Spatial heterogeneity of epibenthos on artificial reefs: fouling communities in the early stages of colonization on an East Australian shipwreck     

Simon J. Walker  Thomas A. Schlacher  & Monika A. Schlacher-Hoenlinger 《Marine Ecology》2007,28(4):435-445

Artificial reefs are spatially complex habitats and serve as good model systems to study patterns of community succession and the response of epibiota to environmental clines over small spatial scales. Here, we quantified spatial heterogeneity in community composition and diversity of fouling communities across a number of environmental gradients that included water depth, surface orientation of habitats, exposure to currents, and shelter. Assemblage structure was quantified by spatially replicated photo transects on a recently scuttled large navy ship off the East Australian coast, lying in 27 m of water. A rich assemblage of epifauna had colonized the wreck within a year, dominated by barnacles, sponges and bryozoans. Community structure varied significantly over small spatial scales of meters to tens of meters. Depth, surface orientation and exposure were the major environmental drivers. Assemblages were substantially less diverse and abundant on the deepest (23 m near the seafloor) part of the hull with residual antifouling paint, on sheltered surfaces inside the wreck, and on the sediment‐laden horizontal surfaces. Overall, the wrecks’ habitat complexity corresponds with small‐scale heterogeneity in the fouling communities. This study supports the notion that wrecks enhance local diversity and biomass within the habitat mosaic of their location, and habitat complexity may be an important mechanism for this, as demonstrated by the large spatial variability in the assemblages documented here.  相似文献   

Rare species, habitat diversity and functional redundancy in marine benthos   总被引：1，自引：0，他引：1  

Kari E. Ellingsen  Judi E. Hewitt  Simon F. Thrush 《Journal of Sea Research》2007,58(4):291-301

Macro-ecological theories relating species richness, abundance, range size, biological traits and environmental tolerance have rarely been tested in marine soft-sediments, despite the spatial extent of these habitats and the inherent richness of resident communities. This study examines the contribution of rare species to marine soft-sediment communities from New Zealand, focussing on the relationships of range size with abundance, environment, habitat diversity and life history traits. 54% of the 351 species sampled exhibited restricted ranges (found at ≤ 2 sites). In contrast to many terrestrial systems, we observed only a weak positive relationship between abundance and frequency of occurrence. Restricted-range species were not randomly distributed, with their distribution related to habitat characteristics, suggesting an important link between habitat diversity and rarity. They exhibited a similar range of traits to the total observed species pool, suggesting that they are not only important to biodiversity but could play a role in stability. Restricted range species were generally not small and this, together with the number of different biological traits represented, suggests that rare species are important to the functioning of marine systems. Thus, our results highlight the importance of considering rare species in habitat-based approaches to conservation.  相似文献   

Geochemistry of suspended matter from the Baltic Sea 2. Results of bulk trace metal analysis by AAS     

Lutz Brügmann  Paul C. Bernard  Rene van Grieken 《Marine Chemistry》1992,38(3-4)

In 1984, on a transect covering the whole Baltic Sea and parts of the adjacent North Sea, 160 water samples were taken and analysed for their concentrations of particulate and dissolved metals. In addition, the suspended materials were investigated for their elemental bulk composition.The particulate fractions represented from about 5% (Cd, Cu and Ni) to 50% (Fe and Pb) of the total (particulate plus dissolved) concentrations. For some elements (Ba, Cd, Cu, Pb and Zn), the particulate matter from the surface microlayer was enriched with respect to those suspended materials taken from 0.2 m depth. This could reflect the atmospheric input of metal-rich aerosols. In anoxic deep waters, maximum contents of Zn (6400 μg g⁻¹), Cu (1330 μg g⁻¹) and Cd (12 μg g⁻¹) were observed in the particulate matter, indicating sulphidic forms. On the other hand, under oxic conditions the distribution coefficients (K_d) decreased with the water depth (Cd, Fe and Pb).Relative to global background levels, the particulate matter contained metal “excesses” amounting to more than 90% of the total contents (Cd, Mn, Pb and Zn). Automated electron probe X-ray microanalysis (EPXMA) revealed that the elemental composition of sediments is mainly governed by post-depositional processes of early diagenesis and is only weakly related to the composition of suspended matter in the overlying water body. For instance, in relation to surface mud sediments of the central Baltic net-sedimentation basins, Zn, Cd, Cu and Mn had 30–100% higher levels in the suspended materials. The general pattern of metal contents of particulate matter taken from 10 m depth on a transect between the Bothnian Bay and the North Sea were—possibly as a result of anthropogenic inputs—rather similar for Pb, Zn and Cu. For Fe and Mn, the distribution patterns along the transect were probably governed by the natural loading characteristics and by the biogeochemistry of those elements.  相似文献   

Multi-proxy evidence of long-term changes in ecosystem structure in a Danish marine estuary, linked to increased nutrient loading   总被引：3，自引：0，他引：3  

Marianne Ellegaard  Annemarie L. Clarke  Nina Reuss  Simon Drew  Kaarina Weckstrm  Stephen Juggins  N. John Anderson  Daniel J. Conley 《Estuarine, Coastal and Shelf Science》2006,68(3-4):567

This paper presents a study of changes in eutrophication over the past 100 years in a fertile estuary. The Danish estuary Mariager Fjord is a long, narrow sill-fjord with a permanently anoxic basin. In 1997 anoxia spread from the basin to the entire inner estuary, killing almost all eukaryotes and prompting debate on the causes. This paper reports a multi-proxy survey of ²¹⁰Pb-dated sediment cores from the anoxic basin. Analyses of diatoms, dinoflagellates, pigments and geochemical proxies were used to determine changes in ecosystem structure over the past 100 years. The aim was to establish ‘base-line conditions’, for management purposes, of the biological structure prior to 1900, and to examine possible causes of changes observed. Geochemical proxies total nitrogen (TN), total carbon (TC) and biogenic silica (BSi) were consistently high throughout the sediment record. Increased concentrations of pigments and natural isotopes (δ¹³C, δ¹⁵N) suggested increasing production and nutrient loading. The main changes in the biological proxies occurred between 1915 and the 1940s, and indicated that the estuary has been somewhat eutrophic since 1900, but that the eutrophication process increased over the past 100 years. A reconstruction of TN concentration by a diatom-based transfer function supports this interpretation, with inferred TN ca. 1900 around 60 μmol l⁻¹, and an increase in TN concentration over the past century to ca. 130 μmol l⁻¹ by 1995. Inferred TN decreased to ca. 100 μmol l⁻¹ by 2001, similar to present day monitoring data.  相似文献   

An evaluation of the Emerald/Western Bank juvenile haddock closed area     

Frank  Kenneth T.; Shackell  Nancy L.; Simon  James E. 《ICES Journal of Marine Science》2000,57(4):1023-1034

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Reply to comment on “Deep groundwater flow and geochemical processes in limestone aquifers: evidence from thermal waters in Derbyshire, England, UK” by John Gunn, Simon H Bottrell, David J Lowe, Stephen R H Worthington. Hydrogeology Journal (2006), 14 (6), 868–881     

John Gunn  Simon H. Bottrell  David J. Lowe  Stephen R. H. Worthington 《Hydrogeology Journal》2008,16(5):1019-1021

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Zircon M257 ‐ a Homogeneous Natural Reference Material for the Ion Microprobe U‐Pb Analysis of Zircon     

Lutz Nasdala  Wolfgang Hofmeister  Nicholas Norberg  James M. Martinson  Fernando Corfu  Wolfgang Dörr  Sandra L. Kamo  Allen K. Kennedy  Andreas Kronz  Peter W. Reiners  Dirk Frei  Jan Kosler  Yusheng Wan  Jens Götze  Tobias Häger  Alfred Kröner  John W. Valley 《Geostandards and Geoanalytical Research》2008,32(3):247-265

We introduce and propose zircon M257 as a future reference material for the determination of zircon U‐Pb ages by means of secondary ion mass spectrometry. This light brownish, flawless, cut gemstone specimen from Sri Lanka weighed 5.14 g (25.7 carats). Zircon M257 has TIMS‐determined, mean isotopic ratios (2s uncertainties) of 0.09100 ± 0.00003 for ²⁰⁶pb/²³⁸U and 0.7392 ± 0.0003 for ²⁰⁷pb/²³⁵U. Its ²⁰⁶pb/²³⁸U age is 561.3 ± 0.3 Ma (unweighted mean, uncertainty quoted at the 95% confidence level); the U‐Pb system is concordant within uncertainty of decay constants. Zircon M257 contains ～ 840 μg g^?1 U (Th/U ～ 0.27). The material exhibits remarkably low heterogeneity, with a virtual absence of any internal textures even in cathodoluminescence images. The uniform, moderate degree of radiation damage (estimated from the expansion of unit‐cell parameters, broadening of Raman spectral parameters and density) corresponds well, within the “Sri Lankan trends”, with actinide concentrations, U‐Pb age, and the calculated alpha fluence of 1.66 × 10¹⁸ g^?1. This, and a (U+Th)/He age of 419 ± 9 Ma (2s), enables us to exclude any unusual thermal history or heat treatment, which could potentially have affected the retention of radiogenic Pb. The oxygen isotope ratio of this zircon is 13.9%o VSMOW suggesting a metamorphic genesis in a marble or calc‐silicate skarn.  相似文献   

O,H, C isotope study of rocks from the KTB pilot hole: crustal profile and constraints on fluid evolution     

K. Simon  J. Hoefs 《Contributions to Mineralogy and Petrology》1993,114(1):42-52

The pilot hole of the Continental Deep Borehole (KTB) drilling project is located in the Bavarian Oberpfalz at the western margin of the Bohemian Massif. The 4-km deep borehole penetrated various paragneisses and minor orthogneisses with intercalations of amphibolites and metagabbros. The different lithologies have systematically different whole-rock oxygen isotope values and give little evidence for large scale water-rock interaction. Minor fluid interaction is well documented during retrograde metamorphism by non-equilibrium fractionations between refractory minerals (quartz, garnet and hornblende) and altered minerals (chlorite/biolite and feldspar). Ubiquitous vein mineralisation indicates fluid-induced retrogression at temperatures between 150°C and 400°C. The D values of hydroxylbearing minerals are very uniform in all lithologic units. The calculated hydrogen isotope composition of the fluid in equilibrium with matrix and vein minerals increases from -45 for metabasic rocks, to -20 for gneisses, to about -5 for vein minerals. The oxygen isotope composition of the fluid has been buffered by the rock and decreases with decreasing temperature because of increasing fractionations at low temperatures and low water-rock ratios. Modern fluids sampled from open cavities within the borehole have isotopic compositions that suggest a continuous fluid evolution during retrogression in a closed system. The ¹³C values of calcite and graphite also indicate closed system mixing processes.  相似文献   

Wollastonite at Nuliyam,Kerala, southern India: a reassessment of CO2-infiltration and charnockite formation at a classic locality     

Simon L. Harley  M. Santosh 《Contributions to Mineralogy and Petrology》1995,120(1):83-94

The occurrence of a charnockitised felsic gneiss adjacent to a marble/calc-silicate horizon at Nuliyam, southern India, has been cited in recent literature as a classic example of the dehydration of crustal rocks resulting from the advective infiltration of CO₂-rich fluids generated from a local carbonate source. Petrographic study of the Nuliyam calc-silicate, however, reveals it to consist of abundant wollastonite and scapolite and contain locally discordant veins rich in wollastonite. At the pressure—temperature conditions proposed for charnockite formation in recent studies, 5 kbar and 725°C, this wollastonite-bearing mineral assemblage was stable in the presence of a fluid phase only if X _CO2 was near 0.25 and could not have coexisted with the fluid causing biotite breakdown and charnockite development in adjacent rocks (X _CO2>0.85). The stable coexistence of wollastonite and scapolite prohibits the calc-silicate from being a source for fluid driving charnockitisation at the required P-T conditions. Textural observations such as the limited replacement of wollastonite by calcite+quartz symplectites and mosaics, are consistent with late fluid infiltration into the calc-silicate. The extensive isotopic, chemical and mineral abundance data of Jackson and Santosh (1992) are re-interpreted and integrated with these observations to develop a model involving the infiltration of an externally derived CO₂-rich fluid during high-temperature decompression. Increased charnockite development next to the calc-silicate has arisen because the calc-silicate acted as a relatively unreactive and impermeable barrier to fluid transport and caused fluid ponding beneath antiformal closures. The Nuliyam charnockite/calc-silicate locality is an example of a structural trap in a metamorphic setting rather than a site where charnockite formation can be attributed to local fluid sources.  相似文献   

Hydrothermal alteration of Variscan granites,southern Schwarzwald,Federal Republic of Germany     

K. Simon 《Contributions to Mineralogy and Petrology》1990,105(2):177-196

Hercynian S-type granites from the southeastern Schwarzwald granite series represent cogenetic biotite-and two-mica granites. Oxygen- and hydrogen-isotope data show that hydrothermal alteration invoking isotopically light surface waters resulted in a drastic reduction in ¹⁸O and D and pronounced disequilibrium between the minerals. Effective water-rock ratios are calculated to be high, about 0.8 vol units. A shift in the¹⁸O/¹⁶O and the chemical composition of the fluid due to water-rock interaction is continuously traced from pure H₂O with meteoric isotopic character in the deep-seated biotite granites to slightly saline water with rock-equilibrated isotopic composition in the two-mica granites at a shallower level. Substantial retrograde hydrometamorphism in the temperature range 500° to 200° C resulted mineralogically in high-temperature chloritization of biotite, and low-temperature muscovitization as well as feldspar alteration, respectively. Another result of the re-equilibration of cations is strong disturbance of the Rb–Sr system which affects measured ages and initial⁸⁷Sr/⁸⁶Sr values. Hydrothermal differentiation and alteration probably overlap to a very large extent magmatic differentiation processes.  相似文献