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1.
Three-week old soybean (Glycine max) plants were subjected to a factorial combination of four regimes of soil matric water potential (ψm=−0·03, −0·5, −1·0 and −1·5 MPa), two levels of supplementary Zn (O and 20 mgl−1) and two levels of foliar IAA application (O and 10 mgl−1). Under control conditions (no Zn, no IAA), increasing soil drying progressively retarded shoot and root growth (length and dry mass production), reduced leaf relative water content (RWC) and decreased the contents of chlorophyll (Chl) and shoot soluble sugars (SS), but increased soluble sugar content of roots and lowered osmotic water potential of shoots and roots (osmotic adjustment). Total free amino acid (TAA) content increased in shoots but decreased in roots whereas contents of soluble proteins (SP) decreased in shoots and roots. The effect of water stress was statistically significant (p<0·05) and had a major effect (as indicated by η2values) on leaf RWC, shoot and root dry masses and osmotic potential. Supplementary Zn improved root growth at all levels of stress and shoot growth under severe stress. Improvement of growth was positively correlated with the internal tissue Zn concentrations (r=0·91 and 0·86 for shoot and 0·94 and 0·82 for root length and dry mass respectively). Exogenous IAA raised (p<0·05) RWC, Chl, DM (slightly), root SS, and SP, whereas shoot TAA was lowered. Effects on root TAA and shoot SS were more complex: they were lowered at zero stress and raised under severe stress. IAA and Zn in combination had additive effects on Chl, growth and osmotic potential, but their combined effects on SP and TAA were more complex. It is concluded that the treatment of soybean plants grown under conditions of low soil water potentials and Zn deficiency with Zn and IAA solutions counteracted the deleterious effects of stress, especially at high stress levels, and helped stressed plants to grow successfully under these adverse unfavourable conditions.  相似文献   

2.
The natural abundance15N/14N method was used to estimate the influence of silvicultural and P fertilization treatments on N accretion, N2fixation and N partitioning among tissues in a mature mesquiteProsopis glandulosavar.glandulosastand in Texas. The silvicultural treatments consisted of understory removal, herbicide treatment of brushy resprouts, thinning trees to single stems and 100 kg ha−1P fertilization. The trees had a mean basal diameter of 17·8 cm with 8 to 35 cm range. The stand was slow growing with the increase in dry matter ranging from 0·465 Mg ha−1year−1to 0·701 Mg ha−1year−1for the 8 years after the treatments were applied. N accretion after 8 years ranged from 3·1 kg ha−1year−1to 4·4 kg ha−1year−1.Due to the range in δ15N of the leaves, twigs, branches and trunk, we used the weighted (by biomass) average δ15N per tree in calculations of the percent N derived from N2fixation (%Ndfa). There was considerable variability in δ15N of the reference plants, i.e. from 3·3 to 5·9. In contrast there was low variability in the background δ15N of nearby soils (7·0±1·0). As the total above-ground biomass δ15N of a grass grown outside the influence of mesquite (7·8±0·58) had the same δ15N as the soil (7·5±1·0), we used the grass outside the influence of mesquite and the weighted tree mean δ15N to calculate % of N derived from N2fixation.The decrease in intraspecific competition by thinning multistemed trees to single stemmed trees was the only treatment that significantly (p= 0·0001) increased growth. Interspecific competition, i.e. understory removal, did not increase growth. There were no significant differences in total N production or N fixation among treatment means. The most striking result was the highly positive correlation between tree δ15N and total N per tree and biomass per tree (R2= 0·90,F= 164·4, df. = 18, mean square error (MSE) = 0·155,p= 0·0001). This implies that the younger trees colonizing infertile soils relied more heavily on N2fixation than larger trees which accumulated 1200 kg ha−1more N under their canopies. The percentage N derived from N2fixation ranged from 63 to 73% in the various treatments. Despite the high percentage of N derived from N2fixation, the N2fixation of the stand was very low, i.e. 1·98 to 2·80 kg N ha−1year−1, due to the low growth of the stand. We believe that comparisons of the whole tree weighted δ15N to background soil δ15N provides a more reasonable approach to estimate % N2fixation than comparisons of leaves of fixers and reference plants.  相似文献   

3.
We studied the relationship between primary productivity and species richness of small mammals at both large and small spatial scales in the arid and semi-arid grasslands of north China. The productivity (x)–species richness (y) pattern at a large spatial scale can be described by a unimodal quadratic regression curve (y=7·41+0·1*x−0·0003*x2,p =0·008, r2=0·788). At a small spatial scale, however, neither linear nor quadratic regression fit the data for 1980 and 1994 (p>0·25). Primary productivity may not be an appropriate predictor of the species richness of small mammals at a small spatial scale. We conclude, therefore, that the primary productivity–species richness pattern of small mammals may be scale-dependent in the arid and semi-arid grasslands of north China. Landscape complexity should be considered in future studies of productivity–richness relationships.  相似文献   

4.
An experiment was carried out at Nahshala Farm, north west of Al-Ain City, U.A.E. during the 1998–2000 growing seasons, using six halophytes (Batis maritima, Distichlis spicata, Juncus roemerianus, Paspalum vaginatum, Salicornia bigelovii and Spartina alterniflora) and two levels of leaching fraction (0·25 and 0·50) under three irrigation salinity levels (10, 20, and 40 gL−1) in a randomized complete block design arranged in split plots. The purpose of the research was to apply the theory of crop salt tolerance on aboveground yields and agronomic characteristics of halophytes to determine their thresholds for salinity and to determine leaching requirements. The results indicated that the halophyte species tested can grow with minimum reduction in the growth potential at <20 gL−1of mean salinity of soil solution. Leaching fraction (LF) of 0·25 at the highest salinity of irrigation water (40 gL−1) was inadequate to attain the steady-state salt balance during the growth period, although the drainage salinity reached more than 90 g L−1. Furthermore, if the same level of LF is used for longer period, soil salinity under this high salt treatment will continue to rise, and plant growth may deteriorate. Leaching fraction of 0·50 is preferable if salinity of irrigation water was more than 20 gL−1and dry matter production is considered, although the amount of water use will be excessive  相似文献   

5.
The migration of a small slipfaceless dome dune close to the northern edge of the Namib Sand Sea has been measured by topographic survey. The dune dimensions are 45-m wide and 1-m high with a volume of 551 m3 that has been calculated as the difference between the dune's surface elevation and an interdune surface extrapolated from measurements around the edge of the dune. The migration direction, 64°, and distance moved, 90 m, are measured against stakes set out in 1976. The dune has moved about 90 m between 1976 and 1999. This is an average linear migration rate of around 4 m year−1, and is equivalent to an annual sand transport rate of about 1.2 tonnes m−1 year−1. The calculated total potential sand flow in this part of the Namib Desert is 119 tonnes m−1 year−1, and the resultant potential sand flow is 63 tonnes m−1 year−1. The dune migration is therefore about 1% of the total potential sandflow and 2% of the resultant indicating that dune migration is only a small part of total potential sand transport. The results suggest that small slipfaceless dome dunes are very inefficient at trapping sand, and that winds blowing across the interdune in this area are undersaturated with sand.  相似文献   

6.
We investigated the size fraction and pigment-derived class compositions of phytoplankton within the euphotic zone of the Antarctic marginal ice zone between 63.3°S and 66.5°S along the 140°E meridian on two consecutive cruises in the late austral summer and early austral autumn of 2003. We observed significant temporal and spatial variations in phytoplankton size and taxonomic composition, although chlorophyll a concentrations were generally below 1 μg l−1 during both periods. Microphytoplankton (>20 μm), mainly diatoms, were prominent in the euphotic zone in the southernmost area around 66.5°S during late summer. In the rest of the study area during both cruises, the phytoplankton community was dominated by pico- and nano-sized populations (<20 μm) throughout the euphotic zone. The small-size populations mostly consisted of diatoms and haptophytes, although chlorophytes were dominant in extremely cold water (−1.5°C) below the overlying warm water around 65.5°S during late summer. From late summer to early autumn, chlorophytes declined in abundance, probably due to increasing temperature within the euphotic zone (−1 to 0°C). These pico- and nano-phytoplankton-dominated populations were often accompanied by relatively high concentrations of ammonium, suggesting the active regeneration of nutrients within the small-size plankton community.  相似文献   

7.
This paper presents a detailed cost accounting of the processes involved in the production of flours from the mesocarps of the fruits of Prosopis alba and P. pallida to be used in human foods. The flours from both species are similar and contain 7–8% protein, 0·8–2% fat, 26–32% total dietary fiber and 48–59% sucrose. The data were based on actual production rates in use at the Universidad de Piura pilot plant using simple machinery. Machinery hourly charges were based on equivalent rental costs in order to avoid interest charge calculations. As opposed to other analyses which only included processing costs delivered to the plant, our analysis ascribed value to the owner of the trees in order to provide incentives to plant and care for the trees. The sensitivity analyses showed that the most economically important factor was the conversion efficiency of pods into flour. When a 40% conversion efficiency, such as that which has been obtained in both Peru and Argentina was obtained, and by-product credits for high fiber containing fraction were assumed, the wholesale selling price with a 5% marketing cost and a 40% profit direct costs was estimated to be $0·66 kg−1 for Peru (using a $0·40 h−1 labor rate ) and $1·76 kg−1 for Argentina (using a $1·5 h−1 labor rate).  相似文献   

8.
A portable wind tunnel was used to test the contribution of biological and physical elements to overall soil aggregation on a soil dominated by biological soil crusts in south-eastern Australia. After moderate disturbance and simulated wind erosion, 90% of surface aggregates on the loamy soil and 76% on the sandy soil were dominated by biological elements (cryptogams). Lower levels of biological bonding were observed on the severely disturbed treatment. Linear regression indicated a significant positive relationship (r2=0·72) between biological soil crust cover and dry aggregation levels greater than 0·85mm. To maintain sediment transport below an erosion control target of 5gm−1s−1 for a 65kmh−1 wind at 10m height, a crust cover of approximately 20% is required. When a multiple regression model which sequentially fitted biological crust cover and dry aggregation greater than 0·85mm was applied to the data, dry aggregation accounted for more of the variation in sediment transport rate than biological crust cover. These data were used to develop a conceptual model which integrates crust cover and dry aggregation, and provides a useful framework within which to predict the likely impacts of changes in soil crust cover and aggregation.  相似文献   

9.
The forage and oilseed halophyte,Salicornia bigeloviiTorr., was grown in gravity-drained lysimeters set in open plots of the same crop over two seasons in a coastal desert environment in Sonora, Mexico. The lysimeters were irrigated daily with seawater (40 g l−1salts) at rates ranging from 46–225% of potential evaporation. Biomass and seed yields increased with increasing irrigation depth over the range of treatments. Biomass yields ranged from 13·6–23·1 t DM ha−1, equivalent to conventional forage crops, on seasonal water application depths of 2·3–3·8 m, but were markedly lower at lower irrigation depths. Increasing the irrigation depth lowered the soil solution salinity, resulting in greater growth and water use, and hence leaching fractions that were nearly even over irrigation treatments, averaging 0·5. Evapo-transpiration rose in direct proportion to the irrigation depth. Potential evaporation was estimated by site pan evaporation and by the Blaney-Criddle and Penman models using climatological data; the methods agreed within 15%. The ratio of evapo-transpiration to potential evaporation increased over the growing season and approached 1·5 by pan on the highest irrigation treatment due to the combined effects of high transpiration and high evaporation from the permanently moist soil surface. The best field predictor of biomass yield was the salinity of the soil moisture in the top 15 cm of soil profile, which constitutes the root zone for this crop. Root zone salinity must be kept at 70–75 g l−1for high yields. Although irrigation and drainage requirements were high compared to conventional crops, seawater irrigation appears to be feasible in medium sand and could augment crop production along coastal deserts. The possibility of using this crop for animal production is discussed.  相似文献   

10.
The objectives of this study were to determine the composition, density and spatial distribution of the soil seed bank of woody species, as well as their regeneration pattern in two different land use systems, controlled (ranch) and open grazing, in an Acacia woodland of the Rift Valley in Ethiopia. We also compared the species composition of the soil seed bank and the above-ground vegetation to find out if differences exist in the soil seed bank and advance regeneration between the two land use systems. The germination requirements of seeds of the woody species were also investigated under laboratory conditions. Acacia senegal, Acacia seyal, Acacia tortilis, Dichrostacys cinerea and Balanites aegyptiaca were encountered in the above-ground vegetation in both systems. Seeds of only A. tortilis (90±32 seeds m−2) from the ranch, and A. senegal (5±3 seeds m−2) and A. tortilis (72±34 seeds m−2) from the open system were found in the soil seed bank along transects with patchy horizontal pattern. The two systems were not significantly different in density of soil seed banks of A. senegal andA. tortilis . Jaccard's similarity index showed that only a few woody species were common in the soil seed flora and above ground vegetation. However, all of the species accumulated seeds (58±43–331±130 seeds m−2) in/on the soil under the canopy. Very large numbers of seeds of A. tortilis (19382±9722 seeds m−2) and D. cinerea (1278±494 seeds m−2) were also found in barns. Most of the seeds recovered from the soil samples (60–80%) were found in the litter layer. Acid and mechanical scarification improved legume germination (36–99% and 60–99%, respectively) over boiling water (0–48%). Treatment means differed significantly for all the legumes (p<0·001) but not for Balanites. Height and diameter class distribution of regeneration of A. tortilis and D. cinerea in both systems and A. senegal in the open system had a negative exponential correlation (rs=−0·5, −0·25 and −0·86, respectively). A. seyal and B. aegyptiaca showed poor regeneration. Horizontal distribution of advance regeneration of all the species was patchy. Advance regeneration of A. seyal, A. tortilis and B. aegyptiaca were not significantly different, while that of A. senegal and D. cinerea were significantly different between the two systems. Poor representation of species in the soil seed bank along transects and in the different height and diameter classes may be attributed to the low density of mature trees as well as the mode and strategy of seed dispersal. Ungulate and wind dispersed species (e.g. A. senegal, A. tortilis and D. cinerea) were highly favoured. Patchiness in the distribution of seeds and advance regeneration was also a result of endozoochory. Dispersal of non-ungulate dispersed seeds (A. senegal, A. seyal and B. aegyptiaca) was restricted to the canopy zone. Piles of seeds ofA. tortilis and D. cinerea that were found in barns were a result of consumption of their pods by cattle. High concentration of seeds in the litter layer may be due to low soil disturbance and larger size of seeds. The height and diameter class distribution of A. senegal (in the open system), A. tortilis and D. cinerea also indicated that the species have good regeneration. Results from the germination tests indicated that seeds of the legumes require pre-sowing treatments to give a rapid, uniform and improved germination. Intervention through artificial regeneration should be employed to improve the density and regeneration capacity of those species with hampered regeneration at both systems.  相似文献   

11.
We investigated the effects of soil water potential (SWP) and temperature on seed germination of six coexisting species of an inter-tropical desert. These species include three life-forms; the shrubs Cercidium praecox andProsopis laevigata ; the columnar succulents Neobuxbaumia tetetzo and Pachycereus hollianus; and the arborescent semi-succulents Beaucarnea gracilis and Yucca periculosa. In the six species germinability increased and germination time and speed of germination (t50) decreased as SWP decreased. The SWP treatments were 0 MPa, −0·12 MPa, −0·2 MPa, −0·41 MPa and −0·66 MPa. There was, however, a SWP threshold below which germination time and t50increased and germinability decreased. The shrubs had the highest germinability whereas the columnar succulents had the lowest. The shrubs also had shorter germination time andt50 than arborescent semi-succulents whereas seeds of the columnar succulents were indeterminate. In all species except P. laevigata germinability increased and the germination time and t50decreased as temperature increased. The temperature treatments were 12°C, 20°C and 26°C. The shrubs had the shortest t50and germination time and the highest germinability at all temperatures. Arborescent semi-succulents had the lowest germinability and longest germination time and t50at the three temperatures treatments. Our results support the hypothesis that in desert environments different plant life-forms utilise different germination strategies to persist.  相似文献   

12.
The selective use by cattle of range resources in the Sahel was assessed in terms of quantity and nutritional quality of the herbage grazed and diet selected. Peak standing crop was 1951 kg ha−1dry matter (DM) with 47·9 kg nitrogen (N) ha−1and 7·4 kg phosphorus (P) ha−1. Esophageally fistulated cattle diets were consistently (p<0·05) higher in N, P and organic matter digestibility (OMD) than the herbage grazed. Cattle consumed 48% of the mean annual herbage production. The results indicate grazing ruminants select a higher quality diet from Sahelian rangelands than can be predicted on the basis of pasture evaluation alone. Therefore, the foraging behaviour of the animals needs to be considered in the evaluation of Sahelian rangelands for animal production.  相似文献   

13.
In-stream macrophytes are typically abundant in nutrient-rich chalk streams during the spring and summer months and modify the in-stream environment by altering river flows and trapping sediments. We present results from an inter-disciplinary study of two river reaches in the River Frome catchment, Dorset (UK). The investigation focused on how Ranunculus (water crowfoot), the dominant submerged macrophyte in the study reaches, modified patterns of flow and sediment deposition. Measurements were taken on a monthly basis throughout 2003 to determine seasonal patterns in macrophyte cover, associated changes in the distributions of flow velocities and the character and amount of accumulated fine sediment within stands of Ranunculus.Maximum in-stream cover of macrophytes exceeded 70% at both sites. Flow velocities were less than 0.1 m s− 1 within the stands of Ranunculus and accelerated to 0.8 m s− 1 outside the stands. During the early stages of the growth of Ranunculus, fine sediment mostly accumulated within the upstream section of the plant but the area of fine sediment accumulation extended into the downstream trailing section of the plant later in the growing season. The fine sediment accumulations were dominated by sand (63–1000 μm) with silts and clays (0.37–63 μm) comprising < 10% by volume. The content of organic matter in the accumulated sediments varied within stands, between reaches and through the growing season with values ranging between 9 and 105 mg g− 1 dry weight. At the reach scale the two sites exhibited different growth forms of Ranunculus which created distinctive patterns of flow and fine sediment deposition.  相似文献   

14.
The world fallout of caesium-137 (137Cs) associated with nuclear weapons testing during the 1950s and 1960s has provided a valuable man-made tracer for studies of soil erosion and sediment delivery. But relatively few researchers have used it to estimate wind erosion. In this paper, the137Cs technique is introduced into the study of wind erosion and its modern processes in the Qinghai-Tibet Plateau. Two137Cs reference inventories of 982·11 and 2376·04 Bq m−2was established preliminarily, which distribute in the south and mid-north parts of the study area respectively. By analysing the patterns of137Cs depth profiles from sampling sites, the aeolian processes of erosion and deposition along nearly 40 years has been revealed, i.e. the shrub coppice dunes(S1) and semi-fixed dunefields (S3) had experienced the alternation of erosion and deposition, while the grasslands (S4, S6 and S7) and dry farmlands (S5) suffered erosion only. By using the137Cs model, the average wind erosion rates for shrub coppice dune (S1), semi-fixed dunefields (S3), dry farmlands (S5) and grasslands (S4, S6 and S7) were estimated to be 84·14, 69·43, 30·68 and 21·84 tha−1a−1respectively, and for the whole Plateau, averaging 47·59 ha−1a−1which can be regarded as the medium erosion standard. These results derived from137Cs for the first time have significant implications for the further research of wind erosion and desertification control in the Qinghai-Tibet Plateau.  相似文献   

15.
Cactus seedlings often establish under nurse plants which modify environmental conditions by increasing moisture and decreasing solar radiation, which may cause beneficial and detrimental effects, respectively, on seedling growth. Three soil moisture treatments (5%, 25% and 60%) and two solar radiation levels (100% exposure=243 μmol m−2 s−1, and 40%=102 μmol m−2 s−1) were used in a factorial design to analyze seedling growth response of three rare cactus species (Mammillaria pectinifera, Obregonia denegrii and Coryphantha werdermannii). The variables evaluated were relative growth rate (RGR), root/shoot ratio (R/S), and K (RGRroots/RGRshoot), obtained from an initial seedling harvest (6-month-old seedlings) and a final harvest 6 months after treatment application. All three species had slow RGRs (0.002–0.012 g g−1 day−1). O. denegrii had the lowest RGR values, but was the only species for which R/S and K varied with soil moisture. While all seedlings responded markedly to soil moisture, no response was observed to radiation treatments. The latter might have been related to the relatively low solar radiation levels present in the greenhouse. Yet, our results suggest that the main benefit nurse plants offer to seedlings is the increase in soil moisture.  相似文献   

16.
Using the UNFCCC as a basis, and the objectives of estimating soil organic carbon (SOC) changes during the period 1900–2100, a spatially explicit database of climate, land cover and soil texture was compiled for a 262,000 km2 region in semi-arid Sudan. The area is characterized by low input cultivation of millet, sorghum and sesamé combined with livestock grazing. By integrating the database with the CENTURY ecosystem model, we were able to estimate historical, current and future pools of SOC as a function of land management and climate.The SOC (upper 20 cm) decrease from 1900 to 2000 was estimated to be 6·8 Mt and the maximum potential carbon sink (SOC increase) for the period 2000 to 2100 was estimated to be 17 Mt. Cropland and grassland lost 293 and 152 t SOC km−2 respectively whereas the savannahs gained 76 t SOC km−2 from 1900 to 2000. The SOC sequestration scenario simulated during 2000–2100 recovered 94, 84 and 75 t km−2 for cropland, grassland and savannah respectively.In addition to climate and soils, cropping intensity, fallow periods, fire frequency and grazing intensity also influence cropland SOC variation. Grassland and savannah SOC variations depend on grazing intensity and fire return interval. Land management may affect future amounts of SOC in semi-arid areas thereby turning them from sources into sinks of carbon. SOC estimates were reasonably consistent with measurements (r2=0·70, n=13).  相似文献   

17.
We set up an automatic weather station over a playa (the flat floor of an undrained desert basin that becomes at times a shallow lake), approximately 65 km east–west by 130 km north–south, located at the U.S. Army Dugway Proving Ground (40°08′N, 113°27′W, 1124 m above mean sea level) in north-western Utah, U.S.A., in 1999. This station measured the incoming (Rsi) and outgoing (Rso) solar or shortwave radiation using two CM21 Kipp & Zonen pyranometers (one inverted), the incoming (Rli or atmospheric) and outgoing (Rlo or terrestrial) longwave radiation, using two CG1 KippZonen pyrgeometers (one inverted), and the net (Rn) radiation using a Q*7 net radiometer (Radiation Energy Balance System, REBS). We also measured the 10-m wind speed (U10) and direction (R.M. Young wind monitor) and precipitation (Campbell Sci., Inc.). The measurements were taken every 2 s, averaged into 20-min, continuously, throughout the year. The annual (August 1999 – August 2000) comparisons of global or solar radiation and windiness with two other stations in central (Hunter) and northern (Logan) Utah, indicate higher solar radiation (Rsi,Dugway=7797 MJ m−2 period−1vs. Rsi, Hunter=7021 MJ m−2 period−1 and Rsi, Logan=6865 MJ m−2 period−1) and much higher annual mean windiness (UDugway=387 km day−1vs. UHunter=275 km day−1 and ULogan=174 km day−1) throughout the period over the playa. These data reveal the possibility of simultaneously harvesting these two sources of clean energies at this vast and uniform playa.  相似文献   

18.
The oxygen-18 and deuterium isotopic compositions of rainfall from eight meteorological stations are discussed. The results show that the stable isotope distribution of the rainfall over the western province display a difference in isotopic ratios, which can be attributed to the altitude effect and water vapour sources, and to some extent, to rainfall amount. The changes of δ18O and δ D contents with altitude are −0·08 and −0·38‰ per 100 m respectively. Comparison of the isotopic analyses of rainfall and local ground-water and springs indicates that the monsoon derived rainfall that prevails during the autumn season plays a crucial role in alluvial aquifer recharge. Although Atlantic water vapour can arrive virtually throughout the area in winter months, its influence seems to be of minor importance to the water balance in the study area.  相似文献   

19.
Sand wedges, formed mainly in a Late Pleistocene alluvial gravel in the Hexi Corridor are described in detail in this paper. 14C ages of the infilling eolian sand indicate that the sand wedges were products of the Last Ice Age. During the period of formation, the mean annual air temperature in the Hexi Corridor was about −5.3 °C, i.e. 12.6–14.6 °C lower than that of the present day. This estimated value is coincident with the fall in mean air temperature of 13 °C, which is predicated from a mirabilite (Na2SO4·10H2O) sedimentary layer, and also agrees with research on the estimated amplitude of air temperature lowering in middle and high latitudes of the Northern Hemisphere during the last glacial period. The annual precipitation in the Western Hexi Corridor at that time was probably 100–200 mm, about 100 mm more than at present.  相似文献   

20.
Four plant communities were determined through floristical and non-metric analysis: a Dolichlasium lagascae community in crevices on sunny rock outcrops; a Cercidium praecox ssp. glaucum community on sunny slopes; an Artemisia mendozana–Adesmia trijuga community on shady slopes; and a Eupatorium buniifolium community on dry riverbanks. The discriminant function analysis showed that the first group corresponds to the sunny rock outcrops discriminated by Na content (r=0·53, p<0·05), the second group is for sunny slopes discriminated by electric conductivity and the third for shady slopes–riparian discriminated by the (Ca+Mg) content (r=0·54, p<0·05). A pattern of primary succession on outcrop surfaces is proposed.  相似文献   

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