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1.
Patagonia exhibits a particularly abundant record of Cretaceous dinosaurs with worldwide relevance. Although paleontological studies are relatively numerous, few include taphonomic information about these faunas. This contribution provides the first detailed sedimentological and taphonomical analyses of a dinosaur bone quarry from northern Neuquén Basin. At Arroyo Seco (Mendoza Province, Argentina), a large parautochthonous/autochthonous accumulation of articulated and disarticulated bones that represent several sauropod individuals has been discovered. The fossil remains, assigned to Mendozasaurus neguyelap González Riga, correspond to a large (18–27-m long) sauropod titanosaur collected in the strata of the Río Neuquén Subgroup (late Turoronian–late Coniacian). A taphonomic viewpoint recognizes a two-fold division into biostratinomic and fossil-diagenetic processes. Biostratinomic processes include (1) subaerial biodegradation of sauropod carcasses on well-drained floodplains, (2) partial or total skeletal disarticulation, (3) reorientation of bones by sporadic overbank flows, and (4) subaerial weathering. Fossil-diagenetic processes include (1) plastic deformation of bones, (2) initial permineralization with hematite, (3) fracturing and brittle deformation due to lithostatic pressure; (4) secondary permineralization with calcite in vascular canals and fractures, and (5) postfossilization bone weathering. This type of bone concentration, also present in Rincón de los Sauces (northern Patagonia), suggests that overbank facies tended to accumulate large titanosaur bones. This taphonomic mode, referred to as “overbank bone assemblages”, outlines the potential of crevasse splay facies as important sources of paleontological data in Cretaceous meandering fluvial systems.  相似文献   

2.
The paper describes an early and primitive sauropod dinosaur, Gongxianosaurus shibeiensis (gen. et sp. nov.), from the Dongyuemiao Member of the Lower Jurassic Ziliujing Formation in Shibei Township, Gongxian County, Sichuan Province, China, which is one of the 5 dinosaur fossils discovered in Gongxian in 1997. Except the skull which is incomplete, the fossils are well preserved. It has some features of both sauropods and prosauropods. It is an intermediate type in the evolution of dinosaurs from prosauropods to sauropods and provides materials for the study of the origin and evolution of the sauropod dinosaur fauna. The discovery of this new sauropod furnishes a way for the stratigraphic correlation between the Early Jurassic Ziliujing Formation in the Sichuan basin and the Lower Jurassic Lufeng Formation in the Central Yunnan basin.  相似文献   

3.
There are 24 known localities for skeletal remains of sauropod dinosaurs in the republics of Central Asia (Kazakhstan, Kyrgyzstan, Tajikistan, and Uzbekistan). Sauropod remains are very rare at all these localities and represented usually only by isolated teeth. Only narrow-crowned teeth are known from the Cretaceous of Central Asia. The oldest record of such teeth is from the Aptian Sultanbobo Formation of Uzbekistan. Exposures of the Turonian Bissekty Formation at the most productive vertebrate locality in the region, Dzharakuduk in Uzbekistan, has yielded many isolated teeth and a few skeletal remains that can be attributed to a non-lithostrotian titanosaur. Similar narrow-crowned, cylindrical teeth from Cenomanian-to Coniacian-age strata in the Kyzylkum Desert, Uzbekistan, may belong to a closely related taxon. Another taxon, with teeth that are pentagonal in cross-section, is known from the Santonian Yalovach and Bostobe formations of Tajikistan and western Kazakhstan, respectively. A femur reported from the Santonian Syuksyuk Formation of southern Kazakhstan possibly belongs to a lithostrotian titanosaur. The change in tooth structure at the Coniacian–Santonian boundary in the region possibly suggests replacement of non-lithostrotian titanosaurs by lithostrotians. The titanosaur from the Bissekty Formation is similar to Dongyangosaurus sinensis from the Cenomanian–Turonian of Zhejiang (China) in the extensive pneumatization of the neural arch on the anterior caudal vertebrae with several fossae. It also resembles Baotianmansaurus henanensis from the Cenomanian of Henan (China) in the possession of very short anterior caudal centra. These three taxa possibly represent an as yet formally unrecognized endemic clade of Asian non-lithostrotian titanosaurs.  相似文献   

4.
Determining the relationships of mid-Late Cretaceous African taxa is central to understanding the timing and resultant palaeobiogeographical patterns of Gondwanan fragmentation. The early Late Cretaceous (Cenomanian) Kem Kem beds of southeastern Morocco preserve a diverse vertebrate fauna, including sauropod dinosaurs. Sauropod material includes the holotype of the rebbachisaurid diplodocoid Rebbachisaurus garasbae and fragmentary remains representing indeterminate titanosauriforms and rebbachisaurids. Here, we describe two new specimens from the Kem Kem beds. A dorsal neural arch with complex internal pneumaticity is tentatively attributed to a somphospondylan titanosauriform. A caudal vertebra possessing several rebbachisaurid synapomorphies is excavated by a large and pervasive lateral pneumatic foramen, a feature undocumented in other rebbachisaurids. However, caudal vertebrae are currently unknown for the sympatric R. garasbae, so this element could be referable to that taxon or a second, previously unknown, rebbachisaurid species. Interestingly, this new caudal vertebra displays a mosaic of features otherwise restricted to limaysaurine or nigersaurine rebbachisaurids, suggesting a placement basal to these clades, which is the position usually recovered for R. garasbae in phylogenetic analyses. A review of the mid-Cretaceous African sauropod fossil record removes the Cretaceous record of dicraeosaurids from Africa, restricting this clade to a single post-Jurassic occurrence in Argentina. All diagnostic sauropod remains can be attributed to titanosauriforms or rebbachisaurids. Whereas rebbachisaurids were seemingly restricted to northwestern Africa and disappeared post-Cenomanian, titanosauriforms were widespread across the African continent and survived until the latest Cretaceous. The development of the mid-Cretaceous Trans-Saharan Seaway might have acted as a dispersal barrier for rebbachisaurids and other vertebrate groups. In contrast, titanosauriforms might have been able to cross this barrier, but it is possible that they were also unable to disperse, and that northwestern African titanosauriforms were not closely related to taxa from the rest of the African continent. New materials and a better understanding of titanosaur interrelationships will be crucial in teasing these scenarios apart.  相似文献   

5.
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

6.
The oldest dinosaur assemblages of Thailand deposit in continental sedimentary rocks of the Nam Phong Formation. Not only Isanosaurus attavipatchi was discovered but at least two more species of basal sauropods were found. A partial skeleton from Phu Hin Tan locality refers to sauropod taxon A. It is different from Isanosaurus and shares some characteristics with basal sauropods in the Early Jurassic. The sauropod taxon B was found in Non Sra Ard locality. Base on postcranial skeleton, sauropod taxon B shares some characteristics of the family Vulcanodontidae. Moreover, several fragments of the sauropods specimen are also found in Pha Khok Wang Yang and Phu Noi localities. All of the evidences above indicate that there are a highly diversity of sauropods in the Nam Phong Formation. At least three sauropod species (including Isanosaurus) were found in the Nam Phong formation. The issue about the age of the Nam Phong has been debated for a long time between the Triassic and Jurassic age. All of sauropodomorphs, in this study, are more likely the Early Jurassic period dinosaurs than the Triassic period dinosaurs. And they have deposited in the upper part of Nam Phong Formation.  相似文献   

7.
A new long-necked sauropod dinosaur, Yunmenglong ruyangensis gen. et sp. nov., is erected on the basis of an incomplete skeleton from the late Early Cretaceous Haoling Formation of the Ruyang Basin, Henan Province. The characters of the anterior cervical vertebrae, the shape of the neural canal of the dorsal vertebra and the ball-shaped distal end of the neural spine of caudal vertebrae with coarse surfaces differ from other long-necked sauropod dinosaurs. The new genus has characters in common with both Euhelopus and Erketu; it represents the first long-necked sauropod dinosaur recorded from central China to date. The rod-like, well-developed epipophyses and the pleurocoels on the cervical vertebrae indicate that it may be close to Euhelopus, an observation also confirmed by a phylogenetic analysis, which shows that Erketu, Yunmenglong and Qiaowanlong form a clade, and are more derived than Euhelopus.  相似文献   

8.
9.
A new species of sauropod dinosaur Huanghetitan ruyangensis is erected based on the following characters: deepest body cavity with a dorsal rib reaching at least 2.93 m long, anterior caudal vertebrae with mushroom-shaped neural spines. Based on this new specimen of Huanghetitan found in the early Late Cretaceous Mangchuan Formation of Ruyang, Henan Province, the family Huanghetitanidae faro. nov. is proposed as a new rank to include only the genus Huanghetitan You et al. 2006. At present, Huanghetitan includes two species: H. liujiaxiaensis You et al., 2006 and H. ruyangensis sp. nov. The systematic relationships of Huanghetitan among sauropod dinosaurs are briefly discussed.  相似文献   

10.
《Cretaceous Research》2012,33(6):774-780
Rebbachisauridae is a poorly understood clade of diplodocoid sauropod dinosaurs, currently known only from the Cretaceous of Africa, Europe and South America. European representatives are particularly rare and fragmentary. Here, we report an anterior caudal vertebra from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, off the southern coast of England. This specimen possesses several features known only in rebbachisaurids and shares two synapomorphies with the Afro-European taxa Demandasaurus darwini and Nigersaurus taqueti, both pertaining to the morphology of the neural spine. These features are the development of triangular lateral processes and the presence of an elliptical fossa on the lateral surface, bounded by the lateral lamina and postspinal rugosity. The Isle of Wight specimen also shares several features solely with Demandasaurus, indicating a close relationship with the Spanish taxon. These include the presence of a hyposphenal ridge, as well as an anteriorly excavated caudal rib that is restricted almost entirely to the neural arch. However, it differs from Demandasaurus in a number of ways, including the lack of excavation on the posterior surface of the caudal rib, the orientation of the neural spine, and the composition and morphology of the lateral lamina. In addition, the Isle of Wight vertebra possesses one potential autapomorphy: bifurcation of the elliptical fossa on the neural spine. However, because of the fragmentary nature of the material, a new name is not erected. Along with Demandasaurus and Histriasaurus boscarollii, this caudal vertebra indicates the presence of at least three European rebbachisaurid taxa and provides new anatomical information on this enigmatic clade of sauropod dinosaurs.  相似文献   

11.
A new genus and species of an Aptian–Albian sauropod from the Itapecuru Formation, northern Brazil, Amazonsaurus maranhensis, is described. It is known from an incomplete, but diagnostic postcranial skeleton. The new taxon possesses several autapomorphies, such as the anterior caudal vertebrae with lateral laminae formed by the coalescence of the spinoprezygapophyseal and postzygodiapophyseal laminae and, to a lesser extent, of the postzygodiapophyseal laminae. It exhibits many synapomorphies supporting its inclusion in the Diplodocoidea. These include high caudal neural arches and anterior caudal neural arches with spinoprezygapophyseal laminae on the lateral aspect of the neural spine. This record is consistent with previous hypotheses on the existence of a community of Afro-South American dinosaurs.  相似文献   

12.
Here we describe a new record of a sauropod dinosaur from the Lower Cretaceous (Hauterivian–Barremian) Rio Piranhas Formation, Sousa Basin, NE Brazil. Dinosaur fossil bones from this deposit were unknown until now. Thus, the discovery of a sauropod fibula from this locality is highly significant. Our discovery represents an indeterminate titanosaur and the earliest stratigraphic occurrence of this group in central Gondwana. When compared to chronocorrelate titanosaur trackmakers of this geological unit, this fossil specimen appears substantially smaller. Histological analysis of the fibula suggests that this is a relatively young individual (approximately 40–50% adult body size) that had passed its most rapid phase of early juvenile growth, but had not yet attained somatic maturity. Thus, the fibula recovered is from a young individual rather than from a small-bodied adult titanosaur.  相似文献   

13.
Eighty two dinosaur trackways were newly discovered in Upper Cretaceous lacustrine deposits on islands in the vicinity of Yeosu, Korea. Most dinosaur tracks occur in marginal lake deposits with polygonal desiccation cracks. The dinosaur tracks at the Yeosu site include 65 ornithopod trackways, 16 theropod trackways and one sauropod trackway. The prevalence of ornithopod tracks and the limited occurrence of sauropod tracks at the Yeosu site evidently reflect decreased sauropod diversity in the Upper Cretaceous. All ornithopod trackways represent bipeds, and most of the ornithopod tracks are similar to Caririchnium from other sites of the Korean peninsula. All fossil wood specimens collected in the study area represent conifers (three species of cupressaceous and two species of taxodiaceous conifers, and a new species) except for one, which is a discotyledon. It is thus inferred that the southwestern part of the Korean Peninsula was primarily covered with mesic forests with taxodiaceous trees during the Late Cretaceous. The K–Ar age of the Yeosu tracksite is determined as 81–65 Ma (Camapnian to Maastrichtian). It indicates that the Yeosu track site contains the last records of dinosaurs living in Asia. Consequently, semi-arid palaeoclimatic conditions, together with a large lake as a persistent water source and rich vegetation of gymnosperm trees as food, resulted in the preservation of abundant dinosaur tracks in the Upper Cretaceous on the Korean Peninsula.  相似文献   

14.
The Fox-Amphoux Syncline of the Aix Basin (SE France) has yielded a diverse Late Cretaceous vertebrate assemblage, including several taxa of dinosaurs. Here, we report on cranial material of titanosaurian sauropods, which consist of a partial braincase, two fragmentary skull roofs, and nine teeth, from the Métisson locality (Var Department). The braincase differs from those found previously in Europe (i.e., Ampelosaurus atacis, Lirainosaurus astibiae, and an unnamed juvenile skull from Romania) as well as from other titanosaurian braincases in having a groove that extends along the ventral surface of the occipital condyle neck (this feature may be autapomorphic). One of the fragmentary skull roofs may belong to the same taxon, whereas the other suggests the presence of a second titanosaur at Métisson very close to Ampelosaurus. Two dental morphologies are present in the sample; one of them includes teeth of different sizes. We suggest that this could be accounted for by age differences within a single taxon, or be due to different positions in the tooth row. The presence of a new, still unnamed titanosaurian taxon in the Ibero-Armorican Island supports previous works indicating a high titanosaurian diversity during the Campanian-Maastrichtian in southern Europe.  相似文献   

15.
A study of the abundant and undescribed isolated and associated bones and teeth from the La Amarga Formation (Barremian of Neuquén, Argentina) permitted the recognition of additional clades of sauropod dinosaurs: basal titanosauriforms, both basal and derived titanosaurs, and rebbachisauroid diplodocoids, which are now added to the already known dicraeosaurids and a recently published basal diplodocoid. These forms substantially increase the knowledge on the Early Cretaceous sauropod diversity in Gondwana.  相似文献   

16.
Dental histology of periodontal tissues (cementum, periodontal ligament and alveolar bone) has been studied in mammals, crocodylians and some basal tetrapods, but these structures have never been studied in titanosaur sauropods. The goal of this work was to study the structures of dental insertion in Titanosaurs. Like many physiological processes, histological analysis of titanosaur teeth shows hard tissue formation, characterized by a circadian rhythm. From thin sections it was possible to observe microstructures such as incremental lines of von Ebner, dentinal tubules and cross striations, all key to the understanding of developmental tooth dynamics. The structural and histological analyses carried out here on teeth of Late Cretaceous titanosaurs reveals the presence of acellular and cellular cementum, periodontal ligament, and alveolar bone, all structures necessary for a truly thecodont dentition. This is the first time documented for a dinosaur via histological tissue, and is an important finding that will help elucidate aspects of dinosaurian dentition, tooth replacement rate, feeding strategy, metabolism, and general biology.  相似文献   

17.
18.
Isolated left prefrontal, left squamosal and atlas of titanosaur dinosaurs are described and compared. They come from the Late Cretaceous Serra da Galga Member of the Marília Formation at the Serra do Veadinho region, Peirópolis (Uberaba County, Minas Gerais State, Brazil). Due to the sparse cranial elements of titanosaurs already known from Brazil, these specimens are noticeable to be presented. In addition, the atlas vertebra is described for the first time for Brazilian titanosaurs. The morphology of the cranial bones closely resembles lithostratian titanosaurs, such as Rapetosaurus, rather than basal titanosaurs. The atlas is similar to that of other titanosaurs, suggesting that the anatomy of this element seems to be more conservative than other vertebral elements, in which vertebral laminae play an important rule in titanosaur taxonomy.  相似文献   

19.
记河南汝阳白垩纪一新的巨型晰脚类恐龙   总被引:1,自引:0,他引:1  
记述了产自河南汝阳地区晚白垩世早期莽川组一新的巨型蜥脚类恐龙--巨型汝阳龙(新属新种):Ruyangosaurus giganteus gen.et sp.nov..它具有以下特征:神经棘低,缺少椎前关节突隔板,神经弓的侧面具有大的、不规则的三角形凹,前关节横突隔板前后伸展,胫骨粗壮,达127 cm等.汝阳龙的发现显示在晚白垩世早期蜥脚类恐龙发生的分异程度要比以前想象的高得多.  相似文献   

20.
Bones in the fossil record sometimes exhibit unusual structures that can be attributed to pathologies, taphonomic alterations, or morphological variation. The goal of this work is to describe three bone abnormalities present in the type specimen of Bonitasaura salgadoi from the Cretaceous of north Patagonia, Argentina. The studied material corresponds to a left femur, a left metatarsal III, and a right prezygapophysis of a mid-caudal vertebra. Macroscopic and/or histological examinations were conducted, and a discussion concerning the origin of each abnormality is provided. The results of this study suggest that the anomalous structures are pathologies. In the femur, an osteoblastic tumor was identified by the presence of a large outgrowth of ovoid appearance with a spiculated microstructural pattern. The metatarsal III shows an enthesophyte (or bone spur) based on its location, shape and growth in parallel to the long axis of the bone element. Finally, the abnormal tissue observed in the prezygapophysis of the caudal vertebra was determined to be an infection by the presence of reactive new bone associated with a local widening of the subperiosteal margin and a sinus of drainage. This is the first report of multiple pathologies in a single specimen of a titanosaur, and it provides new insights about paleopathologies in sauropod dinosaurs.  相似文献   

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