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1.
The presence and localization of organic sulfate within coral skeletons are studied by using X-ray absorption near edge structure spectroscopy (XANES) fluorescence. XANES spectra are recorded from four reference sulfur-bearing organic molecules: three amino acids (H-S-C bonds in cysteine; C-S-C bonds in methionine; one disulfide bond C-S-S-C bonds in cystine) and a sulfated sugar (C-SO4 bonds in chondroitin sulfate). Spectral responses of three coral skeletons show that the sulfated form is extremely dominant in coral aragonite, and practically exclusive within both centres of calcification and the surrounding fibrous tissues of coral septa. Mapping of S-sulfate concentrations in centres and fibres gives us direct evidence of high concentration of organic sulfate in centres of calcification. Additionally, a banding pattern of S-sulfate is visible in fibrous part of the coral septa, evidencing a biochemical zonation that corresponds to the step-by-step growth of fibres.  相似文献   

2.

Elizabeth and Middleton Reefs are atoll-like structures that have developed on top of volcanic edifices and are close to the southern environmental limit of reef development in the southwest Pacific. Reef morphology and vertical accretion rates during the Holocene appear similar to those on other more tropical reefs. Sediment samples were collected from the lagoon of both reefs and around the flanks of Middleton Reef. A distinctly chlorozoan assemblage was observed with coral, molluscs, Halimeda, coralline algae and foraminifers being the dominant sediment constituents. Lagoon sediment samples show little variation within or between reefs, lacking the concentric zonation characteristic of larger atolls. Samples collected from the flanks of Middleton Reef, and subsurface material from vibrocores, differ compositionally from the surficial lagoon sand and were typically more tropical in character. A comparison of the sediment constituents from these reefs with those of samples from within a fringing reef and from the shelf around Lord Howe Island, further south, indicated regional patterns in sediment composition. Halimeda rapidly decreased in abundance with increased latitude, and appeared confined to deeper water, whereas coralline red algae increased significantly. The rapid change in these major sediment contributors is coincident with the general decrease in coral growth rates with latitude. This reinforces the notion that the latitudinal limit of reef development is constrained by factors other than coral growth alone.  相似文献   

3.
The Darwin Mounds are small (up to 70 m in diameter), discrete cold‐water coral banks found at c. 950 m water depth in the northern Rockall Trough, north‐east Atlantic. Formerly described in terms of their genesis, the Darwin Mounds are re‐evaluated here in terms of mound growth processes based on 100 and 410 kHz side‐scan sonar data. The side‐scan sonar coverage is divided into a series of acoustic facies representing increasing current speed and sediment transport/erosion from south to north: pockmark facies, ‘mounds within depressions’ facies, Darwin Mound facies, stippled seabed facies and sand wave facies. Mound morphometric changes are quantified and show a south‐to‐north divergence from an inherited morphology, reflecting the outline of coral‐colonized fluid escape structures, to developed, downstream elongated, elevated mound forms. It is postulated that increasing current speeds and bedload sand transport favour mound growth and development by a process of enhanced sand sedimentation within mounds due to current deceleration by frictional drag around coral colonies. Comparisons are made with similar growth processes attributed to comparably sized cold‐water coral mounds in the Porcupine Seabight, offshore Ireland.  相似文献   

4.
Cold‐water coral mound morphology and development are thought to be controlled primarily by current regime. This study, however, reveals a general lack of correlation between prevailing bottom current direction and mound morphology (i.e. footprint shape and orientation), as well as current strength and mound size (i.e. footprint area and height). These findings are based on quantitative analyses of a high‐resolution geophysical dataset collected with an Autonomous Underwater Vehicle from three cold‐water coral mound sites at the toe of slope of Great Bahama Bank. The three sites (80 km2 total) have an average of 14 mounds km?2, indicating that the Great Bahama Bank slope is a major coral mound region. At all three sites living coral colonies are observed on the surface of the mounds, documenting active mound growth. Morphometric analysis shows that mounds at these sites vary significantly in height (1 to 83 m), area (81 to 6 00 000 m2), shape (mound aspect ratio 0·1 to 1) and orientation (mound longest axis 0 to 180°). The Autonomous Underwater Vehicle measured bottom current data depict a north–south flowing current that reverses approximately every six hours. The tidal nature of this current and its intermittent deviations during reversals are interpreted to contribute to the observed mound complexity. An additional factor contributing to the variability in mound morphometrics is the sediment deposition rate that varies among and within sites. At most locations sedimentation rate lags slightly behind mound growth rate, causing mounds to develop into large structures. Where sedimentation rates are higher than mound growth rates, sediment partially or completely buries mounds. The spatial distribution and alignment of mounds can also be related to gravity mass deposits, as indicated by geomorphological features (for example, slope failure and linear topographic highs) in the three‐dimensional bathymetry. In summary, variability in sedimentation rates, current regime and underlying topography produce extraordinarily high variability in the distribution, development and morphology of coral mounds on the Great Bahama Bank slope.  相似文献   

5.
祝仲蓉 Marsh.  J 《沉积学报》1992,10(1):133-145
更新世以来,剧烈的构造运动已将巴布亚新几内亚合恩半岛东北海岸的晚第四纪珊瑚礁阶地抬升上千米.阶地中造礁珊瑚的成岩变化和成岩产物的组构特征反映了该礁的成岩历史,充分体现该区快速构造上升的影响.海水潜流带和淡水渗流带为上升礁的主要成岩环境.生物钻孔、生物碎屑填隙、珊瑚文石针粗化、珊瑚骨骼的溶解和新生变形转化,以及其不同矿物成分和组构的种种胶结物的胶结作用是造礁珊瑚经历的主要成岩作用.地球化学资料表明其成岩变化发生于开放的化学体系之中.  相似文献   

6.
我国生物礁研究的发展   总被引：7，自引：0，他引：7  
赵焕庭 《第四纪研究》1996,16(3):253-262
黄汲清教授研究我国南方二叠系，引发了我国对古代生物礁的研究工作。６０年代起我国开始对古代生物礁的研究。现代珊瑚礁研究从２０年代末兴起，８０年代以来正在深入发展，同全球变化研究和珊瑚礁开发紧密联系，拓展了研究领域，积累了许多新知。  相似文献   

7.
High-resolution seismic profiles, swath bathymetry, side-scan sonar data and video imageries are analysed in this detailed study of five carbonate mounds from the Belgica mound province with special emphasis on the well-surveyed Thérèse Mound. The selected mounds are located in the deepest part of the Belgica mound province at water depths of 950 m. Seismic data illustrate that the underlying geology is characterised by drift sedimentation in a general northerly flowing current regime. Sigmoidal sediment bodies create local slope breaks on the most recent local erosional surface, which act as the mound base. No preferential mound substratum is observed, neither is there any indication for deep geological controls on coral bank development. Seismic evidence suggests that the start-up of the coral bank development was shortly after a major erosional event of Late Pliocene–Quaternary age. The coral bank geometry has been clearly affected by the local topography of this erosional base and the prevailing current regime. The summits of the coral banks are relatively flat and the flanks are steepest on their upper slopes. Deposition of the encased drift sequence has been influenced by the coral bank topography. Sediment waves are formed besides the coral banks and are the most pronounced bedforms. These seabed structures are probably induced by bottom current up to 1 m/s. Large sediment waves are colonised by living corals and might represent the initial phase of coral bank development. The biological facies distribution of the coral banks illustrate a living coral cap on the summit and upper slope and a decline of living coral populations toward the lower flanks. The data suggest that the development of the coral banks in this area is clearly an interaction between biological growth processes and drift deposition both influenced by the local topography and current regime.  相似文献   

8.
A common assumption in the geological analysis of modern reefs is that coral community zonation seen on the surface should also be found in cores from the reef interior. Such assumptions not only underestimate the impact of tropical storms on reef facies development, but have been difficult to test because of restrictions imposed by narrow‐diameter cores and poor recovery. That assumption is tested here using large‐diameter cores recovered from a range of common zones across three Campeche Bank reefs. It is found that cores from the reef‐front, crest, flat and rubble‐cay zones are similar in texture and coral composition, making it impossible to recognize coral assemblages that reflect the surface zonation. Taphonomic signatures imparted by variations in encrustation, bioerosion and cementation, however, produce distinct facies and delineate a clear depth zonation. Cores from the reef‐front zone (2–10 m depth) are characterized by sections of Acropora palmata cobble gravel interspersed with sections of in‐place (but truncated) A. palmata stumps. Upper surfaces of truncated colonies are intensely bioeroded by traces of Entobia isp. and Gastrochaenolites isp. and encrusted by mm‐thick crustose corallines before colony regeneration and, therefore, indicate punctuated growth resulting from a hurricane‐induced cycle of destruction and regeneration. Cores from the reef crest/flat (0–2 m depth) are also characterized by sections of hurricane‐derived A. palmata cobble‐gravels as well as in‐place A. palmata colonies. In contrast to the reef front, however, these cobble gravels are encrusted by cm‐thick crusts of intergrown coralline algae, low‐relief Homotrema and vermetids, bored by traces of Entobia isp. and Trypanites isp. and coated by a dense, peloidal, micrite cement. Cores from the inter‐ to supratidal rubble‐cay zone (+0–5 m) are only composed of A. palmata cobble gravels and, although clasts show evidence of subtidal encrustation and bioerosion, these always represent processes that occurred before deposition on the cay. Instead, these gravels are distinguished on the basis of their limited bioerosion and marine cements, which exhibit fabrics formed in the intertidal zone. These results confirm that hurricanes have a major influence on facies development in Campeche Bank reefs. Instead of reflecting the surface coral zonation, each facies records a distinctive, depth‐related set of taphonomic processes, which reflect colonization, alteration and stabilization following the production of new substrates by hurricanes.  相似文献   

9.
A new constitutive model for fibre-reinforced cohesive soil is proposed. The model combines a Cam-Clay like bounding surface model with an elastic–plastic one-dimensional fibrous element model. A “smearing procedure”, which can consider any spatial distribution of fibre orientation, is employed to transform discrete tensile forces developed in the fibres into stresses for the composite material. The fibre stress contribution is bounded by both degradation of soil–fibre bonding due to pull-out mechanism and tensile strength of the fibres. Eventual occurrence of fibre breakage is also considered. The model performances are analysed for both consolidation and shearing loading modes, and qualitative comparison is performed with experimental data available in the literature. For consolidation loading, tensile stresses are not developed in the fibres and thus the fibre effect is rather limited. For drained shear loading, addition of fibres can result in a consistent shear strength increase. The beneficial effect of fibres seems to be controlled by two parameters: the fibre tensile stiffness and the fibre/soil strain ratio that accounts for any possible slippage or shear deformation at the fibre/soil matrix interface. For undrained shear loading, the strengthening effect of the fibres appears to be counteracted by the increase in pore water pressure, induced by the additional confining contribution of the fibres. In agreement with published experimental data, the model suggests also that the moisture content is a key factor governing fibre effectiveness for undrained shearing. Finally, analysis of the model predicted critical states for fibre-reinforced cohesive soil is provided.  相似文献   

10.
海岸生物地貌过程研究海岸带生物过程和动力-沉积-地貌过程之间的双向相互作用,是海岸生态系统响应和反馈全球变化的重要机制,被列为全球变化核心项目海岸带陆海相互作用研究的重点内容。20世纪90年代以来的调查研究揭示了华南红树林和珊瑚礁热带生物海岸生物地貌过程的主要特点。造礁石珊瑚的高生长率和珊瑚礁高堆积速率是珊瑚礁生物地貌过程的物质基础;红树林生态系统的高生产力、高归还率和捕沙促淤功能是红树林生物地貌过程的物质基础。热带生物海岸地貌结构显示分带性和生物地貌类型和动力地貌类型的叠加和共存,潮汐水位严格控制群落分布格局并形成重要的生物地貌界限。热带生物海岸生物地貌过程有利于消除或减缓海平面上升的浸淹效应。热带生物海岸不断加剧的人类活动干扰和生态破坏导致生物地貌功能削弱和海岸资源环境恶化。  相似文献   

11.
Soil reinforcement through the inclusion of oriented or randomly distributed discrete elements such as fibres has recently attracted increasing attention in geotechnical engineering. Therefore, the purpose of this paper is to investigate the influence of certain parameters (the strength properties of the fibre, the relative size of the fibres and grains, and the rate of shear) on the shear strength of polypropylene fibre reinforced cohesive soils. A series of consolidated drained or undrained direct shear tests were conducted on unreinforced and reinforced sandy silt and silty clay specimens. Two types of polypropylene fibres with different mechanical indices were used. The fibre content was varied between 0.3% and 1.1% by weight of dry soil. The test results revealed that the inclusion of fibres in soil significantly increases the shear strength. The attainment of the high shear strength is attributed to the micromechanisms involved in the fibre/soil interactions as studied through scanning electron micrographs. The results also showed that the reinforcement effect was more pronounced under undrained shearing conditions. An important outcome from the current work is that, from the data obtained, the strength of the reinforced soil composites is not practically affected by the fibre mechanical indices.  相似文献   

12.
This study presents a new and innovative distribution model of the main reef builders that thrived on bioconstructed margins of Upper Jurassic Intra-Tethys platforms. The proposed model is based on the zonation of an Upper Jurassic reef complex presently exposed in the central Apennines. Excellent exposures allow the reconstruction of the reef profile across the platform margin. Biota are mainly represented by corals and calcified demosponges, distributed with different amounts and aggregating structures along the reef complex. Microencrusters represent a secondary but important component that actively contributes to the reef construction process. Reef builders are distributed in variable quantities and with variable aggregating structures along the reef complex, revealing a strong reef zonation. Like the biota, the sedimentary features (grain size, sorting and reworking degree, bioerosion, cement crusts and cavities) show a distinctive distribution along the reef complex. Three main zones have been recognized along the studied reef complex: an internal, more protected and deeper zone, corresponding to a relatively deep back-reef/lagoon, is dominated by corals and chaetetids; a central zone (reef flat), where well-preserved coral bioconstructions give rise to a true, but discontinuous framework, and a more external zone, corresponding to the reef crest, characterized by the shallowest depths recorded across the reef complex and dominated by stromatopores. The latter forms mound-shaped structures, playing a role essential for the zoned character of the reef complex. These structures actively contribute to the construction of a moderately high topographic relief in the reef crest.  相似文献   

13.
王志金  陈世悦  马帅  汪峰 《沉积学报》2017,35(6):1177-1185
对柴达木盆地东北缘石炭系地层中生物礁的发育特征及区域展布进行了系统阐述。基于野外实地考察与室内镜下分析,采用地球化学与岩石学方法相结合,总结归纳柴达木盆地东北缘石炭系生物礁的岩石学特征、生物特征以及地层特征,分析生物礁主要产出层位及规模大小,探讨其古环境与油气地质意义。研究结果表明:柴达木盆地东北缘石炭系造礁生物主要为四射珊瑚,其次为横板珊瑚,发育锥状、盘状的单体珊瑚和管状、丛状、蜂窝状的群体珊瑚,附礁生物为腕足、等;珊瑚礁在岩层中以点状产出,呈不连续带状分布,依据其在地层中的发育规律与展布特征,分为簇状竖直丛生管状珊瑚、簇状水平延伸管状珊瑚以及簇状顶端向下倾斜管状珊瑚;根据岩石中的生物颗粒含量,生物礁灰岩储层岩石类型分为珊瑚骨架灰岩、珊瑚颗粒石灰岩、生物颗粒质石灰岩、含生物颗粒石灰岩以及泥晶灰岩。柴达木盆地东北缘早石炭世以碳酸盐台地环境为主,晚石炭世以海陆交互的潮坪环境为主。研究区石炭系生物礁主要位于下石炭统怀头他拉组三、四段,在石灰沟、旺尕秀等剖面可见厚层油砂,指示柴达木盆地东北缘石炭系具有良好的生烃潜力。  相似文献   

14.
Porcupine Bank (51–54°N, 12–15°W), approximately 10,000 km2 in area and 150–500 m deep, is effectively removed from present-day terrigenous sedimentation by the broad Irish Shelf to the east, and deep water to the north, west and south. Water temperatures range from 10° to 14.5°C. Carbonate sediments presently accumulate on the bank, mixing with relict quartz sands and lithic pebbles, cobbles and boulders of glacial origin. Traction current structures are absent but biogenic hollows, pits, mounds and trails are prevalent. The lithofacies have a crudely concentric zonation with pelagic foraminiferal oozes in the deepest waters ( > 500 m), passing to foraminiferal (pelagic and benthic) glauconitic quartz sands over the bulk of the bank (500-200 m) where scattered patches of Lophelia pertusa coral branches are found, which in turn pass into a narrow zone associated with the lithic boulder field at the bank crest (< 200 m) which has coarse sands composed of quartz and skeletal fragments of benthic organisms-molluscs, echinoids, bryozoans and serpulids. This facies pattern is closely similar to that on Rockall Bank, 500 km to the north, and supports the construction of a depth-related facies model for modern open-sea temperate carbonate sediments: > 500 m, pelagic foraminiferans; 500-200 m, benthic foraminiferans plus Lophelia coral patches; 200-100 m, bivalve-echinoderm; 100-50 m, bryozoan-bivalve-gastropod-serpulid; <50 m, calcareous red algae-bivalve-gastropod-barnacle.  相似文献   

15.
Peat deposits are comprised of high organic content substances primarily derived from dead plant vegetation. Peat itself is not inert but undergoes continuous biological decomposition that causes progressive destruction of the peat fabric, reductions in fibre and organic contents and biogas generation. Depending on the degree of decomposition, the organic solids can exist as fresh (intact) fibres, slightly decomposed or ultimately completely decomposed (amorphous) material. From a geotechnical perspective, an understanding of the relationship between degree of decomposition and engineering properties, including the level of compressibility, is important in dealing with such problematic deposits. However a review of the literature indicates that such relationships have not been sufficiently investigated. Moreover, potential impacts of uncontrolled or unexpected decomposition in-situ are regularly discounted in geotechnical practice. This paper reviews decomposition effects in peat and potentially significant knock-on effects in terms of the material’s physical properties and compressibility. Progressive reduction in solids volume and deterioration in the integrity of the organic structure due to on-going decomposition may cause significant additional settlement to occur over time. More decomposed peat generally undergoes lower primary consolidation and creep strains and is also less prone to future decomposition, compared with lesser decomposed peat.  相似文献   

16.
The cathodoluminescence of low grade metamorphic quartz is reported in the literature to be uniformly brown. This study describes an investigation of syntectonic quartz fibres from veins and pressure shadows contained in different carbonate units of the Helvetic Alps, Switzerland, and of Anglesey, U.K., with the Scanning Electron Microscope in the cathodoluminescence mode. The cathodoluminescence spectra measured show systematic variations between older and younger parts of the same fibre, and between fibres from different tectonic units. These variations can be related first to variations in the temperature of formation of the quartz fibres, and second to deformation-induced differences in the microstructure of the earlier formed parts of fibres during later imposed strains.  相似文献   

17.
On- and off-mound sediment cores from Propeller Mound (Hovland Mound province, Porcupine Seabight) were analysed to understand better the evolution of a carbonate mound. The evaluation of benthic foraminiferal assemblages from the off-mound position helps to determine the changes of the environmental controls on Propeller Mound in glacial and interglacial times. Two different assemblages describe the Holocene and Marine Isotope Stage (MIS) 2 and late MIS 3 (∼31 kyr BP). The different assemblages are related to changes in oceanographic conditions, surface productivity and the waxing and waning of the British Irish Ice Sheet (BIIS) during the last glacial stages. The interglacial assemblage is related to a higher supply of organic material and stronger current intensities in water depth of recent coral growth. During the last glaciation the benthic faunas showed high abundances of cassidulinid species, implying cold bottom waters and a reduced availability of organic matter. High sedimentation rates and the domination of Elphidium excavatum point to shelf erosion related to sea-level lowering (∼50 m) and the progradation of the BIIS onto the shelf. A different assemblage described for the on-mound core is dominated by Discanomalina coronata, Gavelinopsis translucens, Planulina ariminensis, Cibicides lobatulus and to a lower degree by Hyrrokkin sarcophaga. These species are only found or show significantly higher relative abundances in on-mound samples and their maximum contribution in the lower part of the record indicates a higher coral growth density on Propeller Mound in an earlier period. They are less abundant during the Holocene, however. This dataset portrays the boundary conditions of the habitable range for the cold-water coral Lophelia pertusa, which dominates the deep-water reefal ecosystem on the upper flanks of Propeller Mound. The growth of this ecosystem occurs during interglacial and interstadial periods, whereas a retreat of corals is documented in the absence of glacial sediments on-mound. Glacial conditions with cold intermediate waters, a weak current regime and high sedimentation rates provide an unfavourable environmental setting for Lophelia corals to grow. A Late Pleistocene decrease is observed in the mound growth for Propeller Mound, which might face its complete burial in the future, as it already happened to the buried mounds of the Magellan Mound province further north.  相似文献   

18.
Antitaxial non‐deforming strain fringes from Lourdes, France, show complex quartz, calcite and chlorite fibre patterns that grew around pyrite in a slate during non‐coaxial progressive deformation. Development of these fringes was modelled using a computer program ‘Fringe Growth 2.0’ which can simulate incremental growth of crystal fibres around core‐objects of variable shape. It uses object‐centre paths as input, which are obtained from fibre patterns in thin section. The numerical experiments produced fibre patterns that show complex intergrowth of displacement‐controlled, face‐controlled and intermediate fibres similar to those in the natural examples. The direction of displacement‐controlled growth is only dependent on the relative movement between core‐object and fringe, so that core‐object rotation with respect to the fringe influences the fibre patterns and produces characteristic asymmetric fibre curvature. Object‐centre paths should be used for kinematic analysis of strain fringes instead of single fibres since these paths represent the fringe as a whole. The length along the path can be interpreted in terms of finite strain and path curvature in terms of rigid body rotation of fringes with respect to an external reference frame.  相似文献   

19.
In this study we have used laser ablation inductively coupled plasma mass spectrometry (LA-ICP-MS), to produce a high resolution coral record of rare earth elements (REE), Mn and Ba from coastal Porites corals from the Great Barrier Reef of Australia. Validation of the LA-ICP-MS technique indicated that the method provides accurate and reproducible (RSD = 13-18%) analysis of low concentration REE in corals (∼1 to 100 ppb). The REE composition in coral samples was found to closely reflect that of the surrounding seawater and distribution coefficients of ∼1-2 indicated minimal fractionation of the series during incorporation into coral carbonate. To explore the idea that coral records of REE can be used to investigate dissolved seawater composition, we analyzed two coastal corals representing a total of ∼30 yr of growth, including a 10-yr overlapping period. Comparable results were obtained from the two samples, particularly in terms of elemental ratios (Nd/Yb) and the Ce anomaly. Based on this evidence and results from the determination of distribution coefficients, we suggest that useful records of seawater REE composition can be obtained from coral carbonates. When compared to the REE composition of a mid shelf coral, coastal corals showed a significant terrestrial influence, characterized by higher REE concentrations (greater than 10 times) and light REE enrichment. The REE composition of coastal seawater inferred from the coral record was dependent on seasonal factors and the influence of flood waters. REE fractionation displayed a strong seasonal cycle that correlated closely with Mn concentration. We suggest that higher Nd/Yb ratios and higher Mn concentrations in summer result from scavenging of heavy REE by particulate organic ligands and Mn reductive dissolution respectively, both processes displaying higher rates during periods of high primary productivity. The Ce anomaly also displayed a strong seasonal cycle showing an enhanced anomaly during summer and during flood events. This is consistent with the Ce anomaly being primarily controlled by the abundance of Ce oxidizing bacteria. Based on these arguments, we suggest that the coral record of dissolved REE and Mn may be regarded as a useful proxy for biological activity in coastal seawater.  相似文献   

20.
Hurricane control on shelf-edge-reef architecture around Grand Cayman   总被引：3，自引：0，他引：3  
Rimming the outer shelf of Grand Cayman is a submerged, 87 km long shelf-edge reef that rises to within 12 m of mean sea level. It consists of an array of coral-armoured buttresses aligned perpendicular to shore and separated by steep-sided sediment-floored canyons. Individual buttresses have a diverse coral-dominated biota and consist of three architectural elements: a shield-like front wall colonized by platy corals, a dome-shaped crown colonized by head corals, and a shoreward-projecting spur covered by varying amounts of branching coral. Buttresses are commonly fronted by coral pinnacles that, in some areas, have amalgamated with buttress walls to produce pinnacle-and-arch structures. As margin orientation changes, shelf-edge-reef architecture shows systematic variations that are consistent with changes in fetch and height of hurricane waves. Along margins exposed to fully developed storm waves, shelf-edge-reef buttresses are deep, have large amplitudes, and are dominated by robust head corals. These characteristics are consistent with hurricane-induced pruning of branching corals and the flushing of significant quantities of sand from buttress canyons by return flows. Along margins impacted by fetch-limited storm waves, reef buttresses are shallower, have intermediate-amplitudes, and have a significantly higher proportion of branching corals. These characteristics are consistent with less coral pruning and sand flushing by weaker hurricane waves. Along margins fully protected from storm waves, the buttresses-canyon architecture of the shelf-edge reef breaks down producing a series of shallow, undulating, branching-coral-dominated ridges that merge laterally into an unbroken belt of coral. These characteristics correspond with negligible amounts of pruning and flushing during hurricanes. In addition to differences between margins, local intra-marginal changes in shelf-edge reef architecture are consistent with changes in the angle of hurricane-wave approach. Open sections of the shelf-edge reef, which face directly into storm waves, are pruned of branching corals and the fragments swept back onto the shelf producing extensive spurs. By contrast, on more sheltered, obliquely orientated sections, storm-waves sweep debris along and off shelf producing little or no spur development. Instead, the debris shed seawards accumulates in front of the buttress walls and initiates the development of coral pinnacles. Over time, repeated buttress pruning and canyon flushing during hurricanes not only controls reef architecture but may also influence accretion patterns. Vertical accretion is limited by the effective depth of storm-wave fragmentation. Once this hurricane-accretion threshold is reached the reef moves into a shedding phase and accretes laterally via pinnacle growth, amalgamation, and infilling. Consequently, the reef steps out over its own debris in a kind of balancing act between lateral growth and slope failure — a pattern widely recognized in ancient reefs.  相似文献   

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