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1.
In recent years the Late Cretaceous (Santonian) terrestrial vertebrate locality at Iharkút (western Hungary) has yielded well-preserved remains of lizard taxa besides the remains of fishes, amphibians, turtles, crocodiles, pterosaurs and dinosaurs. Previously the polyglyphanodontine lizard Bicuspidon aff. hatzegiensis has been reported from Iharkút. However, recent excavations at this site produced more lacertilian remains including new polyglyphanodontine material, namely a maxilla and two dentaries which suggest the presence of a new genus in the Iharkút fauna. This previously unknown lizard (described here as Distortodon rhomboideus n. g. n. sp.) is distinct from other polyglyphanodontines such as Bicuspidon, Paraglyphanodon, Polyglyphanodon, Dicothodon and Peneteius. It differs from these genera mainly in having the lingual cusp situated more distally compared to the labial one on its bicuspid teeth located in the distal part of the tooth row, thus the crowns having a unique rhomboidal shape in occlusal view. Distortodon rhomboideus further strengthens the dominance of borioteiioid lizards in the Iharkút fauna. The growing presence of borioteiioids in European localities supports previous theories which suggest some paleobiogeographic connections between the western Tethyan archipelago and North America in the Late Cretaceous. 相似文献
2.
Lepisosteid fishes are well known from the Upper Cretaceous of Europe, but only by fragmentary remains from some Cenomanian and Campanian–Maastrichtian deposits. Here we report various cranial and postcranial remains of gars, discovered in the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút (Bakony Mountains, Hungary). These remains represent one of the most diverse assemblages of lepisosteid fish material from Upper Cretaceous continental deposits of Europe. Based on tooth morphology, scale-microstructure and the features of the supracleithrum we refer these remains to the genus Atractosteus. Besides some uncertain remains from the Cenomanian of France and Spain, the Santonian aged fossils from Iharkút represent the oldest undisputable occurrence of the family Lepisosteidae in the European continental Cretaceous. Using tooth crown morphology, the surface microstructure of the ganoid scales and the anatomy of the supracleithrum a review of the Late Cretaceous lepisosteid record suggests the occurrence of both Atractosteus and Lepisosteus in the European archipelago. 相似文献
3.
A giant humerus (450 mm total length) belonging to one of the largest pterosaur recorded in South America is described. The specimen (UNCUYO-LD 350) was discovered in the Upper Cretaceous (upper Coniacian – lower Santonian) Plottier Formation of the Mendoza Province, northern Neuquén Basin, Argentina. It was found associated with a smaller pterosaur specimen represented by around thirty postcranial bones. The specimen is assigned to Tapejaroidea and show characters of both Tapejaridae and Azhdarchidae. Based on comparisions with other Azdharchidae species, a wingspan of 9.1 m is estimated for UNCUYO-LD 350, showing that giant pterosaurs were present in South America during the Upper Cretaceous. 相似文献
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5.
Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation (Bakony Mountains, western Hungary) are described. Macro- and microfossils collected from two boreholes and from isolated chunks of sediment/matrix dumped on spoil heaps of the Jókai Mine represent pycnodontiform and lepisosteiform fishes, bothremydid turtles, the mosasauroid Pannoniasaurus inexpectatus, the crocodyliforms cf. Theriosuchus, Iharkutosuchus makadii and cf. Allodaposuchus, as well as ankylosaurian and theropod dinosaurs. This unit was deposited in a swampy lacustrine environment, in contrast with the neighbouring and contemporaneous floodplain deposit of the vertebrate-bearing Csehbánya Formation at Iharkút. Despite significant environmental differences, the faunal composition of the Ajka Coal Formation assemblage completely overlaps with that of the Csehbánya Formation, suggesting the occurrence of the same semi-aquatic and terrestrial species in both settings. The ankylosaurian remains further strengthen the previous view that ankylosaurus preferred wetland habitats such as fluvial systems and coastal regions. 相似文献
6.
Abelisauroid dinosaurs normally reached an average body length (BL) of 5–9 m, but there are controversies due to the incomplete or fragmentary nature of most specimens. For Ekrixinatosaurus, for example, BL was estimated as 10–11 m or 7–8 m; for Pycnonemosaurus it was proposed 7–8 m, however its preserved bones are larger than any other described abelisauroid. The lack of a consistent methodology complicates comparisons of estimated BL, so we reevaluated the estimative for the seven most complete specimens of abelisauroids and compared the values against 40 measurements from the skull, vertebrae and appendicular elements using bivariate equations. It allowed estimating the BL of other 30, less complete, specimens of abelisauroids and to evaluate the allometric scaling of the skeletal parts. Strong correlations (R2 > 0.96) were obtained for all vertebrae and hindlimb measurements, as well as skull height, and length of skull roof, lacrimal–squamosal, scapulocoracoid and humerus; other skull and forelimb measurements present weak correlation due to extreme morphological transformations observed in Abelisauridae and were not adequate for BL estimation. Abelisauroids gradually increased in size during evolution: the mean BL was 3.3 ± 2.5 m for basal abelisauroids and Noasauridae, 5.4 ± 1.8 m for basal Brachyrostra and Majungasauridae, and 7.1 ± 2.1 m for Furileusaura. Despite this variation, diversity of BL on each geographic region and stratigraphic epoch was relatively constant (BL usually varied from 4 to 8 m). The smallest noasaurid and abelisaurid are, respectively, Velocisaurus (1.5 ± 0.1 m) and Genusaurus (3.6 ± 0.0 m). The largest abelisaurids is Pycnonemosaurus nevesi (8.9 ± 0.3 m) followed by Carnotaurus (7.8 ± 0.3 m), Abelisaurus (7.4 ± 0.7 m) and Ekrixinatosaurus (7.4 ± 0.8 m). Skull measurement scale negatively at a similar rate but the height scales almost isometrically and the skull roof length scales more negatively; this probably caused a bending on the skull that may explain the upward orientation of the snout in large taxa. 相似文献
7.
A weathered accumulation of turtle eggs, interpreted as remnants of a single clutch composed of at least 16 turtle eggs (MOR 710) from the Campanian (Upper Cretaceous) Judith River Formation of north-central Montana, USA, represents a new oospecies Testudoolithus zelenitskyae. This ootaxon is diagnosed by the following unique combination of characters: spherical eggs 34–39 mm in diameter, 660–760 μm thick eggshell, shell unit height-to-width ratio of 3.15:1–5.5:1, and domed shell units. Estimated egg mass indicates that the egg-laying adult likely possessed a carapace 35.0–54.4 cm in length. Similarities between T. zelenitskyae oosp. nov. and Adocus sp. eggs, along with comparable body size, suggest that this taxon might have produced MOR 710. One egg exhibits abnormal multilayered eggshell, likely resulting from prolonged egg retention by the female turtle. At least five of these eggs, including the multilayered specimen, preserve embryonic remains that demonstrate a late stage of embryonic development. This suggests that death occurred just prior to hatching. 相似文献
8.
A new ornithopod dinosaur from Antarctica, Trinisaura santamartaensis n. gen. et n. sp. is diagnosed by a unique combination of characters that includes a scapula with a spike-like acromial process with a strong and sharp lateral crest and longer than other ornithopods, a humerus with a rudimentary deltopectoral crest represented as a thickening on the anterolateral margin of the humerus, and shaft strongly bowed laterally, and an ischium gently curved along its entire length. The holotype specimen comprises vertebral and appendicular elements. The presence of axially elongate distal caudal vertebrae, pubis with long prepubic and postpubic processes, as well as a femur with a distinct anterior trochanter, pendant 4th trochanter and shallow anterior intercondylar groove constitute a combination of characters present in the Late Cretaceous Patagonian Gasparinisaura, Anabisetia and Talenkahuen. The materials were found on the surface enclosed in a hard sandstone concretion collected near the Santa Marta Cove, James Ross Island, from the lower levels of the Snow Hill Island Formation (Campanian). This is the first ornithopod taxon identified from this unit, and the second ornithischian dinosaur, after the ankylosaur Antarctopelta oliveroi. However, other ornithopod reports from nearby localities of James Ross and Vega islands in outcrops of the overlying Lopez de Bertodano Formation suggest that this clade was widely represented in the Campanian and Maastrichtian of the James Ross Basin, Antarctic continent. 相似文献
9.
We re-define the Cretaceous bony fish genus Rhinconichthys by re-describing the type species, R. taylori, and defining two new species; R. purgatorensis sp. nov. from the lowermost Carlile Shale (middle Turonian), southeastern Colorado, United States, and R. uyenoi sp. nov. from the Mikasa Formation (Cenomanian), Middle Yezo Group, Hokkaido, Japan. Rhinconichthys purgatoirensis sp. nov. is designated on a newly discovered specimen consisting of a nearly complete skull with pectoral elements. Only known previously by two Cenomanian age specimens from England and Japan, the North American specimen significantly extends the geographic and stratigraphic range of Rhinconichthys. The skull of Rhinconichthys is elongate, including an expansive gill basket, and estimated maximum body length ranges between 2.0 and 2.7 m. Rhinconichthys was likely an obligate suspension-feeder due to its derived cranial morphology, characterized by a remarkably large and elongate hyomandibula. The hyomandibula mechanically acts as a lever to thrust the jaw articulation and hyoid arch both ventrally and anterolaterally during protraction, thus creating a massive buccal space to maximize filtering of planktonic prey items. Cladistic analysis supports a monophyly of suspension-feeding pachycormids including Rhinconichthys, but further resolution within this clade will require more information through additional fossil specimens. 相似文献
10.
Two incomplete bones referred to ankylosaur scapulocoracoid and humerus are described, which were discovered from the Early Cretaceous Jiufotang Formation of Dachengzi Town, Kazuo County, Liaoning Province. Ankylosaur dinosaurs have ever been found in the Lower Cretaceous Yixian, Fuxin and Sunjiawan Formations of western Liaoning. The occurrence of ankylosaurs within Jiufotang Formation improves our knowledge on the ankylosaur stratigraphic distribution. Based on the length of scapulocoracoid, the new dinosaur is estimated to exceed 6.0 meters long, representing the largest ankylosaur in western Liaoning. 相似文献
11.
We present a large, fragmentary skull and the humerus of a mosasaur (Squamata, Mosasauroidea) recovered from upper Maastrichtian beds of the López de Bertodano Formation in Marambio (=Seymour) Island, Antarctica. The material belongs to a large, adult individual with marked heterodonty as well as unusual humeral features. Different phylogenetic analyses returned the studied specimen within the Tylosaurinae, while the distinctive features of the skull and humerus allow distinguish it from the unique Antarctic known tylosaurine species, Taniwhasaurus antarcticus (Novas et al., 2002), as well as from other known Late Cretaceous mosasaurids from the Southern Hemisphere, thus, justifying the erection of a new taxon, Kaikaifilu hervei gen. et. sp. nov. The different dental types documented in the specimen studied have been previously recorded through isolated teeth from the same locality and were subsequently referred to several genera. This new find is relevant for assessing the previously known fragmentary records of Antarctic mosasaurids, suggesting that its local diversity could be more reduced than previously interpreted. The new material represents the youngest occurrence of tylosaurines in Antarctica. 相似文献
12.
A new specimen of pterosaurian metacarpal IV from the Early Cretaceous (Berriasian) Durlston Formation of Dorset, southern England, closely resembles those of the Chinese dsungaripterid Dsungaripterus weii Young, 1964 and the Central Asian azhdarchid Azhdarcho lancicollis Nessov, 1984. However, the new specimen lacks the thickened bony wall typical of Dsungaripteridae and is therefore regarded as azhdarchoid. An mcIV with a markedly offset distal condyle with well-developed diaphyseal constriction behind the distal condyle and asymmetrical condylar margins may be an autapomorphy of Dsungaripteridae + Azhdarchoidea, while the same condition with bone thickening may typify Dsungaripteridae. The new specimen also provides osteological evidence supporting claims for large wing-spanned pterosaurs in the Berriasian of southern England based previously only on ichnological finds. The new specimen suggests a wingspan of between 2.9 and 3.7 m and represents the largest pterosaur from the Berriasian. 相似文献
13.
The paper presented is the first comprehensive, fully quantitative, high resolution study of marine palynology from an OAE3 black-shale environment. It is based on 175 m core spanning the upper Turonian to lower Santonian at Tarfaya, Morocco, NW Africa, which has been sampled from centimetre to 3 m intervals. The results are integrated and discussed with lithology and geochemistry data to (1) distinguish between potential changes in production and preservation of total organic carbon (TOC) accumulation and (2) constrain the stratigraphic position of the Oceanic Anoxic Event 3 (OAE3).The succession is characterized by increased total organic carbon (TOC), varying between 1% and 19% (average about 6%). Distinct black-shale horizons of variable thickness appear episodically throughout the succession, with higher frequency in the late Turonian. Higher TOC contents do not strictly correlate to lithologic black-shales or peaks of a specific taxon of organic-walled algae. The palynomorph spectrum is strongly dominated by organic-walled algae, with the ratio of terrigenous sporomorphs to organic-walled algae (t/m index) varying between zero and 0.05 (average 0.01). The dominance of algal organic matter is corroborated by the prevalence of Type I kerogen identified using Rock-Eval pyrolysis. Dinocyst diversity is low, with the absolute taxa number varying from 7 to 27 between single samples. The peridinioid/gonyaulacoid ratio of dinocysts (p/g ratio) shows strong fluctuations, varying between 1 and 283 (average of about 100).The upper Turonian interval is dominated by Bosedinia spp., a dinocyst taxon formerly described as abundant only in lacustrine sediments from the Oligocene and Miocene of SE Asia. This dominance is episodically modified by the increase of the warm-temperate waters dinocysts fraction, here mainly represented by the genera Alterbidinium, Isabelidinium and Spinidinium. Within the Coniacian-Santonian, black-shale horizons are limited in number and are concentrated within the upper Coniacian to lower Santonian interval. The dinocysts show alternating, prominent peak abundances of Palaeohystrichophora spp. and the warm-temperate water dinocysts fraction, here mainly represented by the genera Trithyrodinium and Chatangiella. However, a final episode of increased proportions of Bosedinia spp. is confined to a 5 m thick black-shale horizon closely spanning the Coniacian-Santonian boundary.Changes in the ratio of total sulphur to total organic carbon (TS/TOC) reflect fluctuating oxygen contents of bottom waters throughout the late Turonian to Santonian. These are significantly parallelled by the alternation of dinocysts assemblages suggestive of enhanced upwelling and water column stratification respectively, probably reflecting changes in the mode of TOC accumulation. Accordingly, preservation largely prevails during the late Turonian interval and changes towards increased production within the Coniacian-Santonian. However, a final preservation-event is probably represented by the black-shale horizon closely spanning the Coniacian-Santonian boundary (top Dicarinella concavata foraminifera zone), which may reflect an episodic shutdown of a major upwelling cell. It is thus proposed, that the “culmination” of the OAE3 at Tarfaya may represent intermittent preservation of TOC within an otherwise high productivity environment related to a global cooling trend. 相似文献
14.
In this study, we describe a new stratigraphy of three exposed sections in central Tunisia, integrating Coniacian and Santonian planktic foraminifera and calcareous nannoplankton, supported by ammonite and inoceramid bioevents. In the three sections, the Coniacian/Santonian (C/S) boundary lies slightly above the lowest occurrence (LO) of the calcareous nannofossil Lucianorhabdus cayeuxii, which marks nannofossil Zone CC16 and matches well with the LO of the planktic foraminifera Dicarinella asymetrica. It also lies ∼4–7 m below the LO of the inoceramid Platyceramus cycloides and the ammonite Texanites (Texanites) sp. Comparing these marker C/S bioevents with the global stratotype section, the Olazagutia section (Spain) shows that the stratigraphic range of the bioevents are variable. This observation must be taken into consideration when making regional chronostratigraphic correlations. 相似文献
15.
A new anurognathid pterosaur specimen from the Middle Jurassic Tiaojishan Formation of Qinglong, northern Hebie Province is described. The new specimen is referred to Dendrorhynchoides, based on the general morphology of the skeleton, but it represents a new species, named here as Dendrorhynchoides mutoudengensis sp. nov.. It is characterized by the presence of short, robust and straight teeth, and bearing wing metacarpal approximately 40% of the length of humerus. The new specimen provides further osteological information for anurognathid pterosaurs, especially the presence of a relatively elongate tail. 相似文献
16.
A new species of the foraminiferal genus Pseudorhapydionina is reported from the Santonian shallow-water carbonate and mixed deposits of La Cova Unit, in the Montsec and Pedraforca thrust sheets (Southern Pyrenees, NE Spain). Pseudorhapydionina bilottei sp. nov. differs from the Cenomanian species of the genus in its larger test size and the number of chambers in its early planispiral-involute stage. The discovery of P. bilottei sp. nov in the Santonian indicates that the genus Pseudorhapydionina escaped extinction during the Cenomanian-Turonian boundary (CTB) event in the Pyrenean bioprovince. A new subfamily Pseudorhapydionininae is described. 相似文献
17.
A well preserved skeleton of a new abelisaurid is reported here. The holotype of Viavenator exxoni was found in the outcrops of the Bajo de la Carpa Formation (Santonian, Upper Cretaceous), northwestern Patagonia, Argentina. This new taxon belongs to the South American clade of abelisaurids, the brachyrostrans. The current phylogenetic analysis places it as the basalmost member of a new clade of derived brachyrostrans, named Furileusauria, characterized by distinctive cranial, axial and appendicular anatomical features. The Santonian age of Viavenator allows filling the stratigraphic gap exhibited between the basal brachyrostrans of Cenomanian–Turonian age, and the derived forms from the Campanian-Maastrichtian. The evolution of abelisaurids during the Late Cretaceous, faunal replacements, and the adaptive radiation that occurred during that period of time in South America are discussed. 相似文献
18.
《Proceedings of the Geologists' Association. Geologists' Association》2023,134(3):361-373
Four isolated cervical vertebrae from the Kimmeridge Clay Formation (Upper Jurassic, Kimmeridgian) of Abingdon, Oxfordshire, England are identified as from a pliosaurid plesiosaurian sauropterygian on account of their shortness relative to width and height, their near platycoelous nature and the location of tall rib facets on the centrum body. They are noteworthy for their size, with a maximum width of 269 mm, maximum height of 222 mm and maximum length of 103 mm. Simple scaling and comparisons with cervical vertebrae of Mid Jurassic pliosaurs Peloneustes and Liopleurodon, and the Early Cretaceous Stenorhynchosaurus and Sachicasaurus suggest a total body length of between ~ 9.8 m and 14.4 m for the Abingdon Kimmeridge Clay pliosaur. Likely the true length was towards the higher end of this range.A genus and species cannot be confidently determined on the basis of the described material, but they likely belong to Pliosaurus sp. or a similar animal, for which a precise neck length is not known. We estimate a neck length of 0.77 m for Pliosaurus ?brachyspondylus based on the average cervical lengths provided for specimen CAMSM J.35991. 相似文献
19.
The shallow-marine, mixed siliciclastic-calcareous Late Cretaceous deposits from the Apuseni Mountains have been extensively studied and compared to coeval deposits from the Alpine Gosau. The former are mainly represented by conglomerates, sandstones, marls, and limestones with rudists that unconformably overlie the crystalline basement and its Permo-Mesozoic cover. Our new, detailed investigations on the rudist fauna from Măgura Hill, the type locality of Pseudopolyconites hirsutus (Patrulius) and Miseia costulata Patrulius, indicate a Late Santonian–Early Campanian age for these deposits instead of an Early Santonian one as previously suggested (Patrulius, 1974). This study also mentions for the first time the occurrences of Pseudosabinia klinghardti (Böhm) and Pseudopolyconites parvus Milovanović in the rudist-bearing deposits from the Apuseni Mountains. We include their palaeontological features, as well as the ones for Pseudopolyconites hirsutus. Based on new biostratigraphic data, our study expand the stratigraphic range of Pseudosabinia klinghardti and Pseudopolyconites parvus – previously considered characteristic for the Early Campanian–Maastrichtian interval. Also we add new information on their palaeobiogeographic distribution within the central-eastern Mediterranean area during the Late Cretaceous. 相似文献
20.
Cretornis hlavaci Frič, 1881 from the Upper Cretaceous of Czech Republic (Pterosauria,Azhdarchoidea)
Cretornis hlavaci Frič, 1881 from the Upper Cretaceous (Turonian) of Czech Republic is a valid taxon referred to Azhdarchoidea based on having a saddle-shaped humeral head, pneumatic foramen on proximal humerus present on anterior side and absent on posterior side, elongate deltopectoral crest with subparallel proximal and distal margins, pneumatic foramen absent on distal side of humerus, metacarpals I–III not articulated with carpus and displaced on anterodorsal side of wing metacarpal, and wing metacarpal much longer than humerus. Absence of a pneumatic foramen on posterior side of proximal humerus suggests attribution of Cretornis hlavaci to Neoazhdarchia. It has a unique construction of the distal ulna with a dorsal articulation surface placed distinctly proximal to the tuberculum shared only with the non-azhdarchid azhdarchoid Montanazhdarcho minor from the Campanian of North America. Cretornis hlavaci differs from the latter taxon by the structure of its humerus and distinctly longer wing metacarpal. It is more derived than “Tapejaridae” but shares with Azhdarchidae the deltopectoral crest of the humerus displaced distally from humeral head. Cretornis cannot be assigned to Azhdarchidae because of the oval cross section of the second wing phalanx. A unique rhombic outline of the distal humerus of Cretornis hlavaci is a possible autapomorphy for this taxon. Its wing span estimated as 1.5–1.6 m. This is the first taxon of non-azhdarchid pterosaurs known from the Upper Cretaceous of the Eastern Hemisphere. 相似文献