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1.
Anabranching is characteristic of a number of rivers in diverse environmental settings worldwide, but has only infrequently been described from bedrock-influenced rivers. A prime example of a mixed bedrock-alluvial anabranching river is provided by a 150-km long reach of the Orange River above Augrabies Falls, Northern Cape Province, South Africa. Here, the perennial Orange flows through arid terrain consisting mainly of Precambrian granites and gneisses, and the river has preferentially eroded bedrock joints, fractures and foliations to form multiple channels which divide around numerous, large (up to 15 km long and 2 km wide), stable islands formed of alluvium and/or bedrock. Significant local variations in channel-bed gradient occur along the river, which strongly control anabranching style through an influence on local sediment budgets. In relatively long (>10 km), lower gradient reaches (<0.0013) within the anabranching reach, sediment supply exceeds local transport capacity, bedrock usually only crops out in channel beds, and channels divide around alluvial islands which are formed by accretion in the lee of bedrock outcrop or at the junction with ephemeral tributaries. Riparian vegetation probably plays a key role in the survival and growth of these islands by increasing flow roughness, inducing deposition, and stabilising the sediments. Less commonly, channels may form by eroding into once-continuous island or floodplain surfaces. In shorter (<10 km), higher gradient reaches (>0.0013) within the anabranching reach, local transport capacity exceeds sediment supply, bedrock crops out extensively, and channels flow over an irregular bedrock pavement or divide around rocky islands. Channel incision into bedrock probably occurs mainly by abrasion, with the general absence of boulder bedforms suggesting that hydraulic plucking is relatively unimportant in this setting. Mixed bedrock-alluvial anabranching also occurs in a number of other rivers worldwide, and appears to be a stable and often long-lived river pattern adjusted to a number of factors commonly acting in combination: (1) jointed/fractured granitoid rock outcrop; (2) erosion-resistant banks and islands; (3) locally variable channel-bed gradients; (4) variable flow regimes.  相似文献   

2.
The spatial distribution of valley setting (laterally-unconfined, partly-confined, or confined) and fluvial morphology in the source region of the Yangtze and Yellow Rivers is contrasted and analyzed. The source region of the Yangtze River is divided into 3 broad sections (I, II and III) based on valley setting and channel gradient, with the upstream and downstream sections being characterized by confined (some reaches partly-confined) valleys while the middle section is characterized with wide and shallow, laterally-unconfined valleys. Gorges are prominent in sections I and III, while braided channel patterns dominate section II. By contrast, the source region of the Yellow River is divided into 5 broad sections (sections I-V) based on valley characteristics and channel gradient. Sections I, II and IV are alluvial reaches with mainly laterally-unconfined (some short reaches partly-confined) valleys. Sections III and V are mainly confined or partly-confined. Greater morphological diversity is evident in the source region of the Yellow River relative to the upper Yangtze River. This includes braided, anabranching, anastomosing, meandering and straight alluvial patterns, with gorges in confined reaches. The macro-relief (elevation, gradient, aspect, valley alignment and confinement) of the region, linked directly to tectonic movement of the Qinghai-Tibet Plateau, tied to climatic, hydrologic and biotic considerations, are primary controls upon the patterns of river diversity in the region.  相似文献   

3.
While studies on gravel mantled and mixed alluvial bedrock rivers have increased in recent decades, few field studies have focused on spatial distributions of bedrock and alluvial reaches and differences between reach types. The objective of this work is to identify the spatial distribution of alluvial and bedrock reaches in the Upper Guadalupe River. We compare reach length, channel and floodplain width, sinuosity, bar length and spacing, bar surface grain size, and slope in alluvial and bedrock reaches to identify whether major differences exist between channel reach types. We find that local disturbances, interaction of the channel and valley sides, variation in lithology, and regional structural control contribute to the distribution of bedrock reaches in the largely alluvial channel. Alluvial and bedrock channel reaches in the Upper Guadalupe River are similar, particularly with respect to the distribution of gravel bars, surface grain size distributions of bars, and channel slope and width. Our observations suggest that the fluvial system has adjusted to changes in base level associated with the Balcones Escarpment Fault Zone by phased incision into alluvial sediment and the underlying bedrock, essentially shifting from a fully alluvial river to a mixed alluvial bedrock river.  相似文献   

4.
In February/March 2007 extreme rainfall occurred over a four‐day period in the 7000 km2 East Alligator River catchment in Arnhem Land, northern Australia. The resultant large flood caused extensive bank erosion, channel widening, stripping of point bars and floodplain, resulting in large amounts of sand transport. This sand was largely deposited in the downstream river channel as a sand slug, and as deep overbank sand splays where the valley abruptly widened immediately downstream of an island anabranching, bedrock‐confined reach. Interpretation of a time series of aerial photographs and satellite images from 1950 to 2012 showed that there have been considerable channel changes along the study reach. The aerial photographs show that extensive sedimentation in the same reach as in 2007 also occurred in 1975, 1981 and 1984. Each time, the sand slug was reworked rapidly over succeeding years by subsequent smaller floods, and the channel deepened naturally as sand supply declined due to revegetation of the upstream riparian zone and the reformation and stabilisation of point and lateral bars. Sand slug formation at an intermediate floodout is an episodic process dependent on the supply of large volumes of sand by extensive channel erosion during extreme floods. A conceptual geomorphic model was developed to highlight the differential effectiveness of extreme versus moderate floods.  相似文献   

5.
Leif M. Burge   《Geomorphology》2004,63(3-4):115-130
This study tests the assumption that the characteristics of channels within multiple channel rivers are different from those of single channel rivers. Some river restoration approaches propose radical transformation of river patterns, from multiple to single channels, based on the link between river patterns and their in-channel characteristics. Determining the links between river patterns and their in-channel characteristics is complicated by differences in geology, history, climate and discharge among rivers. Furthermore, multiple channel rivers are composed of a mosaic of channel types with a range of in-channel characteristics. This study minimizes these problems by analysing a single river containing neighbouring single and multiple channel patterns with little change in discharge downstream, and by analysing all channel types. The study addressed two objectives: to determine the hydraulic geometry, energy, and sediment mobility characteristics of neighbouring single and multiple channel river patterns, and to test for statistical differences in these characteristics between patterns. The Renous River shows a wandering pattern for 11.5 km, with multiple channels around semipermanent islands and abandoned channels in the flood plain. The river displays a single channel river pattern where channels are confined by their valley walls, upstream and downstream of wandering. The analysis was conducted at three scales. First, the confined single channel and wandering multiple channel patterns were compared (pattern scale). Second, the confined channel pattern was compared to single and multiple channel sections within the wandering pattern (section scale). Third, all channel types were compared (channel type scale). Multi response permutation procedure (MRPP) and analysis of variance (ANOVA) were used to analyze differences between channels. Difference tests found no simple discrimination between the single and multiple channel river patterns of the Renous River. Tests between the single confined and multiple wandering channel patterns found few differences in the in-channel variables. The tests did find differences between multiple channel sections within the wandering pattern and confined single channels; however, a greater number of differences were found between multiple channel and single channel sections within the wandering pattern, highlighting the variability within the wandering pattern. Two groups emerged when all channel types were tested for differences: perennial main-channels containing the thalweg, and ephemeral side-channels. Therefore, side-channels define the in-channel characteristics of wandering rivers because few differences were found among main-channels in either pattern. This analysis suggests that all channel types, not just main-channels, should be investigated to obtain a complete picture of a river pattern prior to any restoration efforts. Engineers must exercise caution when applying the link between river patterns and in-channel characteristics to river restoration efforts.  相似文献   

6.
Dissolved and total organic and inorganic carbon in some British rivers   总被引:2,自引:0,他引:2  
Rivers transport both organic and inorganic carbon from their sources to the sea. Results of ~800 organic and inorganic analyses from various British rivers of contrasting size and land use are presented here: (1) the headwater River Tern, a rural river of 852 km2 catchment; (2) the Ouseburn, a small urban 55 km2 catchment; (3) the River Tyne, a larger river system of ~3000 km2 catchment; (4) a spatial survey from 205 sample sites on ~60 rivers from SW England. We found that, with the exception of peat-rich headwaters, DIC concentration is always greater than DOC. DIC is primarily in the form HCO3 , with DIC concentrations highest in highly urbanised catchments, typically greater than those observed in catchments with carbonate bedrock, demonstrating a significant and previously unrecognised anthropogenic inorganic carbon input to urban rivers.  相似文献   

7.
Crustal uplift in southern England: evidence from the river terrace records   总被引:6,自引:0,他引:6  
Much of the past work on the Quaternary rivers of northwest Europe has been concerned with river terraces, which characterise almost every valley. While these terraces are undoubtedly striking features of the landscape, the incision achieved by Quaternary rivers is equally significant in terms of river behaviour, and for an understanding of the factors affecting landform development during the Quaternary. This paper examines the incision achieved during the Quaternary by the Thames, in both its upper and lower catchments, and by the Hampshire Avon in southern England. Valley incision rates of ca. 0.07–0.10 m ka−1 have been identified, although in the lower catchment of the Thames, these have been enhanced by additional incision in response to glacio-isostasy and valley shortening. A model is proposed in which regional uplift is recognized as the primary cause of incision by these Quaternary rivers. Possible mechanisms for regional uplift are considered.  相似文献   

8.
The 270 km long section of the Upper Yellow River at the First Great Bend is comprised of single channel and multiple channel systems that alternate among anastomosing, anabranching, meandering and braided reaches. The sequence of downstream pattern changes is characterized as: anastomosing-anabranching, anabranching-meandering, meandering-braided and braided-meandering. Remote sensing images, DEM data and field investigations are used to assess ahd interpret controls on these reach transitions. Channel slope and bed sediment size are key determinants of transitions in channel planform. Anas- tomosing reaches have a relatively high bed slope (0.86‰) and coarser sediment bed material (d50 = 3.5 mm). In contrast, meandering reaches have a low slope (0.30‰) and fine sediment bed material (d50 = 0.036 mm). The transition from a meandering to braided pattern is characterized by an increase in channel width-depth ratio, indicating the important role of bank strength (i.e. cohesive versus non-cohesive versus channel boundaries). Interestingly, the braided-meandering and meandering-braided transitions are coincident with variable flow inputs from tributary rivers (Baihe and Heihe rivers respectively). Theoretical analysis of the meandering-braided transition highlights the key control of channel width-depth ratio as a determinant of channel planform.  相似文献   

9.
Late Quaternary alluvial induration has greatly influenced contemporary channel morphology on the anabranching Gilbert River in the monsoon tropics of the Gulf of Carpentaria. The Gilbert, one of a number of rivers in this region, has contributed to an extensive system of coalescing low-gradient and partly indurated riverine plains. Extensive channel sands were deposited by enhanced flow conditions during marine oxygen isotope (OI) Stage 5. Subsequent flow declined, probably associated with increased aridity, however, enhanced runoff recurred again in OI Stages 4–3 (65–50 ka). Aridity then capped these plains with 4–7 m of mud. A widespread network of sandy distributary channels was incised into this muddy surface from sometime after the Last Glacial Maximum (LGM) to the mid Holocene during a fluvial episode more active than the present but less so than those of OI Stages 5 and 3. This network is still partly active but with channel avulsion and abandonment now occurring largely proximal to the main Gilbert flow path.A tropical climate and reactive catchment lithology have enhanced chemical weathering and lithification of alluvium along the river resulting in the formation of small rapids, waterfalls and inset gorges, features characteristic more of bedrock than alluvial systems. Thermoluminescence (TL) and comparative optically stimulated luminescence (OSL) ages of the sediments are presented along with U/Th ages of pedogenic calcrete and Fe/Mn oxyhydroxide/ oxide accumulations. They show that calcrete precipitated during the Late Quaternary at times similar to those that favoured ferricrete formation, possibly because of an alternating wet–dry climate. Intense chemical alteration of the alluvium leading to induration appears to have prevailed for much of the Late Quaternary but, probably due to exceptional dryness, not during the LGM. The result has been restricted channel migration and a reduced capacity for the channel to adjust and accommodate sudden changes in bedload. Consequent avulsions have caused local stream powers to increase by an order of magnitude, inducing knickpoint erosion, local incision and the sudden influx of additional bedload that has triggered further avulsions. The Gilbert River, while less energetic than its Pleistocene ancestors, is clearly an avulsive system, and emphasizes the importance in some tropical rivers of alluvial induration for reinforcing the banks, generating nickpoints, reworking sediment and thereby developing and maintaining an indurated and anabranching river style.  相似文献   

10.
A geomorphological and sedimentological case study is made of the Sprongdøla, a medium-sized river flowing in a predominantly bedrock channel in the mountains of southern Norway. Particular attention is given to bed material characteristics and potential sediment sources. Downstream patterns in clast size and shape, including sphericity and roundness, are poorly developed in the Sprongdøla. There is an apparent absence of down-stream fining and a large proportion of bedload in the lower channel remains in very angular and angular roundness categories. Covariance analysis of clast shape and roundness allows this to be attributed to the widespread input of coarse, angular rock particles derived from snow-avalanche activity on the valley sides and especially from frost-shattering of bedrock in the channel, which produces characteristically slabby clasts. The latter process is important in channel widening and in steepening the channel walls. Strictly fluvial processes, avalanche activity and frost-shattering are controlled in different ways by related annual environmental cycles. Their interaction in the context of the periglacial river channel produces a unique process-sediment-landform association that distinguishes alpine periglacio-fluvial systems from those associated with glacio-fluvial rivers and fluvial landscapes in the temperate zone.  相似文献   

11.
Ground‐penetrating radar surveys, coring and the analyses of satellite and aerial images have been carried out to study differences in the evolution of meander bends formed in various geological conditions. The research was conducted in the lower course of the Obra River (western Poland) characterized by a complex geology: particular sections of the valley were formed in glacial, stagnant water and fluvioglacial deposits. The research was conducted in four detailed study sites representing different formation conditions for the meander bends. Four types of meanders were distinguished: laterally migrating bends characterized by frequent changes of migration direction accompanied by river bed avulsions and cutoffs; bends with traces of continuous migration limited by stagnant water basin deposits; confined meanders in a narrow valley formed in glacial till characterized by the occurrence of mid‐channel islands; and meanders with traces of complex changes of the river bed migration influenced by anthropogenic intervention followed by intensive overbank deposition. Moreover, traces of an early development of the Obra valley and remains of multi‐channel pattern were discovered. The results also show that despite being formed in different geological conditions, a similar number of the meander migration phases were recorded in the floodplain architecture during the last 7000 years at each of the sites. It is also found that the development of the studied meanders was slow compared with cases of actively migrating meandering rivers.  相似文献   

12.
密云水库以上的潮白河分为白河和潮河两支,分别穿行于燕山山区。两河的上段为顺直河型或游荡河型,下段为典型的深切曲流。河势及河型受地质构造的深刻影响。潮白两河的纵比降大,来水、来沙在时间和空间上的分配不均匀,对不同河型的形成有影响,这也表明密云水库以上的潮白河具有山地河流的特点。  相似文献   

13.
The Spiti River that drains through the arid Trans-Himalayan region is studied here. The relict deposits exposed along the river provide an opportunity to understand the interaction between the phases of intense monsoon and surface processes occurring in the cold and semi arid to-arid Trans-Himalayan region. Based on geomorphological observation the valley is broadly divided into the upper and lower Spiti Valley. The braided channel and the relict fluvio-lacustrine deposits rising from the present riverbed characterize the upper valley. The deposits in the lower valley occur on the uplifted bedrock strath and where the channel characteristics are mainly of meandering nature. Conspicuous is the occurrence of significantly thick lacustrine units within the relict sedimentary sequences of Spiti throughout the valley. The broad sedimentary architecture suggests the formation of these palaeolakes due landslide-driven river damming. The Optically Stimulated Luminescence (OSL) dating of quartz derived from the bounding units of the lacustrine deposits suggests that the upper valley preserves the phase of deposition around 14–6 ka and in the lower valley around 50–30 ka. The review of published palaeoclimatic palaeolake chronology of Spiti Valley indicates that the lakes were probably formed during the wetter conditions related to Marine Isotope Stage III and II. The increased precipitation during these phases induced excessive landsliding and formation of dammed lakes along the Spiti River. The older lacustrine phase being preserved on the uplifted bedrock strath in the lower valley indicates late Pleistocene tectonic activity along the Kaurick Chango normal fault.  相似文献   

14.
Joanna Korpak   《Geomorphology》2007,92(3-4):166
The purpose of this paper is to explain the influence of river training on channel changes in mountain rivers. Also considered are the causes of failure of different training schemes. The research was conducted on the regulated Mszanka and Porębianka Rivers, belonging to the Raba River drainage basin in the Polish Flysh Carpathian Mountains. Channel mapping carried out in 2004 drew attention to the contemporary morphology of the channels and the development of their dynamic typology. General changes in channel morphometry and land cover were identified by comparing cartographic sources from various years. Archive material from Cracow's Regional Water Management Authority (RZGW) was used to analyse the detailed channel changes caused by each regulation structure. The material consisted of technical designs of individual training works, as well as plans, longitudinal profiles and cross-sections of trained channel reaches. A series of minimum annual water stages at the Mszana Dolna gauging station was used to determine the tendency of channel bed degradation over 53 years. During the first half of the 20th century, the middle and lower courses of the Mszanka and Porębianka Rivers had braided patterns. The slopes, mostly covered with crops, were an important source of sediment delivery to the river channels. Today, both channels are single-threaded, narrow and sinuous. Downcutting is the leading process transforming the channels. They cut down to bedrock along about 60% of their lengths. The main type of channel is an erosion channel, which occurs also in the middle and lower courses of the rivers. The channel sediment deficit is an important cause for river incision. Sediment supply to the channels was reduced after a replacement of crops on the slopes by meadows or forests. Gravel mining has also caused channel downcutting. The rapid channel changes began after 1959, as systematic training was introduced. Channel regulation seems therefore to be a major factor determining channel adjustment. Debris dams and groynes were built before 1980 and these caused the greatest change of channel pattern, increase of channel gradient and magnitude of river incision. After that date the measures mostly involved drop structures. From then on, the rate of downcutting decreased considerably, but has not ceased. The rivers continued to incise until bedrock was exposed or training structures were destroyed. After that, a tendency to lateral migration and local braiding were observed in the deepened channel. The channels displayed a tendency to return to their morphology and dynamic from before the training. The results demonstrate that river training distorts the equilibrium of channel systems. A channel becomes divided into artificial reaches, which later follow different evolutionary patterns. Most training schemes on mountain channels are ineffective in the long term, as river managers seem to consider a channel at a reach scale only. Individual channel reaches, however, are not independent but rather form a system that must be managed at the entire channel scale.  相似文献   

15.
Extensive valley fills have formed at the base of the escarpment in granitic catchments along the south coast of NSW. On the 1865 portion plan, the valley fill surface in the upper part of Wolumla Creek, in the Bega River catchment, was intact, but within a few decades of European settlement of the area the valley fill had been incised. Today the incised channel is up to 10 m deep and 100 m wide. The catchment drains an area of just 18.2 km2. Based on detailed field mapping, with extensive drilling and angering, the volume of the intact valley fill in upper Wolumla Creek in 1865 was approximately 5000 × 103 m3. Between 1865 and the present day, approximately 3500 × 103 m3 of this material has been removed, leaving roughly 1500× 103 m3 of material stored on the valley margins. During an initial period of discontinuous gullying, approximately 230 × 103 m3 of sand accumulated as a floodout. Subsequently, the incised channel became continuous, cutting through the floodout; over 50 per cent of floodout deposits were removed. Flushing of the materials released from upland valley fills has been very efficient in the Wolumla Creek catchment, with a sediment delivery ratio of around 70 per cent. The efficient downstream transfer of deposits reflects bedrock confinement in downstream reaches. Extensive volumes of material have accumulated along the lower reaches of the catchment, exacerbating the transformation to the geomorphic character of the lower Bega River.  相似文献   

16.
Extreme flood events are considered by many researchers to be very important in controlling the development of semi‐arid bedrock‐influenced river systems. Accurate gauging of such events is often impossible, however, as gauges are drowned and often damaged during the event. A methodology for estimating flood discharge for bedrock‐influenced channels is presented that reconstructs hydrometric characteristics of the peak flow and relates these to the roughness character of the river channel in question. The method is evaluated using peak water‐surface slope data relating to the extreme floods of February 2000 along the Sabie and Letaba rivers, located respectively in the Mpumalanga and Northern Provinces, South Africa. The data, in the form of strandline measurements, were taken at hydraulically relevant points along the long profile of both rivers. The resultant data are utilised together with published high flow channel resistance figures, based on the channel morphology of the Sabie and Letaba rivers, to generate peak flow estimates for a number of locations along both rivers. Comparisons are made between the frictional discharge peak flow estimates, velocity‐area and hydrologic estimates of peak flow. These comparisons indicate that the method can produce discharge estimates with an accuracy of ±10% and ± 35% respectively.  相似文献   

17.
Rivers flowing from glacier-clad Quaternary volcanoes in southwestern British Columbia have high sediment loads and anabranching and braided planforms. Their floodplains aggrade in response to recurrent large landslides on the volcanoes and to advance of glaciers during periods of climate cooling. In this paper, we document channel instability and aggradation during the last 200 years in lower Cheakamus River valley. Cheakamus River derives much of its flow and nearly all of its sediment from the Mount Garibaldi massif, which includes a number of volcanic centres dominated by Mount Garibaldi volcano. Stratigraphic analysis and radiocarbon and dendrochronological dating of recent floodplain sediments at North Vancouver Outdoor School in Cheakamus Valley show that Cheakamus River aggraded its floodplain about 1–2 m and buried a valley-floor forest in the early or mid 1800s. The aggradation was probably caused by a large (ca. 15–25×106 m3) landslide from the flank of Mount Garibaldi, 15 km north of our study site, in 1855 or 1856. Examination of historical aerial photographs dating back to 1947 indicates that channel instability triggered by this event persisted until the river was dyked in the late 1950s. Our observations are consistent with data from many other mountain areas that suggest rivers with large, but highly variable sediment loads may rapidly aggrade their floodplains following a large spike in sediment supply. Channel instability may persist for decades to centuries after the triggering event.  相似文献   

18.
世界名河及其流域开放开发的国际经验   总被引:4,自引:0,他引:4  
世界名河及其流域自然条件千差万别,其开放开发的空间模式各异,选择最具社会经济特色和分别代表某一开放开发类型的世界四大名河-密西西北河、圣劳伦斯河、亚马逊河和尼罗河,对其流域背景及开发战略进行了比较研究,旨在为我国大江大河及其流域的开放琥发提供借鉴。  相似文献   

19.
Bald cypress (Taxodium distichum) regeneration was examined in the bottomland forest of the lower Hatchie River in western Tennessee. Bald cypress is found mostly in abandoned meanders. Its absence along the active channel indicates that colonization begins after the meander cuts off, creating an oxbow lake. Analysis of the size, the age, and the spatial structure of the stands indicates that the individuals first establish along the margins of the lake, and then colonize the interior as the lake fills in and new surfaces are exposed. Bald cypress will dominate abandoned meanders for several centuries. However, it will not regenerate in stands with a well-developed forest canopy. Bald cypress is eventually replaced by shade tolerant species and, therefore, only rarely occurs on older surfaces. Channel migration and the subsequent creation of oxbows will maintain bald cypress as a major component of bottomland forests. However, many of the major streams in the lower Mississippi River alluvial valley have been channelized. Maintenance of a straightened channel will exclude the creation of oxbows. Therefore, a primary regeneration opportunity for bald cypress disappears in many areas. [Key words: channel migration, exact probabilities, floodplain forest, oxbow lakes, Taxodium distichum, Tennessee.]  相似文献   

20.
The Quaternary evolution and the morpho-sedimentary features of some of the most important rivers in Spain (Ebro and Tagus rivers among others) have been controlled by subsidence due to alluvial karstification of the evaporitic bedrock. The subsidence mechanism may range from catastrophic collapse to slow sagging of the alluvium by passive bending. In the Ebro Basin, the mechanisms and processes involved in karstic subsidence were studied through the analysis of present-day closed depressions as well as through old subsidence depressions (palaeocollapses and solution-induced basins) and associated deformations recorded in the Quaternary alluvial sediments. The Gállego–Ebro river system is presented as a case study of channel adjustments and geomorphic and sedimentary evolution of fluvial systems in dissolution-induced subsidence areas. In this fluvial system, evaporite dissolution during particular Quaternary time intervals (namely early and middle Pleistocene) have lead to the development of a solution-induced basin, approximately 30 km-long by 8 km-wide, filled by Quaternary deposits with a total thickness in excess of 190 m. The main river response to balance the subsidence in the alluvial plain was aggradation in the central reach of the subsiding area, and degradation both in the upstream reach and in the valley sides where alluvial fans and covered pediments may prograde over the fluvial sediments. The main sinking areas are recognized in the sedimentary record by anomalous thickenings in the alluvial deposits and fine-grained sediments deposited in backswamp and ponded areas.  相似文献   

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