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1.
埃迪卡拉纪—寒武纪转换时期小壳化石的突然出现是世人瞩目的重要演化事件之一,它见证了动物的爆发式快速演化过程,标志着以两侧对称动物为特征的寒武纪生态系统的开始。湖北三峡地区寒武系纽芬兰统岩家河组处于幸运阶—第二阶这一关键层段,是研究上述问题的理想地区之一。岩家河组小壳化石研究最近取得的进展,为寒武系纽芬兰统幸运阶/第二阶界线的标定,和寒武纪早期软体动物和刺胞动物的演化及个体发育提供了重要的化石依据,本文对其进行了总结。全球界线层型剖面和点位(GSSP)寒武系第二阶底界划分标志的Watsonella crosbyi和Aldanella attleborensis在岩家河组第5段底部的同时出现,意味着岩家河组第5段已归属寒武系第二阶。这两类软体动物化石均广泛分布于华南、蒙古、西伯利亚和北美等大陆碳酸盐岩相寒武系第二阶地层中,它们均是定义寒武系第二阶底界的有力竞争者。另外,W.crosbyi壳顶下方一对肌肉附着结构的发现提供了重要的软组织结构信息,证实了W.crosbyi为内腹型壳,归属为未扭曲的软体动物太阳女神螺类。新发现六方锥石类Septuconularia yanjiaheensis具侧向压扁、两辐射对称的、14个等大锥面的围鞘,是迄今所描述的六方锥石类中锥面数最多的类型。根据横肋疏密程度,围鞘由下向上可分为三部分,可能代表胚胎期、幼年期和成年期不同的生长阶段。Septuconularia显然由寒武系幸运阶Hexaconularia通过扇面扩增演变而来,证明Hexaconularia可能是Arthrochites和Septuconularia之间的中间过渡类型。新发现的橄榄蛋类Octapyrgites elongatus与幸运阶Olivooides和Quadrapyrgites十分相似,围鞘由方形的壳顶区和波纹状倒宝塔形的远顶区组成。然而Octapyrgites只有8个口折叶,有别于具12个口折叶的Quadrapyrgites。通过与幸运期水母类多种对称方式对比,发现在寒武系第二阶橄榄蛋类数量减少和五辐射对称骨状壳类缺失,表明在幸运期/第二期转换时期,底栖固着型的刺胞动物水母类多样性发生了明显衰落,甚至部分绝灭,而同期的两侧对称动物则迅速辐射演化。 相似文献
2.
扰动生物作为生态系统工程建造者,通过对沉积底质的改造直接影响沉积物和水体之间的相互作用（营养物质的循环、迁移和存储）,对底栖生态系统工程有重要的影响.遗迹化石作为底栖生物活动的载体,为研究寒武纪大爆发之后底栖生态系统工程建造者的演化规律,并评价底栖生态系统工程建造者对生态系统工程产生的影响提供了依据.对河南地区8个寒武系剖面的遗迹化石数据进行统计,应用现代生物学中生态系统工程影响值的概念,从功能群、阶层、生物灌溉3个方面对底栖生态系统工程建造者进行研究.研究结果表明,寒武系第三阶至鼓山阶底栖生态系统工程建造者功能群类型由生物扩散型扰动者往向上/下运输者转变,阶层逐渐加深,生物灌溉越来越普及,生态系统工程影响值不断增大.鼓山阶至第十阶,遗迹化石类型及结构变化不大,生态系统工程影响值趋于稳定.这表明从寒武纪生物大爆发至鼓山阶,底栖生态系统工程建造者的行为策略发生着明显的进化与创新,生态系统工程建造者不仅开拓了重要的生态空间,而且为其他底栖生物和微生物创造了新的生境,在寒武纪海洋生态系统中扮演着重要的角色. 相似文献
3.
近年来随着“鄂宜页1井”高产气井的突破,寒武系水井沱组(第2统第3阶)已成为中扬子区页岩气勘查开发新的主力层系。本文旨在前人研究的基础上,通过对水井沱组中主要生物类型、古生物组合面貌及其时空分布的简要总结,为鄂西地区的页岩气勘探提供对比借鉴。宜昌地区寒武纪地层发育完整,广泛出露于宜昌、秭归、长阳、兴山等地,古生物化石较为丰富,是开展寒武纪地层及古生物研究的经典地区之一。埃迪卡拉纪—寒武纪过渡时期的地质记录中保存了寒武纪大爆发和后生动物多样化事件的化石证据,从寒武系纽芬兰统到第二统,自下而上发现了岩家河生物群、水井沱生物群(包括清江生物群)、石牌生物群和京山布尔吉斯页岩型生物群等,它们化石类群多样,浮游生物和底栖生物丰富,而在长阳发现的清江生物群更是具有典型布尔吉斯页岩型动物群特异埋藏化石库的特点,出现了大量高保真度的软躯体类群,代表了远离海岸的较深水环境,为寒武纪大爆发研究打开了一个新的窗口。从岩性、生物类型以及构造运动来看,宜昌地区水井沱组发育一套以黑色页岩或钙质页岩夹碳质灰岩为主的黑色岩系,其中海绵化石在黑色岩系中呈现出较高的丰度和多样性,显示当时深水低氧的沉积环境。同时繁盛的微体... 相似文献
4.
华南伊迪卡拉纪"庙河生物群"的属性分析 总被引：4，自引：0，他引：4
我国南方伊迪卡拉(震旦)系陡山沱组页岩中,保存了以"庙河生物群"为代表的宏体碳质压膜化石群,以大量底栖固着的多细胞藻类为主,还包含推测的后生动物和海绵动物、可疑的遗迹化石等多门类生物化石,化石类型多样、分异显著,代表着新元古代"雪球"冰期之后和寒武纪早期后生动物大爆发前夕地球早期多细胞生物的一次大规模的进化辐射事件。本文通过比较湖北秭归庙河和贵州江口翁会两个典型产地的宏体化石优势属种的形态特征,以及两地所处的沉积古环境,分析探讨了这些优势类别与现生多细胞生物(包括红藻、褐藻和绿藻三大高级藻类)的亲缘关系。研究表明,寒武纪之前大约35~10Ma间,各门类多细胞藻类广泛发育,与后生动物的先驱分子构成独特的古生态群落。 相似文献
5.
我国南方伊迪卡拉（震旦）系陡山沱组页岩中,保存了以“庙河生物群”为代表的宏体碳质压膜化石群,以大量底栖固着的多细胞藻类为主,还包含推测的后生动物和海绵动物、可疑的遗迹化石等多门类生物化石,化石类型多样、分异显著,代表着新元古代“雪球”冰期之后和寒武纪早期后生动物大爆发前夕地球早期多细胞生物的一次大规模的进化辐射事件。本文通过比较湖北秭归庙河和贵州江口翁会两个典型产地的宏体化石优势属种的形态特征,以及两地所处的沉积古环境,分析探讨了这些优势类别与现生多细胞生物（包括红藻、褐藻和绿藻三大高级藻类）的亲缘关系。研究表明,寒武纪之前大约35～10 Ma间,各门类多细胞藻类广泛发育,与后生动物的先驱分子构成独特的古生态群落。 相似文献
6.
湘西震旦纪武陵山生物群的化石形态学特征和归属 总被引：2，自引：0，他引：2
湘西桃源理公港震旦系留茶坡组中部黑色页岩中的炭质压膜化石,为研究末前寒武纪晚期生物学提供了一个稀有的布吉斯页岩型分类学窗口。根据已知属种和新获材料的研究,确定了17个明显的形态类型,讨论了每种形态类型特征及其可能的生物学属性,指出除个别化石可能为两侧对称动物外,其余绝大多数化石可以解释为原核生物或多细胞藻类,系统发育上与绿藻、褐藻、蓝藻细菌等相关。具固着器、叶状体为球状或具同心纹构造的Apsidiumsphaericus的发现,进一步证明典型埃迪卡拉动物群(或白海生物群)中简单的圆盘状化石可能为多细胞藻类,而不是水母。武陵山生物群中具同心纹结构的大型圆盘状化石的出现及其共生生物特点,指示该生物群是埃迪卡拉期生物分异发展阶段的产物,是末前寒武纪晚期一个以多细胞藻类发育为特色的新型生物群。 相似文献
7.
澄江生物群分子在贵州遵义牛蹄塘组发现 总被引：8，自引：1，他引：7
澄江生物群是认识寒武纪生命大爆发的重要窗口,它的分布范围一直仅限于滇东地区,员近在贵州遵义松林牛蹄塘组底部页岩中发现有Naronia,Archotuba Conoidalis,Isaxys,Perpicaris,Lingulepis,Tsunyidiscus,Scenella,水母状化石(medusiform fossil),海绵动物化石Leptomitus,类似半索动物秆壁虫Rhabdopleura化石,软舌螺类Hyolithids,宏观藻类和疑源类化石Zunyiphyton perelegans Yang et Zhao,1999,Yuknessia sp.,Longenema Ding,1996,Sphaerocongregus variabilis等。其中有一部分是属于澄江生物群分子,这是云南省外澄江生物群分子的新发现。由于生物群之下40m处还产大量海绵动物、高肌虫和藻类化石等组成的松林生物群,其上又有下寒武统明心寺组古杯动物群,因此,这一生物群的发现对研究早寒武世生物的演化也具有一定的科学意义,同时,对澄江生物群的古生态学研究也具有重要意义。 相似文献
8.
Hiemalora是一类广为分布的埃迪卡拉(Ediacaran)纪(震旦纪)圆盘状印痕化石,周围具流苏状或触须状突起,曾被解释为似水母动物,或某种叶状生物的固着器官,但以往罕见其与叶状生物体共同保存。本文报道了华南贵州震旦纪陡山沱组一类保存完好丰富的被确认为藻类的碳膜化石——革辛娜藻Gesinella,具有完整的须状固着构造。这一须状构造的形态和大小与Hiemalora非常相似,致密加厚的基部及其周围辐射状分布的丝状或须根状构造,可以分别和后者的中央盘以及周缘突起的"触须"相比较。陡山沱组的这一类碳膜化石同时保存有明显巨大的形同藻类的叶片和固着器,为研究埃迪卡拉生物群的底栖印痕化石提供了新的认识,即:盘状的Hiemalora可能是多细胞藻类的固着器官,而不一定是最初解释的似水母动物或其它叶状体生物的固着构造。不过,证实这一点尚需更进一步地研究埃迪卡拉纪生物的保存方式和埋藏环境。 相似文献
9.
最早的栉水母动物化石——华南伊迪
卡拉纪的“八臂仙母虫” 总被引：2，自引：1，他引：1
华南伊迪卡拉系陡山沱组保存的磷酸盐化球状化石中有一部分被认为是最早的后生动物胚胎化石, 但由于缺乏成年或幼年过渡期的化石证据, 对这些最早微化石的动物属性还一直存在争议。最近在贵州江口县翁会地区伊迪卡拉(震旦)系陡山沱组的页岩中发现一个与“庙河生物群”相似的宏体碳质压膜化石群, 其中包含一个盘状的宏体化石新类型——八臂仙母虫(Eoandromeda octobrachiata, Tang et al. 2008), 具有八条螺旋状辐射的旋臂, 封闭在一个包膜内。这种八辐射结构在八射珊瑚和栉水母动物中都可见到, 但同时具有的螺旋特性与现生的所有类群都无法对比。在贵州陡山沱组的“瓮安生物群”中发现的球状胚胎化石同样具有螺旋性状, 尽管还没有直接的个体发育证据, 但我们认为新属种可能代表这些螺旋状胚胎化石的成年实体化石。最新报道的南澳大利亚Rawnsley石英岩层中也发现相似的印痕化石, 被归为同一类别, 表明华南的“庙河生物群”与澳大利亚的“伊迪卡拉生物群”时代相当。翁会化石库中少量出露的三辐射化石与俄罗斯“白海生物群”的典型分子—Albumares和Anfesta可以对比。新发现的倾斜保存的八臂仙母虫化石和碳质条带呈子午线状排列的核桃状化石, 可能是该类化石归属栉水母类的有力证据, 这一发现推测可将栉水母类的最早化石记录从“澄江生物群”的早寒武世推前大约3千万年。 相似文献
10.
新元古代与寒武纪之交,即伊迪卡拉纪—寒武纪过渡期,是地球历史上一个重大的地质突变期,出现了一系列地质环境变化事件和生物快速演化事件。云南东部位于扬子地区西南缘,在前寒武纪—寒武纪过渡期属于开放式浅水台地至台缘斜坡环境,是中国伊迪卡拉(震旦)系顶部与底寒武统基本连续沉积的典型地区,也曾是中国震旦系与寒武系界线层型候选剖面所在地。近年来,在此新发现了更为丰富的宏体藻类化石、遗迹化石和伊迪卡拉生物群的疑似后生动物化石等(江川生物群),以及大量以条带状Shaanxilithes为代表的宏体化石层。相关剖面自下而上从灯影组至磷矿沉积层(中谊村段)出露有连续的宏体化石记录,研究后可以细分伊迪卡拉系顶部灯影峡阶的化石带,并与底寒武统晋宁阶、梅树村阶对接,具有作为标准层型剖面的较大潜力,为伊迪卡拉纪末期年代地层单位的建阶分层、对比及确立层型剖面,以及与寒武纪最早期年代地层的界线划分提供了更为理想的标准。 相似文献
11.
Living medusozoans and their Middle Cambrian ancestors are characterized fundamentally by four-fold symmetry. The symmetric pattern of their earlier antecedents during the Ediacaran–Cambrian transition, traditionally expected to be tetramerous, needs to be reconsidered in the light of Cambrian pentamerous fossil medusozoans. Here we present a microanatomic analysis of three tiny tetramerous specimens from the Lower Cambrian Kuanchuanpu Formation in southern China; they display diagnostic characteristics of Cubomedusae, including paired but well-separated interradial tentacles deep in the subumbrellar cavity, interradial septa connected by claustra, coronal muscles at the subumbrella, and an annular velarium suspended by twelve frenula. These fossils likely represent three new taxa of a monophyletic stem group of cubomedusans, which is further confirmed by our phylogenetic analysis based on the data matrix with 25 taxa and 107 characteristics. Among these fossil cubozoans and co-occurring pentamerous forms, the specimen ELISN31-31 is mostly close to the crown-group Cubomedusae with regard to its narrow vascular cavity and the endodermic perradial fusion. Diverse symmetrical patterns among different lineages of early Cambrian medusozoan candidates might have been independently evolved in different classes of medusozoans. 相似文献
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QIN Jiachen LIU Yunhuan SHAO Tiequan WANG Qi ZHANG Yanan ZHOU Xingyu LIU Mingjin 《《地质学报》英文版》2023,97(4):1026-1037
A wide variety of scalidophoran worms have been found in the Cambrian Fortunian in recent years, suggesting that scalidophorans diversified during this age. Here, we report new scalidophoran material obtained from the 2nd member of the Kuanchuanpu Formation(Fortunian Stage) in Zhangjiagou Lagerst?tte, Xixiang County, Shaanxi Province, South China. Three indeterminate forms of scalidophoran worms are described in detail. The morphology of these worms differs from that of any coeval worms and may ... 相似文献
13.
Tae-Yoon S. Park Ji-Hoon Kihm Jusun Woo Yong-Yi Zhen Michael Engelbretsen Jongsun Hong Suk-Joo Choh Dong-Jin Lee 《《地质学报》英文版》2016,90(3):827-837
Five species, Lipopora lissa Jell and Jell, 1976, Lipopora daseia Jell and Jell, 1976, Tretocylichne perplexa Engelbretsen, 1993 from Australia, Cambroctoconus orientalis Park, Woo, Lee, Lee, Lee, Han and Chough, 2011 from China, and Cambroctoconus kyrgyzstanicus Peel, 2014 from Kyrgyzstan, belonging to the Cambrian stem-group cnidarians have been documented in the fossil record. Cambroctoconus coreaensis sp. nov., interpreted here as a stem-group cnidarian, from the Seokgaejae section in the Daegi Formation, Taebaek Group (Cambrian Series 3), Taebaeksan Basin, central-eastern Korean Peninsula, has a slender cup-shaped skeleton. A cladistic analysis produced 21 most parsimonious trees, which invariably placed the six stem-group cnidarians below the crown-group, but their relationships within the stem-group are unresolved. Nine out of the 21 trees suggest a monophyletic relationship for the Cambrian stem-group cnidarians, whereas in six other trees a monophyly of Cambroctoconus and Tretocylichne appeared as the sister-group to the crown-group cnidarians with Lipopora at the most basal branch. This result may reflect the fact that crown-group cnidarians evolved in the Precambrian, and suggests that the diversity of stem-group cnidarians was a result of an independent radiation in the Cambrian. 相似文献
14.
In the recent years, exceptional fossil sites have revealed astonishing details on the anatomy, lifestyles and behaviour of Cambrian animals but surprisingly, very little is known about one of their most vital features, reproduction. We describe here in situ eggs (clusters of 3 to 30 oocytes) in the tube-dwelling priapulid worm Paraselkirkia sinica from the Cambrian Stage 3 Xiaoshiba Lagerstätte (ca. 514 Ma, South China). These oocytes were accommodated within paired tubular ovaries located in the posterior half of the primary body cavity as in modern meiobenthic priapulid worms, thus indicating that the general organization of female tubular gonads in priapulid worms has remained virtually unchanged for half a billion years. Our findings provide for the first time, key information on the reproductive organs and strategies of early ecdysozoans, a huge animal clade that dominated Cambrian marine ecosystems and accounts for a large part of today's biodiversity (e.g. arthropods). Moreover, we also emphasize the critical role of ecology on the reproductive strategies and lifestyles of both modern and Cambrian worms. 相似文献
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SHAO Tiequan WANG Qi LIU Yunhuan ZHANG Yanan LI Hewei HU Bo LI Chuang DONG Zhiguo ZHANG Hongtao 《《地质学报》英文版》2015,89(4):1398-1405
The metazoan Quadrapyrgites consists, from proximal to distal ends, of an oral area, a cone, a collar, and an apex from the Early Cambrian Fortunian Stage in Xixiang Zhangjiagou section, southern Shaanxi Province, South China. Three morphological types of the apex, the most characteristic feature of the taxon, are identified from hundreds of Quadrapyrgites specimens. The Quadrapyrgites growth zone is situated at the oral end, where 12 terminal lobes are generated, uplifted, enlarged, and evaginated to form a crest; A one-by-one of crest number from 2 to 13 has been recognized. The unique growth pattern and ontogenetic sequence refute a cubozoan or cycloneuralian affinity for Olivooides and Quadrapyrgites, and instead defend a coronate scyphozoan hypothesis. An embryonic development and ontogenetic sequence with embryo and crest is constructed. 相似文献
17.
LIU Yunhuan ZHANG Yanan SHAO Tiequan TANG Hanhu JIANG Kaituo WANG Qi CHEN Cheng DONG Junyan LIANG Yongchun DAI Jun XUE Jiaqi 《《地质学报》英文版》2016,90(5):1629-1636
Maikhanellids are a distinct group of Cambrian Fortunian small shelly fossils by the cap-shaped profiles and scaly shell ornamentation. According to the individual shell shape and ornamentation feature types of 50 maikhanellids fossils which were collected from Xixiang botia, we confirmed the family Maikhanellidae content of six genera, which are Maikhanella, Ramenta, Purella, Ramentoides, Yunnannopleura and Mediata. Unlike the halkieriid animals whose shells and spicules are articulated on different body parts, maikhanellids were presumably yielded by siphogonuchitid spicules penetrating the intermediate matrix (the "spicule shell" hypothesis), and this is the reason that some researchers treated maikhanellids and co-occurring siphogonuchitids as synonyms. The affinities of Maikhanellids and siphogonuchitids remain debatable. Here, we report three dimensionally phosphatized maikhanellid shells and siphogonuchitid spicule bundles from the Cambrian Fortunian small shell faunas of South China. They differ from all the previously reported maikhanellids and siphogonuchitids, and may represent new types. The intact and smooth surface underneath the partially preserved scales of the shells challenges the "spicule-shell" hypothesis. The present siphogonuchitid spicules differ from the maikhanellid scales in size, morphology, and .arrangement pattern, thus the spicules might not be derived from the shells. Whether the shells and spicule bundles were articulated on different body parts of the same animals remains unknown. 相似文献
18.
ZHANG Ya''nan LIU Yunhuan SHAO Tiequan WANG Qi QIN Jiachen ZHANG Wei LIANG Tanxing 《《地质学报》英文版》2022,96(2):440-446
A new maikhanellid genus, Totornatus gen. nov., from the Lower Cambrian (Fortunian Stage) of Zhangjiagou section at Xixiang County, southern Shaanxi Province, South China, is based on a new taxon T. strigatus gen. et sp. nov. Although showing similar morphology with a cap-shaped shell and an oval aperture to other members of the family, the new species differs from previous maikhanellids in its smooth apex. The apical field consists of dense, small, round or oval granules, which gradually merge to bigger, elongated, scale-like protrusions toward the apertural margin. On this basis, we establish Totornatus strigatus, which shows a transverse groove in two-ninths of its shell. This result enriches the diversity of the earliest mollusks and maikhanellids. 相似文献
19.
作为观察寒武纪大爆发主幕奥秘的最佳科学窗口,澄江化石库展示了三胚层动物(即两侧对称动物)首次全面大辐射的历史产物,让学术界终于看到了已知最古老的同时包括原口类和后口类两大支系的动物谱系庞大“树冠”的主体轮廓,从而给许多较高级动物类群的起源和早期演化研究提供了十分重要的信息。然而,该化石宝库中却很少见到作为“树干”的双胚层动物(即辐射动物)的化石记录,这给“真动物”的源头探秘蒙上了阴影。尽管广布于前寒武纪末期全球浅海域的“文德生物群”一直被认为是动物界早期一次“失败的演化试验”的牺牲品,但其中一支成功延续到早寒武世的春光虫(Stromatoveris)具有叠覆状梳齿构造,很可能代表着栉水母类的一类原始祖先。文中描述新的羽毛状动物王氏澄江海笔(Chengjiangopennawangii)不同于春光虫,前者具有彼此分离的羽枝,且羽枝上保存了排列规整的珊瑚个体虫室,因而与现代八射珊瑚中海笔类的形态学特征十分一致;新的谱系分析显示,它应该代表刺胞动物门中的一个原始类群。 相似文献