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1.
Contemporary variants of the lichenometric dating technique depend upon statistical correlations between surface age and maximum lichen sizes, rather than an understanding of lichen biology. To date three terminal moraines of an Alaskan glacier, we used a new lichenometric technique in which surfaces are dated by comparing lichen population distributions with the predictions of ecological demography models with explicit rules for the biological processes that govern lichen populations: colonization, growth, and survival. These rules were inferred from size–frequency distributions of lichens on calibration surfaces, but could be taken directly from biological studies. Working with two lichen taxa, we used multinomial‐based likelihood functions to compare model predictions with measured lichen populations, using only the thalli in the largest 25% of the size distribution. Joint likelihoods that combine the results of both species estimated moraine ages of ad 1938, 1917, and 1816. Ages predicted by Rhizocarpon alone were older than those of P. pubescens. Predicted ages are geologically plausible, and reveal glacier terminus retreat after a Little Ice Age maximum advance around ad 1816, with accelerated retreat starting in the early to mid twentieth century. Importantly, our technique permits calculation of prediction and model uncertainty. We attribute large confidence intervals for some dates to the use of the biologically variable Rhizocarpon subgenus, small sample sizes, and high inferred lichen mortality. We also suggest the need for improvement in demographic models. A primary advantage of our technique is that a process‐based approach to lichenometry will allow direct incorporation of ongoing advances in lichen biology.  相似文献   

2.
Botanists make yearly measurements of lichen sizes that describe highly variable radial expansion of young, and old, Rhizocarpon subgenus Rhizocarpon that is a function of thallus size and age. Such non‐uniform growth would negate use of lichens to date geomorphic events, such as landslides and moraines, of the past 1000 years. Fortunately, many crustose lichens tend toward circular shapes, which can be achieved only when overall uniform radial growth prevails. Largest lichen measurements on rockfall blocks that accumulate incrementally as hillslope talus in earthquake‐prone California plot as distinct peaks in frequency distributions. Rockfall surface‐exposure times are known to the day for historical earthquakes and to the year where mass movements damage trees. Lichenometry consistently dates regionally synchronous rockfall events with an accuracy and precision of ±5 years. Only historical records and tree‐ring dating of earthquakes are better. The four crustose lichens used here have constant long‐term growth rates, ranging from 9.5 to 23.1 mm per century. Growth rates do not vary with altitude or climate in a 900 km long mountainous study region in California, USA. Linear growth regressions, when projected to the present, constrain estimates of colonization time and possible styles of initial lichen growth.  相似文献   

3.
Variation in lichen growth rates poses a significant challenge for the application of direct lichenometry, i.e. the construction of lichen dating curves from direct measurement of growth rates. To examine the magnitude and possible causes of within‐site growth variation, radial growth rates (RaGRs) of thalli of the fast‐growing foliose lichen Melanelia fuliginosa ssp. fuliginosa (Fr. ex Duby) Essl. and the slow‐growing crustose lichen Rhizocarpon geographicum (L.) DC. were studied on two S‐facing slate rock surfaces in north Wales, UK using digital photography and an image analysis system (Image‐J). RaGRs of M. fuliginosa ssp. fuliginosa varied from 0.44 to 2.63 mm yr–1 and R. geographicum from 0.10 to 1.50 mm yr–1.5. Analysis of variance suggested no significant variation in RaGRs with vertical or horizontal location on the rock, thallus diameter, aspect, slope, light intensity, rock porosity, rock surface texture, distance to nearest lichen neighbour or distance to vegetation on the rock surface. The frequency distribution of RaGR did not deviate from a normal distribution. It was concluded that despite considerable growth rate variation in both species studied, growth curves could be constructed with sufficient precision to be useful for direct lichenometry.  相似文献   

4.
The recently observed recession of glaciers on King George Island is associated with decades of climate warming in the Antarctic Peninsula region. However, with only 60 years of glaciological observations in the study area ages of the oldest moraines are still uncertain. The goal of the study was to estimate ages of lichen colonization on the oldest moraines of the Ecology and White Eagle Glaciers on King George Island and on the Principal Cone of Penguin Island volcano. The first lichenometric studies on these islands from the late 1970s used rates that had about four to five times slower Rhizocarpon growth rates. We re‐examined the sites and measured 996 thalli diameters to establish the surface ages. To estimate the age we used (1) long‐term Rhizocarpon lichen group growth rates established by authors using data from a previous lichenometric study on King George Island, and (2) previous data of lichen growth rates from other sub‐Antarctic islands. Our results suggest growth rates between 0.5 and 0.8 mm yr–1. According to these rates the ages of the oldest moraine ridges are of the Little Ice Age and were colonized at the beginning of the twentieth century. The mid‐twentieth century age of lichen colonization on the historically active Penguin Island volcano might support the date of the last eruption reported by whalers in the end of the nineteenth and the beginning of the twentieth century.  相似文献   

5.
Certain species of crustose lichens have concentrically zoned margins which probably represent yearly growth rings. These marginal growth rings offer an alternative method of studying annual growth fluctuations, establishing growth rate–size curves, and determining the age of thalli for certain crustose species. Hence, marginal growth rings represent a potentially valuable, unexploited, tool in lichenometry. In a preliminary study, we measured the widths of the successive marginal rings in 25 thalli of Ochrolechia parella (L.) Massal., growing at a maritime site in north Wales. Mean ring widths of all thalli varied from a minimum of 1.02 mm (the outermost ring) to a maximum of 2.06 mm (the third ring from the margin). There is some suggestion that marginal ring width and thallus size are positively correlated; and hence that growth rates increase in larger thalli in this small population. In a further study on recently exposed bedrock adjacent to Breiðarlon, SE Iceland, we examined the potential for using marginal growth rings to estimate thallus age of a lichen tentatively identified as a Rhizocarpon (possibly R. concentricum (Davies) Beltram.) and thus confirm the timing of surface exposure ( c. 50 years). Collectively, these results suggest: 1) the measurement of marginal rings is a possible alternative method of studying the growth of crustose lichens; 2) O. parella may grow differently to other crustose species, exhibiting a rapidly increasing radial growth rate in thalli >40 mm; 3) where lichens with marginal rings grow on recently exposed surfaces (<60 yrs), minimum age estimates can be made using growth rings as an in situ indication of lichen growth rate; 4) it is suggested that this phenomenon could provide a valuable, previously unexploited, in situ lichenometric-dating tool in areas lacking calibration control.  相似文献   

6.
Abstract The age of recent deposits can be determined using an intrinsic characteristic of the lichen ‘population’ growing on their surface. This paper presents a calibrated dating curve based on the gradient of the size‐frequency distribution of yellow‐green Rhizocarpon lichens. The dating potential of this new curve is tested on surfaces of known age in southeast Iceland. This particular size—frequency technique is also compared with the more traditional largest‐lichen approach. The results are very encouraging and suggest that the gradient can be used as an age indicator, at least on deposits formed within the last c. 150 years – and probably within the last c. 400 years – in the maritime subpolar climate of southeast Iceland. Using both lichenometric techniques, revised dates for moraines on two glacier forelands are presented which shed new light on the exact timing of the Little Ice Age glacier maximum in Iceland.  相似文献   

7.
This paper highlights the importance for dating accuracy of initial studies of delay before colonization for both trees and lichens and tree age below core height, particularly in recently deglaciated terrain where colonization and growth rates may vary widely due to differences in micro-environment. It demonstrates, for the first time, how dendrochronology and lichenometry can be used together in an assessment of each other's colonization and growth rates, and then cross-correlated to provide a supportive dating framework. The method described for estimating tree age below core height is also new. The results show that on the east side of the North Patagonian Icefield in the Arco and Colonia valleys, Nothofagus age below a core height of 112 cm can vary from 5 to 41 years and delay before colonization may range from a maximum of 22 years near water to a minimum of 93 years on the exposed flanks of the Arenales and Colonia Glaciers. Tree age plus colonization delay supplied a maximum growth rate of 4.7 mm/year for the lichen Placopsis perrugosa and lichen colonization is estimated to take from 2.5 to approximately 13 years. A minimum lichenometric date of 1883 was estimated for an ice-formed trimline at the junction of the Arenales and Colonia glaciers and a maximum dendrochronological date of 1881 for a water-formed trimline in the Arco valley. Tree and lichen ages around the valley suggest that a glacial outburst drained the 1881 high level lake releasing approximately 265 million cubic metres of water. Repeated flooding, with a minimum of 38 high lake levels, is suggested by horizontal sediment lines on the Arco valley walls and moraine flanks. Dating confirmed diminishing flood levels with a last minor flood in 1963. The wider significance of the work is that it should produce more accurate dating of recent glacier fluctuations around the North Patagonia Icefield, an area where dated reference surfaces are extremely scarce.  相似文献   

8.
The endolithic lichen Lecidea auriculata is known to enhance rock surface weathering on the Little Ice Age moraines of the glacier Storbreen in Jotunheimen, central southern Norway. This study demonstrates the reduction in Schmidt hammer Rvalues that followed the rapid colonization by this lichen of pyroxene‐granulite boulders on terrain deglaciated over the last 88 years. In the absence of this lichen, the characteristic mean R‐value of boulder surfaces is 61.0 ± 0.3; where this lichen is present, R‐values are lower by at least 20 units on surfaces exposed for 30–40 years. A similar reduction in rock hardness on rock surfaces without a lichen cover requires about 10 ka. The rapid initial weakening of the rock surfaces is indicative of rates of biological weathering by endolithic lichens that may be two orders of magnitude (200–300 times) faster than rates of physico‐chemical weathering alone. If not avoided, the effects of this type of lichen are likely to negate the effectiveness of the Schmidt hammer and other methods for exposure‐age dating, including cosmogenic‐nuclide dating, in severe alpine and polar periglacial environments. The results also suggest a new method for dating rock surfaces exposed for <50 years.  相似文献   

9.
This study presents a growth curve developed from direct and indirect growth rates of Rhizocarpon geographicum lichens from study sites on Mounts Baker, Rainier, Adams, and Hood in the northern Cascade Range of the western USA. Our observations of direct growth rates are based on 31 measurements of 11 lichens growing on different surfaces. This direct growth rate dataset is complemented by indirect growth rates based on measurements of the largest lichen observed on 20 different surfaces over 24–33‐yr periods. The direct and indirect datasets produce statistically indistinguishable mean radial growth rates of 0.48 and 0.50 mm yr?1, respectively. Statistical analysis of zero and first order fits of our growth rate data suggests that lichen growth is best characterized by the average of our mean growth rate (zero order) models at 0.49 mm yr?1. Our revised growth curve for the study area extends the applicable range for dating rock surface in the study area to the seventeenth century, approximately 175 years longer than previous calibrated curves.  相似文献   

10.
Glaciers in small mountain cirques on South Georgia respond rapidly and sensitively to changes in South Atlantic climate. The timing and rate of their deglaciation can be used to examine the impact that nineteenth- and twentieth-century climate change has had on the glacial dynamics and terrestrial ecosystems of South Georgia. As part of a reconnaissance study in Prince Olav Harbour (POH), South Georgia, we measured the size of lichens ( Rhizocarpon Ram. em Th. Fr. subgenus. Rhizocarpon group) on ice-free moraine ridges around two small mountain cirques. Our aims were twofold: first, to provide age estimates for lichen colonization, and hence, deglaciation of the moraine ridges, and second, to examine the potential of applying lichenometry more widely to provide deglacial age constraints on South Georgia. In the absence of lichen age-size (dating) curves for South Georgia, we use long-term Rhizocarpon lichen growth-rates from recent studies on sub-Antarctic Islands and the western Antarctic Peninsula to calculate likely age estimates. These data suggest ice retreat from the two outermost moraines occurred between the end of the 'Little Ice Age' (post c. 1870) and the early twentieth century on South Georgia. Lichen colonization of the innermost moraines is probably related to glacier retreat during the second half of the twentieth century, which has been linked to a well-defined warming trend since c. 1950. Patterns of possible nineteenth- and twentieth-century glacial retreat identified in POH need to be tested further by establishing species- and site-specific lichen age-size (dating) curves for South Georgia, and by applying lichenometry to other mountain cirques across South Georgia.  相似文献   

11.
阐述了地衣形态量计法的基本原理和各种不同的具体量计方法及其在同震滑坡研究中的应用方法,指出将该方法应用于同震滑坡研究中,可以帮助提供地震时间、震中、地震频度、地震影响范围和地震断层的一些信息.  相似文献   

12.
Matthews, J. A. Families of lichenometric dating curves from the Storbreen gletschervorfeld, Jotunheimen, Norway. Norsk geogr. Tidsskr. 28, 215–235.

Lichenometric dating, based on Rhizocarpon geographicum, is applied to the establishment of an areal chronology for deglaciation of the Storbreen gletschervorfeld, central southern Norway. A simple approach permitting many lichenometry curves to be constructed in the same area is adopted, each curve differing in the number of sites per surface or the number of lichens per site employed in its construction. Nine lichenometry curves of exponential form are constructed from largest lichens on four past glacier margins of known age, and the age of four margins of unknown age predicted. Median predicted ages are 1811, 1833, 1854 and 1871 and all predictions fall within an overall range of 17 years, 10 years, 10 years and 7 years respectively. The reproducibility of the predicted ages, together with independent supporting evidence, suggests that families of lichenometry curves allow considerable confidence to be placed in the lichenometric dates and are a promising addition to lichenometric dating technique in general.  相似文献   

13.
Scotland, a maritime subpolar environment (55–60°N), has seen relatively few applications of lichenometry – even though it offers much potential. Perhaps surprisingly, direct measurements of Rhizocarpon geographicum growth rates in Scotland are so far lacking. This study reports on the growth of this crustose areolate species from two sites in Assynt, NW Scotland, between 2002 and 2009. Repeat photography of 23 non-competing thalli growing under identical environmental conditions on a single vertical surface over 5 years at Inchnadamph showed growth rates to be a function of size – with larger thalli (10–30 mm) growing significantly faster than the smallest thalli (<10 mm). Mean diametral growth rates in thalli >10 mm are 0.67 mm yr−1 (s.d. = 0.16). Studies on a second vertical surface near Lochinver, over 7 years, yielded complex growth data on a more mature population of R. geographicum thalli (<50 mm in diameter). Here, mean diametral growth rates in the larger thalli (>10 mm) are slower (0.29 mm yr−1; s.d. = 0.12) than those at Inchnadamph. However, at this site, competition with other species rules out any meaningful comparison of growth rates between the two sites. Other growth processes were monitored over the five to seven-year study period, including hypothallus growth, areolae development, thallus coalescence, and inter-species competition – all have important implications for the use of Rhizocarpon species in lichenometry.  相似文献   

14.
Lichenometric measurements using Rhizocarpon ssp. were carried out on 20 talus slopes in the cirques of the Finstertal valley (Austria) at an elevation of 2300–3000 m a.s.l. The aim was to assess activity patterns on selected slopes and between the slopes of the study area, to find evidence of rockfall pulses in the last centuries and to calculate rockwall retreat rates. A calibration curve was derived from five sites of known age and adapted to the prevailing size of talus boulders. We measured the five largest lichens on more than 300 boulders and the percentage coverage of Rhizocarpon‐free clasts on more than 1000 test fields. Most of the investigated talus cones are characterized by moderate rockfall supply, with the apex being more active than the talus foot and moderate redistribution by avalanche and debris flows. Considerably enhanced activity was found under rockwalls influenced by permafrost, particularly on the north faces at an elevation of >2600 m a.s.l. At currently moderately active sites, boulder falls seem to have been slightly more frequent in the late nineteenth and first half of the twentieth century. In positions where permafrost is expected in the rockwalls, a weak maximum in the late nineteenth century and highly active present‐day conditions were found, the latter being assigned to current permafrost melt. Rockwall retreat rates derived from lichen coverage are between 400 and 1500 mm/ka which is in good concurrence with talus volume assessments, but higher than the rates derived from direct rockfall measurements. The rates derived from lichen coverage have to be taken with caution as the effects of debris redistribution are hard to quantify.  相似文献   

15.
Crustose species are the slowest growing of all lichens. Their slow growth and longevity, especially of the yellow-green Rhizocarpon group, has made them important for surface-exposure dating (lichenometry). This review considers various aspects of the growth of crustose lichens revealed by direct measurement including: 1) early growth and development; 2) radial growth rates (RGR, mm yr−1); 3) the growth rate–size curve; and 4) the influence of environmental factors. Many crustose species comprise discrete areolae that contain the algal partner growing on the surface of a non-lichenized fungal hypothallus. Recent data suggest that 'primary' areolae may develop from free-living algal cells on the substratum while 'secondary' areolae develop from zoospores produced within the thallus. In more extreme environments, the RGR of crustose species may be exceptionally slow but considerably faster rates of growth have been recorded under more favourable conditions. The growth curves of crustose lichens with a marginal hypothallus may differ from the 'asymptotic' type of curve recorded in foliose and placodioid species; the latter are characterized by a phase of increasing RGR to a maximum and may be followed by a phase of decreasing growth. The decline in RGR in larger thalli may be attributable to a reduction in the efficiency of translocation of carbohydrate to the thallus margin or to an increased allocation of carbon to support mature 'reproductive' areolae. Crustose species have a low RGR accompanied by a low demand for nutrients and an increased allocation of carbon for stress resistance; therefore enabling colonization of more extreme environments.  相似文献   

16.
The past decade has seen the development and application of over a dozen new methods for quantitative age-determinations of geomorphic surfaces. Some surface exposure dating methods are numerical, including the accumulation of cosmogenic radionuclides 10Be, 14C, 26Al, 36Cl, and 41Ca, accumulation of cosmogenic stable nuclides 3He and 21Ne, 14C dating of organic matter encapsulated in rock coatings, and dendrogeomorphology. Calendar ages are obtained by dendrogeomorphological analysis. Calibrated ages can be obtained by analysis of rock-varnish chemistry, lichenometry, weathering, and soils. Various methods can be used in combination to overcome individual limitations. Whereas conventional methods provide age control on stratigraphic profiles, surface-exposure dating methods are especially suitable for geographic problems, such as analyzing not only temporal, but also spatial variations in the rates of geomorphic processes. [Key words: geomorphology, process, geochronology, cosmogenic nuclides, Quaternary, surface exposure dating, rock varnish, weathering, soils.]  相似文献   

17.
Trimmed lichen communities (lichen limits) are abrupt changes from a lichen community to a scoured bare rock surface and have been used to determine bankfull channel capacity on bedrock channels and their response to the combined disturbances of flow regulation and climate change. They can also be used to set flushing flows in bedrock channels. In sandstone gorges of the Nepean River, Australia, the crustose lichen, Lecidea terrena Nyl, was common at both gorge and cemetery (sandstone headstones) sites, enabling construction of growth curves for above and below dam areas. Growth curves were used to date lichen colonisation of sandstone surfaces in rivers. The oldest, highest lichen limit at all sites represented the pre‐flow regulation lichen community because its characteristics above and below Nepean Dam were similar and were trimmed to a level that produced consistent discharges across a range of catchment areas. They corresponded to return periods of less than 2 years on the annual maximum series and was developed during the flood‐dominated regime (FDR) of 1857–1900. Lichen limits form by the phycobiont dominating the mycobiont and hence degrading lichen thalli due to water inundation causing weak or dead thalli to be scrubbed from the rock surface. Trimming to the unregulated lichen limit represents a small flood of frequent occurrence appropriate for flushing bedrock channels. A lower lichen limit was only found below a diversion weir and was formed by frequent dam spills between 1950 and 1952 during an extraordinary wet period at the start of the FDR between 1949 and 1990. Lichens colonised exposed sandstone between the level of frequent flows from 1949 to 1952, and the high lichen limit. On the Avon River, an additional lower limit reflected a massive downward shift in flow duration following the start of interbasin diversions to Wollongong in 1962.  相似文献   

18.
A unique 25-year lichen growth monitoring programme involving 2,795 individuals of the Rhizocarpon subgenus at 47 sites on 18 glacier forelands in the Jotunheimen–Jostedalsbreen regions of southern Norway is reported. The data are used to address fundamental questions relating to direct lichenometry: spatial and temporal variability in lichen growth rates, climatic effects on lichen growth rates, lichen growth models, and implications for lichenometric dating. Mean annual (diametral) growth rate ranged from 0.43 to 0.87 mm yr−1 between sites, which is attributed primarily to local habitat differences. Interannual variability in annual mean growth rate exceeded 1.0 mm yr−1 at some sites. Annual mean growth rates for all sites combined varied from 0.52 to 0.81 mm yr−1 and was positively correlated with annual mean temperature and winter mean temperature (both r = 0.64, p <0.005) but not with summer seasonal temperature: positive correlations with annual and winter precipitation were less strong and the correlation with summer precipitation was marginally significant (r = 0.41 p <0.10). Growth-rate models characterized by annual growth rates that remain approximately constant or increase with lichen size up to at least 120 mm tended to fit the data more closely than a parabolic model. This is tentatively attributed to a long 'linear/mature' phase in the growth cycle. Comparison with growth rates inferred from indirect lichenometry suggest that such high measured growth rates could not have been maintained over the last few centuries by the largest lichens used in southern Norway for lichenometric dating. Several hypotheses, such as the effects of competition and climate change, which might explain this paradox, are discussed.  相似文献   

19.
This paper presents new lichenometric population data from the Antarctic Peninsula (67°S), and describes a new approach to lichen growth-rate calibration in locations where dated surfaces are extremely rare. We use historical aerial photography and field surveys to identify sites of former perennial snowpatches where lichen populations now exist. As an independent check on lichen mortality by snowkill, and the timing of snow patch disappearance, we use a positive-degree day (PDD) approach based on monthly climate data from Rothera Research Station. We find that maximum growth rates for lichens <40 mm in diameter on Adelaide Island are around 0.8 mm/yr. Furthermore, we propose that our combined methodology may be more widely applicable to the Polar Regions where the construction of lichenometric dating (age-size) curves remains a problem.  相似文献   

20.
Though the ability of lichens to disaggregate and dissolve the substrate is understood, zones of disparate weathering mechanisms examined in this study were previously unobserved. On a steep (60°) northern (340°N) slope on Red Mountain, Arizona, a 5×5 m study plot was chosen for its maximum lichen coverage (33%) of the same lichen genus Xanthoparmelia in varying growth stages, on a sandstone substrate of consistent lithology. From twenty-two (22) representative lichen thalli, portions were removed to examine sub-thallic rhizine density, and others were resin-inbedded for optical, scanning and backscatter electron microscopy.
In every sample, sandstone beneath and adjacent to the lichen thallus displayed disparate weathering mechanisms. Beneath the center of the lichen cortex, where rhizine density was observed to be the greatest, rhizine penetration disaggregated the sand-stone clasts from the matrix, but little chemical dissolution was apparent. This study found physical weathering predominates beneath the lichen cortex, and chemical weathering predominates at the thallus fringe and beyond the thallus boundary. It was generally found that physical weathering decreased (disaggregation) and chemical weathering increased (dissolution) from the cortex center to the thallus edge. Toward the thallus fringe, minimal clast disaggregation was observed, and substrate dissolution was obvious.  相似文献   

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