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1.
We investigated the occurrence of behaviors that maximize predator avoidance and seaward transport in estuarine decapod zoeae
by collecting larvae from discrete depths in a partially mixed estuary, Willapa Bay, Washington, USA, and relating their abundance
and vertical distribution to a suite of environmental variables. Abundances of first zoeae of Neotrypaea californiensis and Pinnotheridae were associated with tidal phase, diel phase, and water height. Both taxa were most abundant during ebb
tides, and abundances increased with water height, suggesting behaviors that enhanced seaward transport. Additionally, N. californiensis were both shallower and more abundant at night, indicative of behaviors to avoid visual predators. Our results suggest that
both tidal transport and predator avoidance are important and sometimes interactive selective forces shaping larval decapod
behavior. 相似文献
2.
Jianhua GAO Yang YANG Yaping WANG Shaoming PAN Rui ZHANG 《Frontiers of Earth Science》2008,2(3):249-261
High-resolution current velocity and suspended sediment concentration (SSC) data were collected by using an Acoustic Doppler
Current Profiler (ADCP) at two anchor stations and a cross-section in the South Channel of the Changjiang River mouth during
meso and neap tides on Nov. 16, 2003. In addition, tidal cycle (13-hour) observation at two stations was carried out with
traditional methods during the spring tide. Results indicated that resuspension occurred not only at the flood and ebb maximum,
but also in the early phase of ebb in the meso and neap tide. When tidal current transited from high to ebb phase, current
speed accelerated. Subsequently, fine-grained sediment with low critical threshold was resuspended and increased concentration.
The river mouth area remained in siltation in the meso and neap tidal phase during the observation season, with calculated
resuspension flux in the order of magnitude of 10−4–10−7 kg·m−2/s. Suspended sediment transport in the South Channel was dominated by freshwater discharge, but the Storks drift,
vertical circulation and vertical shear effect due to tidal oscillation also played an important role in resuspension and
associated sediment transport. In contrast, resuspension sediment flux in the spring tide was larger than that in meso and
neap tide, especially at the ebb maximum and flood maximum. The present study revealed that intensive resuspension corresponded
well with the larger current velocity during winter. In addition, the ‘tidal pumping’ effect and tidal gravity circulation
were also vital for forming the turbidity maximum in the Changjiang River estuary. 相似文献
3.
A tidally-induced frontal system regularly develops in a small area off Newport News Point in the lower James River, one of the tributaries of the Chesapeake Bay. In conjunction with the front, a strong counter-clockwise eddy develops on the shoals flanking the northern side of the channel as the result of tidal interaction with the local bathymetry and estuarine stratification. A three-dimensional hydrodynamic model was applied to simulate the eddy evolution and front development, and to investigate time-varying circulation and material transport over a spring-neap tidal cycle. The model results show that variation of tidal range, together with periodic stratification-destratification of the estuary, has a significant impact on the residual circulation of the lower James River. The net surface water circulation, which takes the form of a counterclockwise eddy on the Hampton Flats, is stronger during neap tide than during spring tide. Strong stratification and weak flood current during neap tide results in a dominant ebb flow at the surface, which delays flooding within the channel and advances the phase lead of flood tide on shoals adjacent to the channel, thus increasing both period and intensity of the eddy. Front development in the area off Newport News Point provides a linkage between shoal surface water and channel bottom water, producing a strong net upriver bottom transport. The existence of the vertical transport mechanism was independently demonstrated through tracer experiments. The impact of the dynamics on larval dispersion was investigated through a series of model simulations of the movement of shellfish larvae over multiple tidal cycles following their release at selected bottom sites. These results show that eddy-induced horizontal circulation and vertical transport associated with the frontal system are important mechanisms for the retention of larval organisms in the James River. 相似文献
4.
We determined tidal, diel (day-night), and diurnal (day to day) patterns of occurrence for the summer zooplankton assemblage in an intertidal salt marsh basin at North Inlet Estuary, South Carolina. In one time series, 153 μm pump and 365 μm net collections were made every 1–2 h during four consecutive tidal cycles. Taxonomic composition remained unchanged throughout most of the 48-h period, but densities and proportionalities of individual taxa were highly variable. Recurring patterns of abundance were observed and taxon-specific relationships with the tidal and diel cycles were indicated. Zooplankton were not uniformly distributed within flooding-ebbing water masses and distributions could not be explained by simple passive advection with the tides. Diel differences in densities of copepods and bivalves resulted from behavioral responses to changing light conditions. Large pulses of crab and shrimp larvae originating from nocturnal hatching events within the intertidal basin exited but did not return during the next flood tide. Higher densities of postlarval decapods on flood tides indicated settlement and recruitment to the shallow basin. In a second time series, replicated collections of the 153 μm and 365 μm assemblages were made during the daytime ebb tide every 1–3 d from May through October 1991 to determine relationships between diurnal changes in depth, salinity, and temperature and zooplankton composition and abundance. Diurnal variations in densities and proportionalities were less than those observed during the 48-h study and patterns were not regular. For most taxa, relationships between depth and abundance were the same in both time series. During periods of reduced salinity, densities of copepods,Uca zoeae, and barnacle nauplii decreased and densities ofUca megalopae andPenaeus postlarvae increased. However, zoeae emerged and postlarvae recruited throughout the 5-mo period, indicating that considerable flexibility in responses and tolerances existed within the populations. The diversity of life-history strategies and behavioral adaptations found among the zooplankton assures continuous occupation of flooded intertidal habitats. We suspect that the evolution and maintenance of temporally staggered recurring patterns of occurrence results in reductions in the competition for resources. 相似文献
5.
Depth-specific sampling at a single location was used to examine the vertical distribution of pediveliger larvae of bivalve mollusks in the York River, an estuary of Chesapeake Bay, Virginia. The water column at the sampling site was usually well mixed, lacking consistent temperature or salinity gradients for larvae. Four species showed strong vertical stratification when collected simultaneously at three depths. Pediveliger stage larvae of a clam (Cyrtopleura costata), an oyster (Crassostrea virginica), and a shipworm (Bankia gouldi) were most abundant near the benthos, and least abundant near the surface. A mussel (Geukensia demissa), showed the reverse trend, with most pediveliger larvae near the surface. Tidal stage had a slight effect on two species (C. costata andC. virginica), but only to increase the relative abundance of larvae near the benthos during flood tide. Otherwise, neither tidal phase nor light levels (night versus day) had detectable effects on distribution patterns. Sampling very close to the sediment-water interface provided no evidence that pediveliger larvae spent a significant proportion of their time in this location. While the behavior of precompetent bivalve larvae may tend to retain them within an estuary, that of competent-to-metamorphose pediveliger larvae does not appear to have that function. Pediveliger larvae may regulate their depth to best locate potential settlement substrate. Such an hypothesis is consistent with adult habitat zones of at least some of these species in Chesapeake Bay. Removing competent-to-settle larvae from analyses of larval distributions in estuaries will enhance or clarify depth distribution patterns observed for earlier-stage larvae. 相似文献
6.
A 16-month study of estuarine habitats in poly-, meso-, and oligohaline salinity regimes near Charleston Harbor assessed the distribution and abundance of megalopae and early crab stages of the blue crab,Callinectes sapidus. Blue crab were sampled with a plankton net and a cylindrical drop sampler. Blue crab were most abundant in plankton collections at night, accounting for 68% of the megalopae and over 88% of the juveniles collected in day and night tows combined. At night, densities of megalopae were greatest in surface samples, whereas densities in daylight collections were greater on the bottom. Juvenile densities were greatest on the bottom in both day and night collections, although catch rates at night were more variable than those of the megalopae. This suggests that megalopae, and possibly juvenile stages, experience a diel vertical migration. Results indicate that ingress to estuarine nursery areas occurs at the megalopal stage. Megalopal densities were highest at the polyhaline site, while juvenile blue crab were most abundant in the oligohaline area. Habitat utilization by juvenile blue crab was estimated using a cylindrical drop sampler and Venturi suction pump on three bottom types in the intertidal zone. Densities were greatest over the sandy-mud substrate, although catch rates were much lower than those reported for other geographical areas. These results suggest that juvenile blue crab do not occur in abundance on the marsh surface but remain on the creek bottom, possibly because creek physiography and large tidal amplitudes may restrict accessibility to the marsh surface. 相似文献
7.
Measurements of velocity, salinity, and suspended solids concentration have been used to investigate the intra-tidal variation of vertical and transverse shear-induced dispersion. For the study research the interaction of the longitudinal density gradient and vertical shear during the early part of the ebb tide accounted for much of the net longitudinal dispersion of solute landward. The same mechanism also is shown to lead to a net particulate transport landward. The landward flux, however, takes place during the flood tide. The field data are also used to elucidate the tidally averaged tidal pumping mechanism. 相似文献
8.
We describe the tidal circulation and salinity regime of a coastal plain estuary that connects to the ocean through a flood
tide delta. The delta acts as a sill, and we examine the mechanisms through which the sill affects exchange of estuarine water
with the ocean. Given enough buoyancy, the dynamics of tidal intrusion fronts across the sill and selective withdrawal (aspiration)
in the deeper channel landward appear to control the exchange of seawater with estuarine water. Comparison of currents on
the sill and stratification in the channel reveals aspiration depths smaller than channel depth during neap tide. During neap
tide and strong vertical stratification, seawater plunges beneath the less dense estuarine water somewhere on the sill. Turbulence
in the intruding bottom layer on the sill promotes entrainment of fluid from the surface layer, and the seawater along the
sill bottom is diluted with estuarine water. During ebb flow, salt is effectively trapped landward of the sill in a stagnant
zone between the aspiration depth and the bottom where it can be advected farther upstream by flood currents. During spring
tide, the plunge point moves landward and off the sill, stratification is weakened in the deep channel, and aspiration during
ebb extends to the bottom. This prevents the formation of stagnant water near the bottom, and the estuary is flooded with
high salinity water far inland. The neapspring cycle of tidal intrusion fronts on flood coupled with aspiration during ebb
interacts with the sill to play an important role in the transport and retention of salt within the estuary. 相似文献
9.
采用大凌河实测长系列河道径流及水下地形等数据, 通过建立水动力数值模型, 综合考虑潮流和河道径流的影响, 对大凌河口的潮流进行模拟, 探讨大凌河口及辽东湾北部海域水动力过程的主要特征. 结果表明: 大凌河口附近海域的海流以潮流为主, 具有明显的往复性质. 潮流总的运动趋势是, 涨潮主流向为北东, 落潮主流向为南西. 大潮流速大于小潮流速, 涨潮流历时与落潮流历时几乎相当. 最大涨潮流速约为0.52 m/s, 最大落潮流速约为0.4 m/s, 潮流涨落平均潮流强度的分布大体和该海域等深线相适应. 相似文献
10.
A rhodamine dye tracer study was conducted over eight tidal cycles to investigate mixing and tidal exchange processes in Perch Pond, a Cape Cod embayment subject to recurrent blooms of the toxic dinoflagellate, Gonyaulax tamarensis. Dye injected at the inlet to Perch Pond during flood tide became well-mixed within the pond in one day and was removed at an effective first order rate of 0.36 d?1, equivalent to a 70% utilization of the maximum possible tidal exchange. This relatively high flushing efficiency can be attributed to a density-driven circulation within the pond, consisting of a subsurface inflow of high salinity dense water on the flood tide followed by removal of lighter surface layers through the shallow inlet during ebb tide. The formation of a frontal convergence near the inlet on flood tide is consistent with the observed distribution of G. tamarensis cysts and shelifish toxicity. It is also clear that phytoplankton like G. tamarensis, whose maximum growth rates approximate the rate of tidal flushing, can only bloom within the embayment by avoiding the outflowing surface waters. Mixing within the pond is probably less efficient and population losses greater during dry periods when the pond salinity is higher and the stratification weaker. 相似文献
11.
In October of 2004, a 3-d observational program to measure flow and sediment resuspension within a coastal intertidal salt
marsh was conducted in the North Inlet/Winyah Bay National Estuarine Research Reserve located near Georgetown, South Carolina.
Current and acoustic backscatter profiles were obtained from a moored acoustic Doppler current profiler (ADCP) deployed in
a shallow tidal channel during the spring phase of the tidal cycle under high discharge conditions. The channel serves as
a conduit between Winyah Bay, a large brackish estuary, and North Inlet, a saline intertidal coastal salt marsh with little
freshwater input. Salinity measurements indicate that the water column is vertically well mixed during flood, but becomes
vertically stratified during early ebb. The stratification results from brackish (15 psu) Winyah Bay water entering North
Inlet via the tidal channel, suggesting an exchange mechanism that permits North Inlet to receive a fraction of the poor water
quality and high discharge flow from upland rivers. Although maximum flood currents exceed maximum ebb currents by 0.2 m s−1, suspended sediment concentrations are highest during the latter ebb phase and persist for a longer fraction of the ebb cycle.
Even though the channel is flood-dominated, the higher concentrations occurring over a longer fraction of the ebb phase indicate
net particulate transport from Winyah Bay to North Inlet during spring tide accompanied by high discharge. Our evidence suggests
that the higher concentrations during ebb result from increased bed friction caused by flow asymmetries and variations in
water depth in which the highest stresses occur near the end of ebb near low water despite stronger maximum currents during
flood. 相似文献
12.
The utilization of an intertidal salt marsh creek in South Carolina during January 1977 was determined by sampling every third ebb tide for 13 days. All fishes leaving the creek during that period were captured in a channel net. This procedure produced a time-series of samples which permitted analysis of the fish community occupying the intertidal creek at all times of day and night. A total of 14,730 larval, juvenile, and adult fishes comprising at least 22 species in 16 families were collected. The most common larval and juvenile fishes wereLeiostomus xanthurus, Mugil spp.,Myrophis punctatus leptocephali,Lagodon rhomboides, Paralichthys spp., andMicropogon undulatus. Catch sizes for all species varied widely between samples. No diurnal-nocturnal pattern in catches was evident forL. xanthurus, Mugil spp.,L. rhomboides andM. undulatus. M. punctatus was taken in large numbers only when the flood tide occurred during the day, while moreParalichthys spp. larvae were taken in late afternoon-evening flood tide samples. The most common invertebrate,Palaemonetes pugio, was taken in large quantities only in late afternoon-night flood tide samples. Three diversity indices were computed for each sample. Values for all indices varied widely between successive samples. The results emphasize a high degree of utilization of the intertidal creek habitat by larval and juvenile fishes. The diurnal-nocturnal activity patterns of some species, and the wide variation in catch size of the other species can permit use of the intertidal salt marsh habitat with reduced competition for available space and energy. 相似文献
13.
G. Curtis Roegner David A. Armstrong Barbara M. Hickey Alan L. Shanks 《Estuaries and Coasts》2003,26(4):1058-1070
We investigated the distribution of meroplankton and water properties off southern Washington and simultaneously measured time series of larval abundance and water properties in two adjacent estuaries, Grays Harbor and Willapa Bay. The cruise period, in late May 1999, coincided with large variation in the alongshore wind stress that caused dynamic change in the position of the Columbia River plume, coastal upelling and downwelling, and offshore phytoplankton production. In the coastal ocean, meroplankton groups responded differently to this wind event and the associated advection of water masses. Dungeness crab (Cancer magister) megalopae were largely indifferent to the wide salinity variation, and were found throughout the surveyed area in both plume and recently upwelled waters. Megalopae of kelp crab (Pugettia producta) and hermit crab (Pagurus spp). were more abundant in upwelled water and low numbers were caught in the plume water. Barnacle cyprids appeared to track the advective transport suggesting that they may be more passively dispersed. Within the estuaries, hydrography responded rapidly and synchronously to variation in wind stress. Intrusions of both plume and newly upwelled waters were detected at estuarine sites, depending on the type of water present at the coast, indicating a tight link between the estuaries and the coastal ocean in this region. A 90-d record ofC. magister megalopae abundance was made at 3 estuarine sites using light traps. The bulk of theC. magister recruitment was limited to a relatively brief period in late May through June. Within this window, megalopae occurred in distinct pulses of 3–5 d interspaced with periods of low or zero abundance.C. magister megalopae recruited to the estuaries over a wide range of wind forcing, and were transported into the estuary within varied water types. There were no periodic patterns indicative of spring-neap tidal variations in the abundance time series. Abundance was only weakly cross-correlated between the adjacent Grays Harbor and Willapa Bay estuaries, which contrasts with the more synchronous estuarine-coastal linkages measured for water properties. These results suggest the interaction of larval aggregation size in the ocean with estuary-ocean exchange processes likely controls patterns of estuarine recruitment. 相似文献
14.
为研究磨刀门盐水混合层化特征,基于SCHISM模型,建立了三维盐度数值模型,根据实测资料对其进行验证。结合水体势能异常理论,对枯季磨刀门河口混合层化的时空变化特征及深槽与浅滩的层化机制差异进行分析。结果表明:磨刀门河口小潮时水体层化最强,中潮时水体层化最弱,且拦门沙至挂定角段水体层化始终较强。磨刀门深槽水体层化主要受纵向平流、纵向水深平均应变和垂向混合影响,而浅滩水体层化则受横向平流、横向水深平均应变和垂向混合影响;磨刀门河口表、底层水体湍动能耗散率较高,而中间水层存在低耗散区,且涨潮时湍动能耗散率比落潮时大。 相似文献
15.
现场试验表明,三角架观测系统稳定性良好,获取了边界层内多层位、连续的温、盐、流速、浊度同步观测数据,适用于浅海近底部沉积动力过程高分辨率观测及物质输运研究。观测结果显示:观测期间,边界层内存在向陆的余流,并呈现逐渐减小的趋势,其主要由涨、落潮流的不对称造成,大风天气和密度环流亦是影响余流强弱的重要因素;观测期间多数时刻底部切应力大于起动切应力,底质沉积物可产生明显的搬运甚至再悬浮;悬沙浓度对沉积动力的响应在涨、落潮,大、小潮阶段均有各自的特点,水动力的变化、潮流加/减速时间的长短、床面泥沙的供应量、上部水体泥沙的沉降是导致悬沙浓度变化的主要原因;底部边界层内,涨、落潮期间不对称输沙导致潮周期内悬沙净向河口湾内输运。 相似文献
16.
Habitat selection capabilities of the recruiting larval stages of marine invertebrates are limited, in part, by their ability to maneuver in flowing water. Distributional and experimental evidence suggest that blue crab (Callinectes sapidus) megalopae may preferentially settle into vegetated habitats. However, the behavior and swimming capabilities of megalopae in flowing water have not previously been investigated. Laboratory experiments were conducted in a small, recirculating seawater flume to determine the swimming response of megalopae to varying flow velocities. Nighttime trials were conducted at six flow velocities: 0, 1.9, 3.6, 4.8, 6.3, and 9.3 cm s?1. Behavior and swimming velocities of field-collected C. sapidus megalopae were video recorded. Megalopae exhibited negative phototaxis and were found in the water column at all flows in the dark. The maximum sustained swimming speed observed was 12.6 cm s?1 and the mean swimming speed in still water was 5.0 cm s?1, with short bursts in excess of 20 cm s?1. Megalopae frequently oriented into the current and were capable of swimming upstream against the current at flow speeds <4.8 cm s?1; at greater velocities they were not able to do so. The results suggest that at low to moderate current velocities C. sapidus megalopae have the ability to actively move in search of settlement sites and to maintain their positions in desirable sites rather than relying strictly on passive movements by currents. 相似文献
17.
J. C. Joyeux 《Estuaries and Coasts》1999,22(4):889-904
The larvae of winter spawning fishes immigrating through Beaufort Inlet into the Pamlico Sound estuarine system (North Carolina, United States) were passively sampled during 14 flood tides and nine of the following ebb tides. Five taxa were abundant in the catches. Pelagic species were represented by Atlantic menhaden, Brevoortia tyrannus. Nonpelagic taxa were represented by Atlantic croaker, Micropogonias undulatus, spot, Leiostomus xanthurus, pinfish, Lagodon rhomboides, and flatfishes of the genus Paralichthys. The sampling was continuous and the sample duration varied between 4 min and 32 min. The longest samples furnished the most accurate and precise estimates of the mean tidal abundance. Sampling 10 min each hour of the tide was the most efficient protocol for determining the mean tidal abundance. The abundance patterns was shown to differ according to the pelagic or nonpelagic behavior of the larvae. Analyses suggested the nonpelagic taxa rely on astronomical tides and vertical migrations synchronous with the direction of the tide flow to be transported upstream in the estuary. These larvae were not dependent upon the strength of the tide to penetrate the inlet. In contrast, menhaden larvae seemed to rely primarily on strong food tides to enter the estuarine area, probably because the vertical positioning of the species within the water column is not dependent upon the direction of the tide. However, if larvae were present outside of the inlet, strong flood tides indiscriminately brought pelagic and nonpelagic species into the estuary. 相似文献
18.
The particulate organic matter (POM) in hydrodynamically variable habitats such as the lower reaches of estuaries can change in its content and quality on very short time scales (example, hourly), and these changes can potentially influence higher-level consumers in river-estuary-marine systems. Estuarine water samples were collected hourly for 12 h downstream in a small river to evaluate the fatty acid composition of POM over a tidal cycle. Fatty acid constituents of POM collected during the flood tide were dominated by the saturated, higher plant and bacterial fatty acids, whereas unsaturated, polyunsaturated, essential, and diatom-associated fatty acids dominated the POM collected during the ebb tide. Elevated algal biomass (as indicated by high chlorophyll a concentrations), diatom, and freshness indices in the POM indicated enhanced fresh autochthonous-origin materials that dominated the mixed organic pool during the ebb tide compared to more degraded detritus during the flood tide. Tidal retention of organic matter and algal primary production were the most influential factors that differentiated the fatty acid composition of estuarine POM over the short time scale. The results of this study have important implications on the quality of POM at the time of sampling, especially in estuaries where mixed organic pools have multiple inputs and are strongly influenced by tidal cycles. 相似文献
19.
In the Grays Harbor estuary, juvenile Dungeness crab (Cancer magister Dana) are found at higher densities in epibenthic shell deposits compared to open mud flat. Differences in predation rate between habitats have been suggested to be due to habitat preference and differential survival. Megalopae preferred shell over open space in still-water conditions. However, it is not known whether megalopae are able to select shell in flowing water since larval preference is known to differ between still and flowing water. Here we report the first experimental study of swimming behavior of Dungeness crab megalopae in a range of current velocities (0–40 cm s?1) equivalent to natural flow in Grays Harbor estuary. Experiments were conducted in daylight using a recirculating flume. Megalopae swimming speeds ranged from 8.5 cm s?1 (8 body lengths s?1) in still water to 44.8 cm s?1 (44 body lengths s?1) at flow speeds of 40 cm s?1, Neither swimming behavior nor sheltering behavior in shells showed any flow-related pattern. Megalopae spent a large proportion of time swimming against the current and made headway upstream against all current velocities tested. The results suggest that Dungeness crab megalopae are able to maneuver and actively search for settlement sites under current velocities found in natural habitats, including intertidal shell deposits, and support the hypothesis of active selection of shell by megalopae. 相似文献
20.
Diel vertical migration (DVM) and seasonal timing of three sympatricCancer spp. larvae were investigated in southeastern Alaska. Diel sampling was conducted at a station in Icy Strait at 0400, 0700, 1000, 1300 h in the first day, and 1300, 1600, 1900, and 2200 h in the second day from June to September 2001. Larvae were collected with Bongo nets (333 and 505 μm mesh) towed vertically from 20 m depth to the surface. During flood tides surface temperature increased slightly and salinity decreased; the reverse occurred during ebb tides. Zooplankton biomasses were higher in the early morning and evening than during the midday. A total of 4,482 larvae were examined for the study.Cancer oregonensis larvae were most abundant (4,238), followed byC. magister (205) andC. productus (39). Larval density peaked in June while no larvae were found in September.C. magister larvae had a crepuscular migration; larval abundance peaked in early morning and evening.C. oregonensis larvae comprised 94.6% of allCancer spp. larvae collected and had a strong DVM in June. Earlier zoeal stages ofC. oregonensis were relatively more abundant during midday, while later stages were more abundant at night. In July, later larval stages ofC. oregonensis were distributed at the surface all day with a peak at 2200 h. Larvae ofC. productus first occurred in July and peaked in August.C. productus had DVM but many were at the surface diurnally.C. magister andC. oregonensis larvae co-occurred seasonally as they do in the lower latitudes, e.g., Washington and British Columbia; however,C. productus appeared approximately 2 mo later thanC. magister andC. oregonensis in southeastern Alaska. 相似文献