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1.
We describe a juvenile specimen of a titanosaurian sauropod dinosaur consisting of two dorsal and three caudal vertebral centra, an ilium fragment, and an ischium unearthed in 1991 from Site Km 153.5 at BR-050 highway in the Serra da Galga region, municipality of Uberaba, state of Minas Gerais, Brazil. The outcrop at the site is assigned to the Serra da Galga Member of the Marília Formation (Bauru Basin: Bauru Group; Upper Cretaceous: Maastrichtian). Although the material is very incomplete, features such as strongly procoelous caudal centra suggest an affinity with the titanosaurian clade Lithostrotia. The extensive vertebral pneumaticity with deep pleurocoels and well-developed camerae supports the hypothesis that, in titanosaurs, the air sac system was already present and fully developed even at early ontogenetic stages.  相似文献   

2.
Continental Cretaceous deposits exposed in central Patagonia, Argentina, preserve a rich and important record of titanosaurian evolution that spans the entire Upper Cretaceous. Recently, a new lithostragraphic unit, the Lago Colhué Huapi Formation (Coniacian – Maastrichtian), was located in the Golfo San Jorge Basin. Here, we describe new titanosaurian sauropod material from that formation (UNPSJB-Pv 1051). The material consists of a partially articulated left hind limb and disarticulated but associated skeletal elements. They are confidently referred to Titanosauria, and within that clade, to Lithostrotia, probably occupying a derived position. We interpret the bone concentration as an assemblage of hydraulic origin deposited in a fluvial channel. This new material begins the incipient fossil record of the Lago Colhué Huapi Formation, thereby also increasing the titanosaurian diversity of the latest Cretaceous. Additionally, the new titanosaurian enhances evidence in support of strong faunal similarities among the Allen and Marilia formations, chronological equivalents in the Cretaceous of South America. Furthermore, the Lago Colhué Huapi specimen adds to the Late Cretaceous record of Titanosauria and augments our knowledge of central Patagonian terrestrial vertebrate assemblages during this interval.  相似文献   

3.
记述了产自山东诸城上白垩统王氏群中一个较完整的蜥脚类恐龙肱骨(ZJZ-57)。它具有一些进步特征,如肱骨三角胸嵴向远端扩展,肱骨近端外侧1/3处发育隆凸,表明其属于较进步的巨龙类成员。另外,它具有一些独特的鉴定特征组合区别于其他任何巨龙类:近端横向强烈扩展,近端最大宽度与骨干长度之比为0.55;骨干粗壮,粗壮指数为0.39。综合研究表明,ZJZ-57所属个体代表了一种新的巨龙类恐龙:臧家庄诸城巨龙(Zhuchengtitan zangjiazhuangensis gen.et sp.nov.),它可能与Opisthocoelicaudia的亲缘关系较接近。这是山东晚白垩世地层中首次发现的巨龙类恐龙。  相似文献   

4.
《Cretaceous Research》2012,33(6):774-780
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

5.
Determining the relationships of mid-Late Cretaceous African taxa is central to understanding the timing and resultant palaeobiogeographical patterns of Gondwanan fragmentation. The early Late Cretaceous (Cenomanian) Kem Kem beds of southeastern Morocco preserve a diverse vertebrate fauna, including sauropod dinosaurs. Sauropod material includes the holotype of the rebbachisaurid diplodocoid Rebbachisaurus garasbae and fragmentary remains representing indeterminate titanosauriforms and rebbachisaurids. Here, we describe two new specimens from the Kem Kem beds. A dorsal neural arch with complex internal pneumaticity is tentatively attributed to a somphospondylan titanosauriform. A caudal vertebra possessing several rebbachisaurid synapomorphies is excavated by a large and pervasive lateral pneumatic foramen, a feature undocumented in other rebbachisaurids. However, caudal vertebrae are currently unknown for the sympatric R. garasbae, so this element could be referable to that taxon or a second, previously unknown, rebbachisaurid species. Interestingly, this new caudal vertebra displays a mosaic of features otherwise restricted to limaysaurine or nigersaurine rebbachisaurids, suggesting a placement basal to these clades, which is the position usually recovered for R. garasbae in phylogenetic analyses. A review of the mid-Cretaceous African sauropod fossil record removes the Cretaceous record of dicraeosaurids from Africa, restricting this clade to a single post-Jurassic occurrence in Argentina. All diagnostic sauropod remains can be attributed to titanosauriforms or rebbachisaurids. Whereas rebbachisaurids were seemingly restricted to northwestern Africa and disappeared post-Cenomanian, titanosauriforms were widespread across the African continent and survived until the latest Cretaceous. The development of the mid-Cretaceous Trans-Saharan Seaway might have acted as a dispersal barrier for rebbachisaurids and other vertebrate groups. In contrast, titanosauriforms might have been able to cross this barrier, but it is possible that they were also unable to disperse, and that northwestern African titanosauriforms were not closely related to taxa from the rest of the African continent. New materials and a better understanding of titanosaur interrelationships will be crucial in teasing these scenarios apart.  相似文献   

6.
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

7.
兰州—民和盆地位于中国西北横跨甘肃和青海两省的毗连地区,其中广泛发育出露良好下白垩统河口群。近年来,在该盆地东部的兰州盆地产出了一个前所未知的脊椎动物组合,其中以诸多的足印化石和世界已知最大牙齿的植食性恐龙——巨齿兰州龙(Lanzhousaurus magnidens)为代表,还包括一类宽臀的基干巨龙型类蜥脚类——刘家峡黄河巨龙(Huanghetitan liujiaxiaensis),以及许多其他尚未发表的恐龙、鱼类和龟类。本文报道在该地区河口群中所发现的第二种新的蜥脚类恐龙——炳灵大夏巨龙(新属新种)(Daxiatitan binglingi gen.et sp.nov.)。炳灵大夏巨龙的正型标本现存于甘肃省地矿局第三地质矿产勘查院古生物研究开发中心(兰州)。该标本包括最后的10枚颈椎,10枚背椎,2枚近端尾椎,部分颈肋和背肋,1枚脉弓,右肩胛骨,右乌喙骨和右股骨。估计其颈部长度为12.2m,整个身体长度为26m,使其成为中国乃至亚洲最大的恐龙之一。大夏巨龙是巨龙类的一个基干分子[其支干定义为:与盐龙(Saltasaurus)而不与腕龙(Brachiosaurus)更具亲缘关系的巨龙型类],其以如下特征区别于其他巨龙类分子:颈椎的前关节面后移;肩胛骨骨体的远缘与其长轴斜交;股骨远端髁相对于股骨体向背侧方倾斜10°,而且从远端看,该远端髁处于后内—前侧方向。大夏巨龙独特的股骨构造指示出一种强烈外撇的走步姿态。同时,这种姿态也得到了保存于刘家峡恐龙国家地质公园同一层位(河口群上段)的强烈外转的、宽间距的蜥脚类恐龙行迹的进一步佐证。炳灵大夏巨龙的发现丰富了兰州盆地脊椎动物组合,对研究巨龙类恐龙的起源和早期演化具有重要意义。  相似文献   

8.
Here we describe the first sauropod skeletal remains from the Italian peninsula that also represent the earliest record of titanosaurs in Southern Europe. Scattered bones, including an almost complete anterior caudal vertebra, were found in Cretaceous (Aptian–Albian) marine deposits, some 50 km East of Rome. The vertebra shows a bizarre and perhaps unique orientation of the zygapophyseal articular facets that renders their interpretation problematic. Phylogenetic retrofitting tests support the placement of the Italian titanosaur among basal lithostrotians. Palaeobiogeographic analysis based on the resulting phyletic relationships suggests an Afro-Eurasian route for the ancestors of the Italian titanosaur, a scenario compatible with the palaeogeographic evolution of the Italian microplates during the Cretaceous. Together with previously recorded titanosaurian-like ichnites from a Cenomanian locality in Latium, this new find suggests a quite long emersion for the Apenninic carbonate platform. We suggest that the Italian titanosaur was member of a population that crossed the western Tethys Sea through a “filtering bridge” composed of a chain of ephemeral islands and peninsulae, known as Periadriatic (Adria) carbonate platforms, that connected sporadically Africa and Europe since the Early Cretaceous.  相似文献   

9.
A New Titanosaurian Sauropod from Late Cretaceous of Nei Mongol, China   总被引:2,自引:0,他引:2  
A specimen collected from the Upper Cretaceous Erlian Formation of Nei Mongol (Inner Mongolia), China, represents a new genus and species of titanosaurian sauropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises several dorsal, sacral, and caudal vertebrae, several dorsal ribs, one anterior chevron, and much of the pelvis. Diagnostic features of the new species include a thick ridge extending down from the postzygapophysis on the lateral surface of the neural arch of the posterior dorsal vertebrae, a transversely oriented accessory lamina present between the anterior centroparapophyseal limina and the lateral centraprezygapophyseal lamina of the posterior dorsal vertebrae, long, anteroventrally directed caudal rib that bears two distinctive fossae on its posterior margin on the anterior caudal vertebrae and a prominent vertical ridge above the pubic peduncle on the medial surface of the ilium, among others. The ilium is pneumatic, a feature not common among non-avian dinosaurs. The new taxon has an unusual combination of primitive and derived character states. Preliminary character analysis shows a complex character distribution within the Titanosauriformes. Recent titanosauriform discoveries suggest that a significant radiation occurred in Asia early in the titanosauriform evolution.  相似文献   

10.
A new long-necked sauropod dinosaur, Yunmenglong ruyangensis gen. et sp. nov., is erected on the basis of an incomplete skeleton from the late Early Cretaceous Haoling Formation of the Ruyang Basin, Henan Province. The characters of the anterior cervical vertebrae, the shape of the neural canal of the dorsal vertebra and the ball-shaped distal end of the neural spine of caudal vertebrae with coarse surfaces differ from other long-necked sauropod dinosaurs. The new genus has characters in common with both Euhelopus and Erketu; it represents the first long-necked sauropod dinosaur recorded from central China to date. The rod-like, well-developed epipophyses and the pleurocoels on the cervical vertebrae indicate that it may be close to Euhelopus, an observation also confirmed by a phylogenetic analysis, which shows that Erketu, Yunmenglong and Qiaowanlong form a clade, and are more derived than Euhelopus.  相似文献   

11.
A new somphospondylan sauropod (Dinosauria, Titanosauriformes): Gannansaurus sinensis gen. et sp. nov. from the Late Cretaceous Nanxiong Formation of Ganzhou Basin, Jiangxi Province of southern China is erected based on one nearly complete dorsal vertebra and a mid-caudal vertebra. It is characterized by two centroprezygapophyseal lamina fossae; large, square-shaped concavity formed by centroprezygapophyseal lamina and dorsal margin of the centrum; large infraparapophyseal fossa; three openings present within the large lateral concavity, which occupy 65% length of the centrum; the posterior centroparapophyseal lamina (PCPL) and the anterior centroparapophyseal lamina (ACPL) weakly developed; the posterior centrodiapophyseal lamina (PCDL) separated by an infradiapophyseal fossa into two branches dorsally; parapophyseal and diapophyseal laminae cross to form "K" configuration. It shares some characters with the Early Cretaceous form-Euhelopus, indicating that it is more closely related to Euhelopus rather than to other titanosauriform sauropods. This means Gannansaurus may have a close phylogenetic relationship with Euhelopus.  相似文献   

12.
Doklady Earth Sciences - The previously unknown second cervical vertebra (axis) of the sauropod Sibirotitan astrosacralis Averianov et al., 2018 from the Early Cretaceous locality Shestakovo 1...  相似文献   

13.
记述了产自河南汝阳盆地早白垩世晚期相互关联的禽龙类9个中前部尾椎椎体化石。由于化石只保存不完整的尾椎部分,其属种难以断定。但它的发现说明,汝阳巨型蜥脚类恐龙动物群中,除了植食性的大型蜥脚类恐龙外,也存在大型鸟脚类恐龙——禽龙类。保存的尾椎椎体系列与发现于比利时的早白垩世贝尼萨尔禽龙相比,不论是大小还是形态构造上均极为相似,从而判断这9个连续的尾椎体分别为第7到第15尾椎。进而推断,河南汝阳盆地产出巨型蜥脚类恐龙动物群化石的沉积地层形成于早白垩世晚期。  相似文献   

14.
The upper Campanian-lower Maastrichtian site of Lo Hueco (Cuenca, Spain) has provided a set of well-preserved partial skeletons in anatomical connection or with a low dispersion of their skeletal elements. One partial skeleton is herein described and a new titanosaurian sauropod is established, Lohuecotitan pandafilandi. This titanosaur is diagnosed by eight autapomorphic features: dorsally and ventrally widened or bifurcated posterior centrodiapophyseal lamina in anterior and middle dorsal vertebrae; short postspinal lamina with a transversely expanded distal end represented by smooth scars in the dorsal vertebrae; anteriormost caudals with the medial spinoprezygapophyseal and medial spinopostzygapophyseal laminae ventrally connected with the prespinal and postspinal laminae, respectively; anterior caudal neural spines with a dorsal projection of the prespinal and postspinal laminae; anterior caudal neural spines bears a “greek-cross”-like cross-section; middle caudal centra having two round and rough structures in the dorsal edge of the posterior articulation, which extends to the dorsal surface of the centrum; the articular ends of the rami of the haemal arches are divided in two articular surfaces; and tuberosity between the anterior and the lateral trochanter of the fibula. The herein performed phylogenetic analysis considered L. pandafilandi as a member of Lithostrotia more derived than Malawisaurus. The known palaeodiversity of the Late Cretaceous Ibero-Armorican titanosaurs is increasing, and further analyses focused on this group will be necessary to better understand the evolutionary history of European titanosaurs and to clarify their relationships within Titanosauria.  相似文献   

15.
The first Podocarpaceae wood record is described from the Mulichinco Formation (Valanginian, Lower Cretaceous), Neuquén Basin, Argentina. The specimen was directly associated with a middle caudal vertebra of a diplodocid sauropod dinosaur. A new species – Podocarpoxylon prumnopityoides – is proposed based on features that include the presence of abietinean wood type (tracheid radial pitting), plus podocarpoid (cupressoid type) and some dacrydioid (taxodioid type) cross-field pits, diffuse axial parenchyma and low rays. This combination of anatomical characters is comparable to both Prumnopitys and Podocarpus, whereas the type of pits in the cross-fields resembles some members of the extant Prumnopitys. This is the first unequivocal record of the Family Podocarpaceae in the Valanginian of South America and confirms the hypothesis that the divergence between the “Podocarpoid-Dacrydioid” and “Prumnopityoid” clades occurred earlier than the Early Cretaceous.  相似文献   

16.
Bones in the fossil record sometimes exhibit unusual structures that can be attributed to pathologies, taphonomic alterations, or morphological variation. The goal of this work is to describe three bone abnormalities present in the type specimen of Bonitasaura salgadoi from the Cretaceous of north Patagonia, Argentina. The studied material corresponds to a left femur, a left metatarsal III, and a right prezygapophysis of a mid-caudal vertebra. Macroscopic and/or histological examinations were conducted, and a discussion concerning the origin of each abnormality is provided. The results of this study suggest that the anomalous structures are pathologies. In the femur, an osteoblastic tumor was identified by the presence of a large outgrowth of ovoid appearance with a spiculated microstructural pattern. The metatarsal III shows an enthesophyte (or bone spur) based on its location, shape and growth in parallel to the long axis of the bone element. Finally, the abnormal tissue observed in the prezygapophysis of the caudal vertebra was determined to be an infection by the presence of reactive new bone associated with a local widening of the subperiosteal margin and a sinus of drainage. This is the first report of multiple pathologies in a single specimen of a titanosaur, and it provides new insights about paleopathologies in sauropod dinosaurs.  相似文献   

17.
We describe new and redescribe previously discovered sauropod dinosaur material from the Upper Cretaceous (middle Cenomanian–Turonian) Bajo Barreal Formation of south-central Chubut Province, central Patagonia, Argentina. The remains consist of associated and isolated axial skeletal elements recovered from three separate localities, and are herein assigned to the morphologically aberrant Rebbachisauridae clade. Several of the fossils exhibit osteological characters that were previously undocumented in rebbachisaurids, enhancing our understanding of the morphological diversity of this enigmatic sauropod group. In particular, the Bajo Barreal material demonstrates the occurrence within Rebbachisauridae of bifurcate neural spines in cervical vertebrae and well-developed, presumably pneumatic fossae in caudal vertebrae; among Diplodocoidea, these distinctive morphologies had previously been recognized only in flagellicaudatans. Furthermore, the Bajo Barreal fossils constitute the southernmost known occurrences of Diplodocoidea, adding to the globally sparse Late Cretaceous record of these sauropods and augmenting our knowledge of central Patagonian terrestrial vertebrate assemblages during this interval.  相似文献   

18.
A new gigantic sauropod, Huabeisaurus allocates gen. et sp. nov., about 20 m in length and 5 m in height, was discovered in the Upper Cretaceous Huiquanpu Formation, Tianzhen County, Shanxi Province. It is notably different from Diplodocidae, Titanosauridae and Nemegtosauridae in the following aspects: the teeth are strong, peg-like with a length ratio of the tooth crown to tooth root at about 3 to 1; the cervical vertebrae are long with forked spines; the spines in dorsal vertebrae are relatively high, unbifiarcated; the caudal vertebrae are amphicoelous, with anterior neural spines and unbifurcated spines and chevrons; the femur is straight and long, narrow and flat and the tibia and fibula are long and flat. These characters show that the described genus should represent a new family, Huabeisauridae fam. nov. The discovery enriches the sauropod dinosaur record in China, and is quite significant to the study of the taxonomy, evolution, migration, extinction and palaeobiogeographic provincialism of the  相似文献   

19.
A new dinosaur sauropod from the Golmayo Formation (upper Hauterivian-lower Barremian) in Soria province (Spain) is described as Soriatitan golmayensis gen. et sp. nov. The new material consists of one tooth, three dorsal vertebrae with ribs, a partial sacrum, five caudal centra, two caudal vertebral spines, one chevron, a humerus, an ulna, a radius, two partial ilia, two ischia, a fragment of pubis, and a partial femur. Cladistic analysis identified the material as belonging to Brachiosaruidae within Titanosauriformes. Sauropod material from the Golmayo Formation shares a diagnostic feature with Abydosaurus, Cedarosaurus, Tastavinsaurus and Venenosaurus including anteriorly deflected anterior–middle caudal neural spines and with Cedarosaurus a proximodistally straight lateral margin between the proximal head and the shaft of the humerus. Eight characters are potential autapomorphies in the sauropod from Spain, including the presence of a large rectangular ventral ridge below the preacetabular process of the ilium, the same length of the pubic and ischiadic blades in the ischium, and two lateral ridges in the lateral area of the deltopectoral crest of the humerus. The presence of Early Cretaceous brachiosarids in both, North America and Europe, give support to the hypothesis of a connection between the tectonic plates of these continents at some point during the Early Cretaceous.  相似文献   

20.
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