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1.
An isolated jaw fragment from the Late Cretaceous (Cenomanian) Cambridge Greensand Member of the West Melbury Marly Chalk Formation previously identified as a cestraciontid shark fin spine is referred to the pterosaur clade Azhdarchoidea on account of its lateral and occlusal foramina and edentuly. The specimen differs from the azhdarchoid Ornithostoma sedgwicki from the same deposit in having flat lateral surfaces and an acute dorsal/ventral apex. The specimen is similar in overall morphology to CAMSM B40085 from the same horizon and probably represents the corresponding jaw but from a different individual. Likely these specimens represent a new taxon but are considered too fragmentary to diagnose at present. A remarkably similar and distinctive morphology is found in unnamed pterosaur jaws from the Kem Kem Group (?Albian-Cenomanian) of Morocco, supporting the idea of faunal similarity between these two distant localities.  相似文献   

2.
Pterosaurs are a rare component of the Early Cretaceous (Albian) Gault Formation of southern England. The only named taxon reported, ‘Pterodactylusdaviesii Owen, 1874, is widely regarded as a nomen dubium or as Pterodactyloidea incertae sedis, while most other material can be referred to Pterodactyloidea indet. Here we describe a fragmentary humerus and elongate mid-series cervical vertebra both from the Gault Formation of Kent, southeast England that can be referred to the edentulous pterodactyloid clade Azhdarchoidea. The cervical vertebra is identified as being from a non-tapejaromorph azhdarchoid on account of its reduced neural spine, a neural arch confluent with the centrum, a neural canal that is subsumed into the centrum and the lack of foramina on the lateral surfaces of the centrum. The humerus is referred to Azhdarchoidea on account of its sub-rectangular distal end.  相似文献   

3.
The Late Cretaceous (Santonian) continental vertebrate locality of Iharkút, western Hungary has provided numerous azhdarchid pterosaur remains including the recently described, Bakonydraco galaczi. Since the first report of these fossils, additional remains have been discovered that improve considerably our knowledge of some aspects of the anatomy of azhdarchid pterosaurs. New cranial material described here indicates an edentulous, non-crested premaxillary rostrum in Bakonydraco similar to that of Quetzalcoatlus and reveals that this rostrum was considerably thinner and more lightly built than the relatively massive, pointed mandibular symphysis. In addition, the contact surface of the upper and lower jaws of Bakonydraco at least in the symphyseal region was more similar to Tapejara wellnhoferi than to other azdarchids: the contact is irregular and the premaxilla does not fit closely the rostral part of the mandible. Among the postcranial material the atlas-axis complex possessing lateral pneumatic foramina is of great importance because it further supports the notion of interspecific variability of vertebral pneumaticity among pterosaurs.  相似文献   

4.
A new specimen of pterosaurian metacarpal IV from the Early Cretaceous (Berriasian) Durlston Formation of Dorset, southern England, closely resembles those of the Chinese dsungaripterid Dsungaripterus weii Young, 1964 and the Central Asian azhdarchid Azhdarcho lancicollis Nessov, 1984. However, the new specimen lacks the thickened bony wall typical of Dsungaripteridae and is therefore regarded as azhdarchoid. An mcIV with a markedly offset distal condyle with well-developed diaphyseal constriction behind the distal condyle and asymmetrical condylar margins may be an autapomorphy of Dsungaripteridae + Azhdarchoidea, while the same condition with bone thickening may typify Dsungaripteridae. The new specimen also provides osteological evidence supporting claims for large wing-spanned pterosaurs in the Berriasian of southern England based previously only on ichnological finds. The new specimen suggests a wingspan of between 2.9 and 3.7 m and represents the largest pterosaur from the Berriasian.  相似文献   

5.
Seven coleoid jaws recovered from Santonian to lower Campanian (Upper Cretaceous) strata in Hokkaido, Japan were taxonomically studied. Based on the comparison with the jaws of modern and fossil coleoids, six of the seven jaw fossils are referred to the following two genera and three species, including one possible new species: Nanaimoteuthis jeletzkyi and N. yokotai of the order Vampyromorpha, and Paleocirroteuthis sp. nov. (?) of the order Cirroctopodida. The other single lower jaw is seemingly similar to those of modern octopods and teuthids with respect to the shape of the inner lamella, but its order-level assignment could not be determined because of its imperfect preservation. N. jeletzkyi has been described in the Upper Cretaceous fore-arc basin deposits in Hokkaido (Yezo Group) and Vancouver Island, Canada (Nanaimo Group), whereas N. yokotai occurs only in the Yezo Group. These findings, complemented by previous reports of coleoid jaws, gladii, and phragmocones from the Yezo and Nanaimo Groups, demonstrate that a highly diversified, non-belemnitid coleoid fauna including large teuthids had already appeared during the post-Albian Late Cretaceous, in the North Pacific region.  相似文献   

6.
A jaw bone attributed to the mid-Cretaceous azhdarchid pterosaur Alanqa saharica Ibrahim et al., 2010 is described. The new specimen from the Kem Kem beds of south eastern Morocco is unusual for the presence of expanded rostral bones that protrude above the occlusal surface of the jaws. The function of this unusual osteological modification is uncertain, but it is considered likely that it served a role in food processing, or perhaps jaw stability.  相似文献   

7.
姬书安  张立军  路芳 《地质学报》2023,97(6):1723-1740
中国辽宁西部早白垩世翼龙类化石非常丰富,其上部的九佛堂组翼龙类组合以含有丰富的无齿的古神翼龙科、朝阳翼龙科等进步类型而与下部的义县组翼龙类组合相区别。本文描述了辽宁西部建昌盆地九佛堂组一新的大型翼龙类化石,其以上下颌无齿、吻端指数为3.7、前颌骨背支细长且封闭鼻眶前孔背缘、很大的鼻眶前孔后端超过上下颌关节位置、轭骨的上颌骨突基部宽大而应被归入朝阳翼龙科(Chaoyangopteridae)神州翼龙属(Shenzhoupterus)。其以较大的体形(翼展2.05 m)、平直的上下颌咬合面、轭骨的眶后骨突较泪骨突长且基部略宽、第4~7颈椎长度依次减小等特征,区别于朝阳神州翼龙(Shenzhoupterus chaoyangensis),而被命名为三亚神州翼龙(新种)(Shenzhoupterus sanyainus sp. nov.)。这是辽宁西部早白垩世翼展最大且不具牙齿的翼龙类属种,它的发现丰富了九佛堂组翼龙类组合内容,对认识朝阳翼龙科的骨骼形态与生态习性亦具有积极意义。  相似文献   

8.
《China Geology》2020,3(1):1-7
A new dsungaripterid pterosaur, Ordosipterus planignathus gen. et sp. nov., is established on the incomplete articulated lower jaws from the Lower Cretaceous Luohandong Formation in Otog Qi, Ordos Region, Inner Mongolia, China. It differs from other dsungaripterids mainly by having broad and low dentary at and just behind the mandibular symphysis, flat dentary dorsal plane forming the distinct lateral ridge with the curved dentary lateral side, and lower alveoli arranged along the dentary dorsolateral margin with wide spacing that increases from rostral to caudal. It represents the first diagnostic pterosaur from the Ordos Region in Inner Mongolia, and further enlarges the geographical distribution of the family Dsungaripteridae from northwestern China (together with western Mongolia) to central North China.  相似文献   

9.
Cretornis hlavaci Frič, 1881 from the Upper Cretaceous (Turonian) of Czech Republic is a valid taxon referred to Azhdarchoidea based on having a saddle-shaped humeral head, pneumatic foramen on proximal humerus present on anterior side and absent on posterior side, elongate deltopectoral crest with subparallel proximal and distal margins, pneumatic foramen absent on distal side of humerus, metacarpals I–III not articulated with carpus and displaced on anterodorsal side of wing metacarpal, and wing metacarpal much longer than humerus. Absence of a pneumatic foramen on posterior side of proximal humerus suggests attribution of Cretornis hlavaci to Neoazhdarchia. It has a unique construction of the distal ulna with a dorsal articulation surface placed distinctly proximal to the tuberculum shared only with the non-azhdarchid azhdarchoid Montanazhdarcho minor from the Campanian of North America. Cretornis hlavaci differs from the latter taxon by the structure of its humerus and distinctly longer wing metacarpal. It is more derived than “Tapejaridae” but shares with Azhdarchidae the deltopectoral crest of the humerus displaced distally from humeral head. Cretornis cannot be assigned to Azhdarchidae because of the oval cross section of the second wing phalanx. A unique rhombic outline of the distal humerus of Cretornis hlavaci is a possible autapomorphy for this taxon. Its wing span estimated as 1.5–1.6 m. This is the first taxon of non-azhdarchid pterosaurs known from the Upper Cretaceous of the Eastern Hemisphere.  相似文献   

10.
Three pieces from cervical half-rings of an immature nodosaur, part of a nodosaurid presacral rod and some post-cranial osteoderms from the Cretaceous of Cambridge were studied at the Booth Museum of Natural History, Brighton, UK. Two of the three half-ring elements show dorsal ridge morphologies distinct from each other, and all three have unfused sutured lateral borders. It is possible they may be derived from the same animal. Comparison with other material from the Cretaceous of Europe, USA and Asia indicates the presence of a large nodosaurid in the Cambridge Greensand fauna, with cervical half-ring morphologies similar to North American taxa, but unlike any previously known from the European Cretaceous.  相似文献   

11.
A new ctenochasmatoid pterosaur, Gladocephaloideus jingangshanensis gen. et sp. nov. from the Yixian Formation of western Laioning Province is erected based on a complete skull and partial postcranial skeleton. It is characterized by following features: about 50 total teeth with sharp tips; small nasoantorbital opening, occupying approximately 13% of the skull length; ratio of prenarial length to skull length approximately 0.63. The diagnoses of the Ctenochasmatoidea and Gallodactylidae are amended based on the new taxon. Gladocephaloideus jingangshanensis is the first gallodactylid pterosaur found in Asia. Its discovery not only provides much more osteological information about the Gallodactylidae but also indicates that the ctenochasmatoid pterosaurs were highly diverse in the Early Cretaceous. The filamentous structures preserved near the dorsal and posterior margins of the posterior portion of the skull and around the neck indicate that it had an epidermal covering and may have been a warm-blooded animal.  相似文献   

12.
Ceratopsians were herbivorous dinosaurs that dominated many of the terrestrial ecosystems in Asia and North America during the Cretaceous. The bizarre variety of skulls and lower jaw morphologies as well as the inferred ecological abundance of many species in this clade indicate that Ceratopsia was a successful group. Here we analyzed 126 lower jaws from 50 ceratopsian species, using two-dimensional geometric morphometrics and finite element analysis to investigate differences in shape and structural performance of this part of the feeding apparatus across Ceratopsia. Morphological differences in lower jaws across ceratopsian clades are said to originate from feeding adaptation.Our results show that the stress (physical loadings modeled in response to biting) in lower jaws was quite similar between “basal” and “derived” taxa, whereas major differences among clades occur for stress values associated with the coronoid process. The basal ceratopsians Hualianceratops and Yinlong had a highly stressed and primitive lower jaw, indicating that those animals may have fed on relatively soft foliage and fruits. A similar condition was found for basal neoceratopsians and protoceratopsids. Psittacosaurids possessed a well-integrated and compact lower jaw able to withstand high stress, at the cost of having a highly stressed coronoid process. Leptoceratopsids were characterized by the opposite condition. Taxa such as Leptoceratops, Prenoceratops, Zhuchengceratops and Cerasinops appear to have had a comparatively efficient feeding apparatus. Ceratopsidae represents the clade with the most efficient masticatory apparatus within Ceratopsia, even if the horizontal ramus of the lower jaw appears less able to withstand high levels of stress as compared with other ceratopsians. Additionally, we found the dentary and surangular–angular complex co-evolved to generate a masticatory apparatus able to withstand high stress, particularly in Protoceratopsidae and Triceratopsini. The major phenotypic evolutionary rate and morphological changes occurred during the mid- to Late Cretaceous, when intense climate change and angiosperm diversification could have affected the evolution of ecological diversity and feeding biomechanics in Ceratopsia.  相似文献   

13.
Early Cretaceous sediments of Aptian–Albian age outcrop at Munday’s Hill Quarry, Bedfordshire, England. Previous papers describing the section have resulted in different terminologies being applied. The Lower Cretaceous in Bedfordshire is represented by sediments belonging to the Lower Greensand Group and the Gault Clay Formation. Within the Lower Greensand Group in the study area the Woburn Sands Formation, are of Aptian–Albian age. Selected samples have been analysed for palynology. The analysis reveals diverse palynomorph assemblages, including well-preserved dinoflagellate cysts and sporomorphs. Comparison of the assemblages with published records indicates that the lower samples are of Late Aptian age. Forms recorded include common Kiokansium unituberculatum, Cerbia tabulata, Aptea polymorpha and Cyclonephelium inconspicuum. An Early Albian age is indicated for the uppermost sample.  相似文献   

14.
Nineteenth-century references to clavate borings in woody substrates in the Lower Greensand of the Isle of Wight used a variety of names, but Teredo (a wood-boring bivalve, not a boring), Teredolithes (a junior synonym of Teredolites) and Gastrochaena (a bivalve borer of rock and shelly substrates, not a boring in wood) are all nomenclatorially incorrect. Borings in a beach clast derived from the Lower Greensand Group and recently collected from Sandown Bay, Isle of Wight, are referred to Teredolites isp. cf. T. longissimus Kelly and Bromley. This specimen confirms the presence of Teredolites in the Lower Greensand Group and demonstrates a common ichnological problem of beach clasts; borings, either fossil or modern, are incompletely preserved, making confident classification below the level of ichnogenus problematic.  相似文献   

15.
Fragmentary remains, including cervical vertebrae and limb bones, of a large pterosaur from the Upper Cretaceous of Tous, province of Valencia (Spain), are described. The material was recovered from lacustrine beds in the upper part of the Calizas y Margas de Sierra Perenchiza Formation, which is probably Maastrichtian in age. Six fragments of vertebrae allow a reconstruction of the anatomy of the mid-series cervicals of the animal. The general morphology of the cervical vertebrae is closely similar to that of the long-necked Azhdarchidae. Compared to other azhdarchids, the Valencia pterosaur shows minor differences from the genera Azhdarcho and Quetzalcoatlus, and is here provisionally referred to as Azhdarchidae indet. A wingspan of about 5.5 m is calculated by comparison with other known azhdarchids. This is the second azhdarchid pterosaur described from the Iberian peninsula. It confirms the wide distribution of this group of large pterosaurs at the end of the Cretaceous.  相似文献   

16.
The Upper Cretaceous of Africa has produced a diverse fauna of mosasaurs, including the highly specialized, long-jawed Pluridens. The type of Pluridens walkeri comes from the Maastrichtian Farin-Doutchi Formation of Niger, with a second, referred specimen coming from the Campanian section of the Campanian-Maastrichtian Nkporo Shale near Calabar, in southern Nigeria. Comparisons of this referred specimen with the holotype suggest that it represents a distinct and more primitive species. The Calabar jaw resembles P. walkeri in being long and narrow anteriorly with a shallow subdental shelf, and in having small, numerous, recurved teeth with medially positioned replacement pits. However, it lacks many of the derived features that characterize Pluridens walkeri, such as the extremely long and straight jaw, the extreme lateral protrusion and subcircular section of the dentary, strong transverse expansion of the dental thecae, and extreme reduction and increase in number of the teeth. The Calabar Pluridens is therefore referred to a new species, Pluridens calabaria. Following recent studies, Pluridens is considered to represent a highly derived member of the Halisaurinae. The marked differences between the Campanian and Maastrichtian species of the genus underscore the rapid pace of mosasaur evolution during the Cretaceous.  相似文献   

17.
The Mendip Hills, located on the north-western margin of the Wessex Basin, clearly show the onlap of Upper Triassic to Middle Jurassic sediments onto folded Palaeozoic strata. Recent field mapping on the crest of the Beacon Hill pericline at Tadhill, near Frome, augmented by a suite of shallow boreholes, proved up to 6.2 m of glauconitic grey and green silty sand. These glauconitic sands rest unconformably on Silurian volcanic rocks and Devonian sandstone. Lithological and ipalaeontological analyses of these glauconitic sands indicate that they are part of the Lower Cretaceous Upper Greensand Formation. This provides the first evidence for the Albian transgression across the Mendip Hills. The implications for the Cretaceous overstep on the margins of the Wessex Basin, and the analogies with the Upper Greensand succession in Devon are discussed.  相似文献   

18.
A new pterosaur, Microtuban altivolans gen. et sp. nov., is described from the Sannine Formation of northern Lebanon. The specimen is the first pterosaur from the Early Cenomanian (Late Cretaceous) locality of Hjoûla and is regarded as the most complete pterosaur fossil discovered from Africa. While postcranial characters indicate a possible relationship with members of the Thalassodromidae or Chaoyangopteridae, the specimen possesses an exceptionally short wing-finger phalanx 4, forming only 1.1% of the total length of the wing-finger. Its appearance along with an unnamed ornithocheiroid from the slightly younger locality of Hâqel suggests that a number of pterosaur taxa existed within the local area, perhaps living on exposed carbonate platforms.  相似文献   

19.
Post-Wealden dinosaur remains are rare in the UK, so any material from late Early or Late Cretaceous deposits is potentially of palaeoecological and palaeobiogeographical significance. Four dinosaur specimens collected from the Woburn Sands Formation (Aptian) of Upware, Cambridgeshire were described briefly by Walter Keeping in 1883, but have not been reappraised since. These specimens are identified herein as a ?turiasaurian sauropod tooth and indeterminate iguanodontian ornithopod remains (a tooth, middle caudal vertebra, pollex ungual). Although collected from the Woburn Sands Formation, it is likely that all of these fossils were reworked from older (now absent) sediments and they have usually been regarded as either ‘Wealden’ or Neocomian in age, presumably due to previous identifications of some of these specimens as Iguanodon. However, consideration of UK dinosaur faunas and regional geology indicates that these fossils could potentially be older. Further work is needed on the derived terrestrial fossils of the Lower Greensand Group in order to constrain their ages more precisely so that they can be incorporated into broader studies of regional diversity and palaeoecology.  相似文献   

20.
A new pterosaur Moganopterus zhuiana gen. et sp. nov. is erected based on a complete skull with lower jaws and anterior cervical vertebrae. It is characterized by much elongated upper and lower jaws with at least 62 total, long, curved teeth with sharp tips, a well developed parietal crest extending posterodorsally, forming an angle of 15 degrees with the ventral margin of the skull, the ratio of length to width of cervical vertebrae greater than 5:1. The skull length is 750 mm, and it is the largest toothed pterosaur found so far in the world. Based on this new pterosaur, the Boreopteridae can be divided into two subgroups: Boreopterinae sub-fam. nov. and Moganopterinae sub-fam. nov., which is also confirmed by the phylogenetic analysis.  相似文献   

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