distribution were related to habitat characteristics (fluid exits, depth and structure orientation). Gradients in community structure were observed, which coincided with an increasing distance from the fluid exits. A biological zonation model for the Eiffel Tower edifice was created in which faunal composition and distribution can be visually explained by the presence/absence of fluid exits.  相似文献   

17.
The dimensions of sand ripples in full-scale oscillatory flows   总被引：1，自引：0，他引：1  
New large-scale experiments have been carried out in two oscillatory flow tunnels to study ripple regime sand suspension and net sand transport processes in full-scale oscillatory flows. The paper focuses on ripple dimensions and the new data are combined with existing data to make a large dataset of ripple heights and lengths for flows with field-scale amplitudes and periods. A feature of the new experiments is a focus on the effect of flow irregularity. The combined dataset is analysed to examine the range of hydraulic conditions under which oscillatory flow ripples occur, to examine the effects of flow irregularity and ripple three-dimensionality on ripple dimensions and to test and improve existing methods for predicting ripple dimensions.The following are the main conclusions. (1) The highest velocities in a flow time-series play an important role in determining the type of bedform occurring in oscillatory flow. Bedform regime is well characterised by mobility number based on maximum velocity in the case of regular flow and based on the mean of the highest one tenth peak velocities in the case of irregular flow. (2) For field-scale flows, sand size is the primary factor determining whether equilibrium ripples will be 2D or 3D. 2D ripples occur when the sand D50 ≥ 0.30 mm and 3D ripples occur when D50 ≤ 0.22 mm (except when the flow orbital diameter is low). (3) Ripple type (2D or 3D) is the same for regular and irregular flows and ripple dimensions produced by equivalent regular and irregular flows follow a similar functional dependence on mobility number, with mobility number based on maximum velocity in the case of regular flow and based on the mean of the highest one tenth velocities in the case of irregular flow. For much of the ripple regime, ripple dimensions have weak dependency on mobility number and ripple dimensions are similar for regular and irregular flows with the same flow orbital amplitude. However, differences in ripples produced by equivalent regular and irregular flows become significant at the high mobility end of the ripple regime. (4) Ripple dimensions predicted using the Wiberg and Harris formulae are in poor agreement with measured ripple dimensions from the large-scale experiments. Predictions based on the Mogridge et al. and the Nielsen formulae show better overall agreement with the data but also show systematic differences in cases of 3D ripples and ripples generated by irregular flows. (5) Based on the combined large-scale data, modifications to the Nielsen ripple dimension equations are proposed for the heights and lengths of 2D ripples. The same equations apply to regular and irregular flows, but with mobility number appropriately defined. 3D ripples are generally smaller than 2D ripples and estimates of 3D ripple height and length may be obtained by applying multipliers of 0.55 and 0.73 respectively to the 2D formulae. The proposed modified Nielsen formulae provide an improved fit to the large-scale data, accounting for flow irregularity and ripple three-dimensionality.  相似文献   

18.
This study provides an analysis of vesicomyid bivalve–symbiont community distribution across cold seep and hydrothermal vent areas in the Guaymas Basin (Gulf of California, Mexico). Using a combination of morphological and molecular approaches including fluorescent in situ hybridization (FISH), and electronic microscopy observations, vesicomyid clam species and their associated symbionts were characterized and results were analyzed in light of geochemical conditions and other on‐site observations. A greater diversity of vesicomyids was found at cold seep areas, where three different species were present (Phreagena soyoae [syn. kilmeri], Archivesica gigas, and Calyptogena pacifica). In contrast, A. gigas was the only species sampled across the hydrothermal vent area. The same haplotype of A. gigas was found in both hydrothermal vent and cold seep areas, highlighting possible contemporary exchanges among neighboring vents and seeps. In either ecosystem, molecular characterization of the symbionts confirmed the specificity between symbionts and hosts and supported the hypothesis of a predominantly vertical transmission. In addition, patterns of clams could reflect potential niche preferences for each species. The occurrence of numerous traces of vesicomyid movements on sediments in the sites colonized by A. gigas seemed to indicate that this species might have a better ability to move. Furthermore, variation in gill sulfur content could reveal a higher plasticity and sulfur storage capacity in A. gigas. Thus, the distribution of vesicomyid species across the chemosynthetic areas of the Guaymas Basin could be explained by differences in biological traits of the vesicomyid species that would allow A. gigas to more easily exploit transient and punctual sources of available sulfide than P. soyoae.  相似文献   

19.
Recently it has been observed that a strong quantitative relationship exists between asymptotic fluxes of particulate organic carbon (POC) to the deep ocean and asymptotic fluxes of “ballast” minerals (opal; calcium carbonate; dust). It has further been suggested that this relationship might provide a mechanistic basis for improved representations of remineralization in ocean carbon models. Since the depth scale of remineralization z* is the ratio k/v of a remineralization rate k and a settling velocity (SV) v, a mechanistic understanding of settling velocities will be crucial in developing such models.Historically, there have been two approaches to estimating the speed with which POC is transported to the deep ocean. First, settling speeds of single particles have been observed directly in both field and laboratory settings; estimates of fecal pellet sinking velocities tend to be higher and more variable than those of aggregates. Second, estimates have been made of the velocity at which temporal patterns in flux propagate between pairs of sediment traps separated in depth (the “benchmark approach”); recent studies have shown these results to be variable and to depend on mineral ballasting. Here we present SV estimates using a relatively new technology: indented rotating sphere (IRS) sediment traps run in settling velocity (SV) mode. In this approach, particles are separated into SV classes during settling to collection cups. In MedFlux, SV data were collected concurrently with time-series (TS) data; the latter were used to construct benchmark estimates for comparison to the SV estimates. From the SV data, the range of modal settling velocities (sinking velocities having the largest time-averaged mass flux densities on a logarithmic scale of SV) in the fast-sinking fraction was estimated to be 287–503 m/d; the average of these modal values is 353 m/d, with standard deviation 76 m/d. In contrast, mean settling velocities of the fast-sinking fraction depend on the range of settling velocity classes included in the estimate. If only SV classes settling at >50 m/d are included, the range of SVs at MedFlux was 214–298 m/d, with average mean value 242 m/d and standard deviation 31 m/d. These mean-velocity results are in excellent agreement with benchmark estimates of signal propagation velocities at Medflux (220±65 m/d); they are also well within the range of other recent benchmark studies. The agreement between the benchmark estimates and mean settling velocity estimates at MedFlux, but not with modal velocities, argues that the benchmark method estimates mean settling velocities.  相似文献   

20.
In order to estimate the contribution of cold Pacific deep water to the Indonesian throughflow (ITF) and the flushing of the deep Banda Sea, a current meter mooring has been deployed for nearly 3 years on the sill in the Lifamatola Passage as part of the International Nusantara Stratification and Transport (INSTANT) programme. The velocity, temperature, and salinity data, obtained from the mooring, reflect vigorous horizontal and vertical motion in the lowest 500 m over the ～2000 m deep sill, with speeds regularly surpassing 100 cm/s. The strong residual flow over the sill in the passage and internal, mainly diurnal, tides contribute to this bottom intensified motion. The average volume transport of the deep throughflow from the Maluku Sea to the Seram Sea below 1250 m is 2.5 Sv (1 Sv=106 m3/s), with a transport-weighted mean temperature of 3.2 °C. This result considerably increases existing estimates of the inflow of the ITF into the Indonesian seas by about 25% and lowers the total mean inflow temperature of the ITF to below 13 °C. At shallower levels, between 1250 m and the sea surface, the flow is directed towards the Maluku Sea, north of the passage. The typical residual velocities in this layer are low (～3 cm/s), contributing to an estimated northward flow of 0.9–1.3 Sv. When more results from the INSTANT programme for the other Indonesian passages become available, a strongly improved estimate of the mass and heat budget of the ITF becomes feasible.  相似文献   

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