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1.
The seasonal and spatial distribution (density and biomass) of five size classes of two catfish species (Cathorops spixii and Cathorops agassizii) were studied along an estuarine ecocline to test the relative importance of the nursery function of each habitat. Seasonal vs. area interactions were significant for all size classes of both species. During the early rainy season, the middle estuary is an important nursery habitat for juveniles of both species. When environmental conditions change during the late rainy season, the C. spixii primary nursery habitat shifts to the lower estuary. During this period, juveniles of C. agassizii remain in the middle estuary. Another important ecological area is the upper estuary, which becomes a breeding, spawning and hatchery area during the late dry season for both species. The nursery function of habitats shifts according to the seasonal fluctuation of salinity and dissolved oxygen, and each species responds differently to this change.  相似文献   

2.
Estuarine habitat occupied by Alligator mississippiensis, a primarily freshwater species, is spatially and temporally heterogeneous largely due to a salinity gradient that fluctuates. Using long-term night light survey data, we examined seasonal patterns in alligators’ habitat use by size classes in midstream and downstream estuary zones of Shark River, Everglades National Park, in southern Florida. We observed predominantly large-sized alligators (total length?≥?1.75 m); observations of alligators in the small size classes (0.5 m?≤?total length?<?1.25 m) were rare especially in the higher-salinity downstream zone. The density of alligators in the downstream zone was lower than that of the midstream zone during the dry season when salinity increases due to reduced precipitation. Conversely, the density of the large size alligators was higher in the downstream zone than in the midstream zone during the wet season, likely because of reduced salinity. We also found a significant declining trend over time in the number of alligators in the dry season, which coincides with the reported decline in alligator relative density in southern Florida freshwater wetlands. Our results indicated high adaptability of alligators to the fluctuating habitat conditions. Use of estuaries by alligators is likely driven in part by physiology and possibly by reproductive cycle, and our results supported their opportunistic use of estuary habitat and ontogenetic niche shifts.  相似文献   

3.
Identification of food web linkages is a major aim in ecology because it provides basic information on trophic flows and the potential for interspecific interactions. In addition, policy and restoration measures mandated to conform to ecosystem-based management principals can benefit from information on temporal and spatial variability in community-level interactions. Here, we analyzed guild structure of the demersal fish assemblage in Puget Sound, WA, a temperate estuarine system on the US west coast. Using diet information from 2,401 stomachs collected across three seasons (fall, winter, and summer), we identified guild membership for 21 fish species, examined seasonal guild switching, and tested for seasonal shifts in predation and for differences in the degree of diet overlap at the assemblage level. We accounted for ontogenetic variation in diet by dividing species into large (L) and small (S) size classes when appropriate. Using cluster analysis and a permutation approach, we identified seven significant guilds that were typified by predation on benthic invertebrates, pelagic invertebrates, and piscivory. Of the 18 species with more than one season of diet information, six switched guilds (Pacific sanddab L, sturgeon poacher, Pacific tomcod S, speckled sanddab, rex sole, and rock sole S). At the assemblage level, we tested for seasonal differences in prey use between seasons by performing an analysis of similarities based on Bray–Curtis diet similarities and found no significant difference. However, diet overlap was significantly higher in the summer than the fall and winter (with summer > fall > winter) indicating that diets within the assemblage converged in the summer. These results indicate that analyses of guild structure and diet overlap can reveal seasonal variation in community trophic structure and highlight intra-annual food web variation in the Puget Sound demersal fish community.  相似文献   

4.
Fish diets play a critical role in our understanding of aquatic trophic dynamics and are an important component in developing ecosystem-based approaches to fisheries management. Although large nektonic fishes exert top-down predator effects on the food web and typically support viable commercial and recreational fisheries, little is known about the diet of this guild. We evaluated the diets (6327 stomachs) of four nektonic predatory fishes (Pomatomus saltatrix [78–395 mm], Cynoscion regalis [91–520 mm], Morone americana [156–361 mm], and Morone saxatilis [82–785 mm]) in Delaware Bay and in the adjacent ocean. To assess ontogenetic, geographic, and interspecific variation in diets, observations from individual fish stomachs were clustered into species-size class groups, and dietary overlap was estimated using multivariate analyses. A shift in diet composition, as well as diversity, occurred along the estuarine gradient and into the adjacent ocean. Some prey were shared by most predators, including some crustaceans (dominated by Callinectes sapidus, mysids, and Palaemonetes spp.), fundulids (dominated by Fundulus heteroclitus), engraulids (dominated by Anchoa mitchilli), and clupeids (dominated by Brevoortia tyrannus). However, inter- and intra-specific variation in diet was observed as well. In particular, M. americana consumed fewer engraulids and clupeids, and many more and diverse types of invertebrates, while P. saltatrix consumed more clupeids and less invertebrates. The lack of overlap in diet between the four predators evaluated, and between size groups for each predator, supports previous evidence that these groups feed in trophic guilds defined by species and by size within a species. The highly variable diets for these predators suggest high resolution spatial data are necessary in order to quantify their most important prey and their role in coastal ecosystems.  相似文献   

5.
Diversity of Mn oxides produced by Mn(II)-oxidizing fungi   总被引：1，自引：0，他引：1  
Manganese (Mn) oxides are environmentally abundant, highly reactive mineral phases that mediate the biogeochemical cycling of nutrients, contaminants, carbon, and numerous other elements. Despite the belief that microorganisms (specifically bacteria and fungi) are responsible for the majority of Mn oxide formation in the environment, the impact of microbial species, physiology, and growth stage on Mn oxide formation is largely unresolved. Here, we couple microscopic and spectroscopic techniques to characterize the Mn oxides produced by four different species of Mn(II)-oxidizing Ascomycete fungi (Plectosphaerella cucumerina strain DS2psM2a2, Pyrenochaeta sp. DS3sAY3a, Stagonospora sp. SRC1lsM3a, and Acremonium strictum strain DS1bioAY4a) isolated from acid mine drainage treatment systems in central Pennsylvania. The site of Mn oxide formation varies greatly among the fungi, including deposition on hyphal surfaces, at the base of reproductive structures (e.g., fruiting bodies), and on envisaged extracellular polymers adjacent to the cell. The primary product of Mn(II) oxidation for all species growing under the same chemical and physical conditions is a nanoparticulate, poorly-crystalline hexagonal birnessite-like phase resembling synthetic δ-MnO2. The phylogeny and growth conditions (planktonic versus surface-attached) of the fungi, however, impact the conversion of the initial phyllomanganate to more ordered phases, such as todorokite (A. strictum strain DS1bioAY4a) and triclinic birnessite (Stagonospora sp. SRC1lsM3a). Our findings reveal that the species of Mn(II)-oxidizing fungi impacts the size, morphology, and structure of Mn biooxides, which will likely translate to large differences in the reactivity of the Mn oxide phases.  相似文献   

6.
Effects of petroleum covered substrate on intertidal oyster spat (Crassostrea virginica) set were measured at three intertidal elevations in a southeastern North Carolina estuary.Mercenaria mercenaria shells were coated with Bunker C crude oil or a 40∶1 mixture of gasoline: 2-cycle engine oil and placed intertidally for seven 13-d periods. Spat densities were significantly lower on oil treatments versus control and gas-treated shells in the high intertidal zone. This was principally attributed to an increased sediment coat on oiled shells. Maximum spat size was smaller on oil-treated shells at all elevations when compared to gas and control shells, indicating that setting may be delayed on oiled shell. For all experimental 13-d periods in the low intertidal zone and for three periods in the mid-tidal zone, barnacle densities (primarilyBalanus improvisus andB. eburneus) were significantly greater on oiled shells than on control shells.  相似文献   

7.
Early Cambrian univalve molluscs are predominantly represented by microscopic forms (body length of 1–3 mm), preserved mainly as phosphatised internal moulds with limited definable features. Macromolluscs (≥ 5 mm) are generally rare, occur in low abundance and are poorly preserved, often lacking apical features and ornament which hinders taxonomic assessment. New and previously undescribed material from lower Cambrian Hawker Group carbonates of the Flinders Ranges in South Australia includes four new taxa, Minastirithella silivreni gen. et sp. nov., Galeacalvus coronarius gen. et sp. nov., Helcionella histosia sp. nov., and Ilsanella enallaxa sp. nov. Three-dimensional silicified shell material preserved with intact apices offers new insight into protoconch morphology, growth habit (isometric vs. allometric) and developmental mode. This material supports previous suggestions that some micromollusc taxa may in fact be early ontogenetic stages (juveniles) of larger macroscopic taxa; such that the millimetric size range of helcionelloids conforms to the dimensions of earliest apical portion in some macromolluscs documented herein. However, taphonomic limitations associated with phosphatisation show that the morphology (especially height vs. width) of the apex can greatly influence the probability of steinkern formation and preservation potential for both micro- and macro-scale helcionelloids. Artificial inflation in the appearance of millimetric helcionelloids with an optimal morphology for phosphatisation is thus directly linked to taphonomic biases in the fossil record.  相似文献   

8.
We examined ontogenetic, interspecific, and seasonal trophic patterns among sympatric sunfish species, redspotted sunfish,Lepomis miniatus; redear sunfish,Lepomis microlophus; and bluegill,Lepomis macrochirus, in an estuarine bayou. In particular we studied these feeding patterns in relation to relative abundances of prey from different benthic feeding habitats. All three sunfishes showed ontogenetic divergence in their trophic niches, reflecting different ecomorphological specializations. Small fishes depended on zooplankton, whereas larger fishes of all three species shifted their diets to benthic macrofauna. A potential for trophic resource partitioning was reflected by dietary differences among the three sunfish species. One impalied mechanism for resource partitioning was feeding habitat, as redear sunfish frequently used sediment-associated prey, while bluegill showed greater use of water-column-associated prey, and redspotted sunfish often used SAV-associated prey. However, all three sunfishes apparently used each feeding habitat to some degree; and, trophic differences were more clearly based on prey type than on feeding habitat. Redear sunfish, which can crush hard-shelled prey, exhibited the most distinctive diet. An apparent seasonal shift in feeding habitat occurred in autumn/winter, as indicated by increased overlap between diets and SAV. This shift was facilitated by changes in the relative abundances of several common prey types between benthic habitats. The relative abuandance and use of freshwater and estuarine-derived prey also varied seasonally, suggesting a possible trophic benefit of consistent prey availability in the estuarine bayou.  相似文献   

9.
This paper forms the first part of a revision of the ammonite faunas of the Cenomanian stratotype in the environs of Le Mans, Sarthe, France. The history of research on ammonites in the area is briefly noted, as is the lithostratigraphy. The systematics of the heteromorph ammonites (Ancyloceratina) is described, and the following taxa documented: Hamites simplex d'Orbigny, Sciponoceras baculoides (Mantell), Sciponoceras gracile (Shumard), Anisoceras plicatile (J. Sowerby), A. aff. plicatile, Idiohamites alternatus vectensis Spath, I. ellipticus radiatus Spath, Hypoturrilites gravesianus (d'Orbigny), H. tuberculatus (Bosc), H. mantelli (Sharpe), H. sp., Neostlingoceras carcitanense (Matheron), N. aff. carcitanense, Turrilites costatus Lamarck, T. acutus Passy, T. scheuchzerianus (Bosc), T. boerssumensis Schlüter, Mariella dorsetensis (Spath), M. cenomanensis (Schlüter), M. group of cenomanensis (Schlüter)-lewesiensis (Spath), M. spp., M. (Plesioturrilites) sp., Scaphites equalis J. Sowerby, S. obliquus J. Sowerby, S.(?) sp., and S. sp. juv.  相似文献   

10.
The impact of mesozooplankton (>210 μm, mostly adult copepods and late-stage copepodites) and micrometazoa (64–210 μm, mostly copepod nauplii) on phytoplankton size structure and biomass in the lower Hudson River estuary was investigated using various14C-labeled algal species as tracers of grazing on natural phytoplankton. During spring and summer, zooplankton grazing pressure, defined as %=mg C ingested m?2 h?1/mg C produced m?2 h?1 (depth-integrated rates)×100, on total phytoplankton ranged between 0.04% and 1.9% for mesozooplankton and 0.1% and 6.6% for micrometazoa. The greatest grazing impact was measured in fall when 20.2% and 44.6%, respectively, of the total depth-integrated primary production from surface water phytoplankton was grazed. Mesozooplankton exhibited some size-selective grazing on phytoplankton, preferentially grazing the diatomThalassiosira pseudonana over the larger diatomDitylum brightwelli, but this was not found for micrometazoa. Neither zooplankton group grazed on the dinoflagellateAmphidinium sp. We conclude that metazoan zooplankton have a minimal role in controlling total phytoplankton biomass in the lower Hudson River estuary. Differences in the growth coefficients of various phytoplankton size-fractions—not grazing selectivity—may be the predominant factor explaining community size-structure.  相似文献   

11.
《China Geology》2021,4(2):197-204
The authors reassessed the taxonomic distinction of Iteravis huchzermeyeri and Gansus zheni, which are two species of Ornithuromorpha based on specimens from the same locality in western Liaoning and derive from the Jehol Biota. The detailed comparisons of the holotype and referred specimens of both species, reveal no anatomical features that distinguish these taxa as separate species. Some minor differences are considered to relate to ontogenetic or interspecific differences. The stratigraphic occurrence for both specimens is the Lower Cretaceous Jiufotang Formation. Accordingly, the authors conclude that Iteravis huchzermeyeri has priority, by 15 days, for this taxon and that Gansus zheni is a junior synonym. The diagnosis of Iteravis huchzermeyeri is revised based on further study on all specimens referred to this species. Its generic distinction from Gansus is maintained thereby removing a potential genus-level correlation linking Xiagou Formation in Gansu Province with the Jiufotang Formation in Liaoning Province.  相似文献   

12.
As top predators, marine mammals and seabirds play an important role in structuring marine ecosystems, and their trophic role have become a major issue. In this study, we quantify the trophic impact of marine mammals and seabirds in the Río de la Plata estuary based on a previous trophic mass balance model of this estuary. The model comprised 37 functional groups: 3 species of marine mammals (2 cetaceans: Tursiops truncatus, Pontoporia blainvillei and 1 pinniped: Otaria flavescens), 1 coastal bird group comprising 11 species, 17 fish, 12 invertebrates, 2 zooplanktons, 1 phytoplankton, detritus, and 5 fishing fleets. Model results showed that seabirds, O. flavescens and P. blainvillei, are apex predators, with high levels of niche overlap among them, suggesting competition for similar resources such as fish species. Marine mammals and seabirds produce negative effects on commercially important species, while, at the same time, indirect positive effects (increase of the biomass) were also detected in some groups related to trophic cascade effects.  相似文献   

13.
This paper deals with the spatial and seasonal recycling of organic matter in sediments of two temperate small estuaries (Elorn and Aulne, France). The spatio-temporal distribution of oxygen, nutrient and metal concentrations as well as the organic carbon and nitrogen contents in surficial sediments were determined and diffusive oxygen fluxes were calculated. In order to assess the source of organic carbon (OC) in the two estuaries, the isotopic composition of carbon (δ 13C) was also measured. The temporal variation of organic matter recycling was studied during four seasons in order to understand the driving forces of sediment mineralization and storage in these temperate estuaries. Low spatial variability of vertical profiles of oxygen, nutrient, and metal concentrations and diffusive oxygen fluxes were monitored at the station scale (within meters of the exact location) and cross-section scale. We observed diffusive oxygen fluxes around 15 mmol m?2 day?1 in the Elorn estuary and 10 mmol m?2 day?1 in the Aulne estuary. The outer (marine) stations of the two estuaries displayed similar diffusive O2 fluxes. Suboxic and anoxic mineralization was large in the sediments from the two estuaries as shown by the rapid removal of very high bottom water concentrations of NO x ? (>200 μM) and the large NH4 + increase at depth at all stations. OC contents and C/N ratios were high in upstream sediments (11–15 % d.w. and 4–6, respectively) and decreased downstream to values around 2 % d.w. and C/N ≤ 10. δ 13C values show that the organic matter has different origins in the two watersheds as exemplified by lower δ 13C values in the Aulne watershed. A high increase of δ 13C and C/N values was visible in the two estuaries from upstream to downstream indicating a progressive mixing of terrestrial with marine organic matter. The Elorn estuary is influenced by human activities in its watershed (urban area, animal farming) which suggest the input of labile organic matter, whereas the Aulne estuary displays larger river primary production which can be either mineralized in the water column or transferred to the lower estuary, thus leaving a lower mineralization in Aulne than Elorn estuary. This study highlights that (1) meter scale heterogeneity of benthic biogeochemical properties can be low in small and linear macrotidal estuaries, (2) two estuaries that are geographically close can show different pattern of organic matter origin and recycling related to human activities on watersheds, (3) small estuaries can have an important role in recycling and retention of organic matter.  相似文献   

14.
Properties of fluorescent dissolved organic matter in the Gironde Estuary   总被引：5，自引：0，他引：5  
The isolation, characterization and study of the properties of aquatic dissolved organic matter (DOM) still represent a challenge because of the heterogeneity, complexity and low concentration of organic material in natural waters. Based on its ability to interact with contaminants and thus to modify their transport and bioavailability, DOM is of interest for environmental purposes. The objective of this work was to better characterize DOM in the Gironde Estuary (southwestern France). The estuary represents an exchange zone between the continent and the Atlantic Ocean and conditions the transfer of organic and inorganic substances from the continental to the oceanic environment. Several samples were collected along the estuary during three cruises in 2002 and 2006. They were analysed using excitation–emission matrix (EEM) spectroscopy, a sensitive technique that allows direct analysis of water samples. Fluorescent DOM and dissolved organic carbon (DOC) did not behave conservatively in this estuarine system, i.e. the organic material did not undergo simple dilution from the upstream to the downstream part of the estuary. A seasonal variability in DOC content was pointed out, whereas few seasonal variations in DOM fluorescence were observed. DOM sources and processing in the estuary were further evaluated by determining two fluorescence indices – the humification index (HIX) and the index of recent autochthonous contribution (BIX). By applying these indices, the relative degree of humification (HIX) and autotrophic productivity (BIX) could be assessed. Based on the fluorescence and DOC results, the estuary was divided into three zones depending on salinity (S) and characterized by specific DOM: (i) A turbid zone of low salinity (S < 5) and high suspended particulate matter concentration with increase in the intensities of the α′ and α fluorophores, characteristic of humic-like compounds. (ii) A mid-estuarine zone (5 < S < 25) characterized by low autotrophic productivity and containing strongly degraded organic material, as shown by the low values of BIX and high values of HIX. (iii) A higher salinity area (S > 25) characterized by increased autotrophic productivity and a marked marine influence, and associated with high and low values of BIX and HIX, respectively. The HIX and BIX indices were shown as useful tools for readily defining and classifying DOM characteristics in estuarine waters.  相似文献   

15.
During the 14-month collection period,Hemicyclops always appeared as naupliar stages andSaphirella as immature copepodite stages at 3 stations in the middle and lower estuary.Hemicyclops nauplii were abundant during late spring and summer, with a seasonal maximum of 7448/m3 in late spring at the upstream station.Saphirella stages were observed throughout the year, but higher population densities generally occurred in the summer and fall; the highest density was 3413/m3. Both species were usually more abundant upstream than downstream. The larvae may be life stages of a single bottom-dwelling species,Hemicyclops adhaerens.  相似文献   

16.
Suspended particle samples from the Yellow River estuary were sorted into five grain size fractions to explore the effect of grain size distribution on organic matter content and composition. The n-alkanes and PAHs were determined for each size fraction. PAHs and n-alkanes were more abundant in the finer fractions and the loading decreases steadily with increasing of grain size. However, the total n-alkanes or PAHs normalized to organic C were lower in the smaller size fractions than those in the larger size fractions, suggesting n-alkanes or PAHs may be diluted by the addition of organic matter or gradually decreased by degradation in the smaller size fractions. The particulate n-alkanes in the Yellow River estuary consist of a mixture of compounds from terrigenous and riverine biogenic n-alkanes and more biogenic n-alkanes accumulate in finer particles. Particulate PAHs are related to combustion/pyrolysis processes of coal/wood, and the relative contribution of petrogenic PAHs increase with increasing grain size. The total particulate n-alkane and PAH discharges passing the Lijin Station are about 3.94 t d−1 and 0.52 t d−1, respectively. Fine particles (<32 μm) play a significant role in organic matter transfer.  相似文献   

17.
The behavior of the leatherjack, Oligoplites saurus, was studied in the field. Four behavioral patterns were identified: (1) cleaning behavior on the redfin needlefish, Strongylura notata, (2) intraspecific schooling, (3) interspecific schooling with the rough and tidewater silversides, Membras martinica and Menidia peninsulae, pinfish, Lagodon rhomboides, and menhaden, Brevoortia sp., (4) swimming along in a “leafmimic” position or next to floating algae or debris. Fishes exhibiting any of these behaviors fed primarily on plankton. Schooling leatherjackets also fed on scales of Strongylurus notata, Brevoortia sp., Menidia peninsulae and Lagodon rhomboides. These observations are supported by stomach content analyses. It is suggested that scale-feeding may be a preadaptation for cleaning behavior among leatherjackets.  相似文献   

18.
The vertical distribution and feeding type of nematodes in sediments of Chetumal Bay, Mexico, were studied in five intertidal transects along the urbanized zone in June and December 1995. Sediments were collected with a PVC corer to 6-cm sediment depth and cut immediately into three equal 2-cm depth fractions. Nematode density varied from 7.4 × 103 to 5.3 × 105 m?2 in June and from 1.7 × 104 to 7.2 × 105 m?2 in December. In June, the epistrate feederPseudochromadora sp. was the most abundant in the deepest sediment fraction (4–6 cm), whereas epistrate feeders,Neotonchoides sp.,Desmodora sp., and the deposit feederBathylaimus australis were dominant in the top most sediment (0–2 cm). In December, deposit feeders,Desmolaimus zeelandicus, Parodontophora sp., and the epistrate feederOncholaimus oxyuris were the most abundant in the deepest sediment, whereasNeotonchoides sp. andPseudochromadora sp. dominated the first 2 cm of sediments. Highest nematode density was recorded in the uppermost sediment layer (0–2 cm). Feeding types showed different abundance among transects and between months. There was a seasonal change in vertical distribution of nematodes, with the highest abundance in the deepest sediment layer in December, possibly due to the effect of wind waves on sediments of Chetumal Bay. The trophic composition of the nematode fauna in Chetumal Bay showed a dominance of deposit feeders and epistrate feeders, most likely in response to organic enrichment that is typical of eutrophic environments.  相似文献   

19.
The fossil echinoderms of the Lower Carboniferous (Mississippian) of Derbyshire remain understudied, principally due to the nature of the preservation rather than any lack of biodiversity. Echinoids and crinoids are described in float blocks of limestone from near Hurdlow, Derbyshire, which have been etched naturally after many years of being washed by weakly acidic rain. Surface detail is variable, commonly poor, but rare specimens retain enough features for tentative identification. Two species of echinoids are identified from rare disarticulated plates, namely Melonechinus? sp. (ambulacral plates) and archaeocidarid sp. (interambulacral plate); more and superior material will be necessary to confirm this division. Crinoids include a cladid brachial ossicle sp. with a distinctive sculpture; a columnal of a monobathrid camerate platycrinitid sp.; Annulocolumnus (col.) sp. cf. A. annulus Donovan, a columnal morphotaxon with an unusually broad axial canal; and Cyclocyclicus (col.) spp. This is the tenth echinoid site to be recognized from the Mississippian of the White Peak. Examination of etched float blocks provides a further method of investigation of the echinoderm fauna of this and other limestone areas.  相似文献   

20.
Spatial and temporal variations in the abundances and distributions of oligochaetes of a southwestern Louisiana estuary were examined as part of a long term study of community structure of benthic macroinvertebrates. Quantitative samples were collected at monthly intervals from nine stations for two years and an additional 17 stations were sampled once. A tubificid oligochaete,Tubificoides denouxi n. sp., is described from the five species collected. The two predominant oligochaetes,Tubificoides heterochaetus andT. denouxi, were congeneric and exhibited completely allopatric distributions. Two oligochaete species with-restricted distributions,Monopylephorus helobius andLimnodriloides sp., were sympatric withT. denouxi, whileThalassodrilides belli, although less abundant, was sympatric with bothT. denouxi andT. heterochaetus. Sexually mature specimens ofT. denouxi andT. belli were collected only in the summer,T. heterochaetus was sexually mature in both winter and summer collections, andMonopylephorus helobius was sexually mature in spring and summer collections. Many of the studies of Oligochaeta have concluded that correlation exists between sediment grain size and species demography. Our data demonstrate a strong relationship between salinity and the abundance and distribution of estuarine species.Tubificoides denouxi was found only within the salinity range of 14.8 to 22.0‰ salinity,T. heterochaetus was found only within the range of 2.3 to 14.1‰, andT. belli had a salinity distribution intermediate between the previous species. No relationship was found between sediment grain-size analysis, water depth or hydrographic variables and species distribution.  相似文献   

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