How Well Does Chlorophyll Explain the Seasonal Variation in Phytoplankton Activity?     

Maren Moltke Lyngsgaard  Stiig Markager  Katherine Richardson  Eva Friis Møller  Hans Henrik Jakobsen 《Estuaries and Coasts》2017,40(5):1263-1275

The seasonal variation in phytoplankton activity is determined by analysing 1385 primary production (PP) profiles, chlorophyll a (Chl) concentration profiles and phytoplankton carbon biomass concentrations (C) from the period 1998–2012. The data was collected at six different stations in the Baltic Sea transition zone (BSTZ) which is a location with strong seasonal production patterns with light as the key parameter controlling this productivity. We show that the use of Chl as a proxy for phytoplankton activity strongly overestimates the contribution from the spring production to annual pelagic carbon flow. Spring (February and March) Chl comprised 16–30% of the total annual Chl produced, whereas spring C was much lower (8–23%) compared to the annual C. Spring PP accounted for 10–18% of the total annual PP, while the July–August production contributed 26–33%, i.e. within the time frame when zooplankton biomass and grazing pressure are highest. That is, Chl failed in this study to reflect the importance of the high summer PP. A better proxy for biomass may be C, which correlated well with the seasonal pattern of PP (Pearson correlation, p < 0.05). Thus, this study suggests to account for the strong seasonal pattern in C/Chl ratios when considering carbon flow in coastal systems. Seasonal data for PP were fitted to a simple sinusoidal wave model describing the seasonal distribution of PP in the BSTZ and were proposed to present a better parameterizaton of PP in shallow stratified temperate regions than more commonly applied proxies.  相似文献   

龙门山小鱼洞断裂在汶川地震中的地表破裂及地质意义   总被引：8，自引：5，他引：3       下载免费PDF全文

李勇  黄润秋  A.L.Densmore  周荣军  闫亮  张毅  N.Richardson  董顺利  乔宝成  M.A.Ellis  何玉林  陈浩  马博琳 《第四纪研究》2009,29(3):502-512

在龙门山中段小鱼洞地区映秀－北川断裂和彭县－灌县断裂发生了同震位移,同时在该地区新出现了一条北西向的同震破裂带——小鱼洞断裂的破裂带,并分割了两条近于平行的北东向逆冲－走滑型的主断裂。汶川特大地震发生后重点对小鱼洞断裂的地表破裂开展了详细的野外地质填图,利用全站仪和GPS对地表破裂进行了精确的测量。初步调查的结果表明,小鱼洞断裂位于映秀－北川断裂与彭县－灌县断裂之间,走向近于北西向,延伸约15km,以脆性破裂为特征,穿过小鱼洞大桥,并切割了多种类型的地貌单元,使道路发生拱曲、破坏和桥梁垮塌或移位。单个破裂长度在几米到300m不等,其南西盘为上升盘,北东盘为下降盘,平均垂直位错为1.0m,平均水平位错为2.3m,垂直与水平位错量之比1 ∶1～1 ∶2,显示为逆冲分量小于或等于左旋走滑分量,并以左旋走滑作用为主。同时,研究表明小鱼洞断裂属于映秀－北川断裂与彭县－灌县断裂之间的捩断层,其主要特征包括： 1)是在汶川地震中由于龙门山逆冲体之间的差异逆冲运动过程中而形成的断裂; 2)其走向近于北西向,垂直于龙门山北东向的主干断裂,而平行于逆冲体的逆冲运动方向; 3)具有高角度断面的断层,以左旋走滑作用为主。  相似文献   

Effects of sediment on fish communities in East Cape streams,North Island,New Zealand     

J. Richardson  I. G. Jowett 《新西兰海洋与淡水研究杂志》2013,47(2):431-442

We examined the effects of sediment on fish communities at 38 sites in the East Cape region of New Zealand. Many streams in this region are subject to high sediment loads, a natural situation that has been exacerbated by the conversion of native forest to pasture. The fish community consisted of diadromous native species common throughout New Zealand that require access to and from the sea. We found that site altitude and distance inland were important factors in determining the abundance and composition of the fish community. We used four measures as indices of sediment load: suspended sediment concentration, median substrate size, substrate stability, and the ratio of the wetted width to the width of the active (non‐vegetated) stream channel. Although all indices were correlated, the ratio of wetted width to width of active channel was most closely related to fish abundance and diversity. Fish abundance and diversity reduced as sediment load increased among streams, with up to nine fish species in streams with low sediment loads and only two species in streams with high sediment loads. In‐stream habitat also varied with sediment load; as load increased, depth and substrate size decreased and velocity increased. These results suggest that activities that increase sediment loads in rivers will have a negative impact on native fish communities in New Zealand.  相似文献   

Evidence for ecosystem engineering in a lentic habitat by tadpoles of the western toad     

Sylvia L. R. Wood  John S. Richardson 《Aquatic Sciences - Research Across Boundaries》2010,72(4):499-508

A number of aquatic organisms have demonstrated an ability to modulate sediment deposition in tropical and temperate stream systems, but little is known about the impact of similar benthic grazing processes in temperate pond systems. Using exclosure experiments in mesocosms, we used tadpoles of the western toad (Anaxyrus boreas) to investigate two questions: (1) can grazing activities of tadpoles of the western toad in ponds reduce sediment accrual on grazing surfaces (e.g. via bioturbation or ingestion); and (2) do these grazing activities contribute to enhanced periphyton biomass and chlorophyll a content of benthic matter. Mesocosms received pulses of fine sediment weekly over 8 weeks to simulate allochthonous surface inputs following rainstorms. Accumulated sediment and periphyton matter on exposed and grazing-restricted substrate surfaces were measured weekly. Tadpole grazing activity significantly reduced the standing stock of periphyton on open grazing surfaces by 80% in controls. Under sediment-additions, tadpoles effectively removed settled particulate matter from exposed grazing surfaces, but no corresponding increase in periphyton biomass was observed. These results suggest that tadpoles can impact the dynamics of sediment distribution in ponds as well as streams, providing support for a role as “ecosystem engineers” in these habitats.  相似文献   

Shocks and sheaths in the heliosphere     

John D. Richardson 《Journal of Atmospheric and Solar》2011,73(11-12):1385-1389

This paper compares three kinds of shocks and sheaths; the bow shock and magnetosheaths of planets, interplanetary coronal mass ejection (ICME) shocks and sheaths, and the termination shock and heliosheath. We compare the energy balance across these shocks, fluctuations in the sheaths, asymmetries, and shock evolution. All three shocks and sheaths display asymmetries imposed by outside magnetic fields. Energy balance is often achieved through reflection of ions of the shocks, but in some cases no reflected ions are observed. The shocks may have similar small scale fluctuations which drive density changes in the sheaths.  相似文献   

Structure and evolution of mass transport deposits in the South Caspian Basin,Azerbaijan     

Steve E. J. Richardson  Richard J. Davies  Mark B. Allen  Simon F. Grant 《Basin Research》2011,23(6):702-719

The Quaternary to late Pliocene sedimentary succession along the margin of the South Caspian Basin contains numerous kilometre‐scale submarine slope failures, which were sourced along the basin slope and from the inclined flanks of contemporaneous anticlines. This study uses three‐dimensional (3D) seismic reflection data to visualise the internal structure of 27 mass transport deposits and catalogues the syndepositional structures contained within them. These are used to interpret emplacement processes occurring during submarine slope failure. The deposits consist of three linked structural domains: extensional, translational and compressive, each containing characteristic structures. Novel features are present within the mass transport deposits: (1) a diverging retrogression of the headwall scarp; (2) the absence of a conventional headwall scarp around growth stratal pinch outs; (3) restraining bends in the lateral margin; (4) a downslope increase in the throw of thrust faults. The results of this study shed light on the deformation that occurred during submarine slope failure, and highlight an important geological process in the evolution of the South Caspian Basin margin.  相似文献   

Spectral Irradiance, Phytoplankton Community Composition and Primary Productivity in a Salt Marsh Estuary, North Inlet, South Carolina, USA     

Evelyn Lawrenz  Erik M. Smith  Tammi L. Richardson 《Estuaries and Coasts》2013,36(2):347-364

We investigated spatial and temporal changes in spectral irradiance, phytoplankton community composition, and primary productivity in North Inlet Estuary, South Carolina, USA. High concentrations of colored dissolved organic matter (CDOM) were responsible for up to 84 % of the attenuation of photosynthetically available radiation (PAR). Green-yellow wavelengths were the predominant colors of light available at the two sampling sites: Clam Bank Creek and Oyster Landing. Vertical attenuation coefficients of PAR were 0.7–2.1 m^?1 with corresponding euphotic zone depths of 1.5–6.7 m. Phytoplankton biomass (as chlorophyll a [chl a]) varied seasonally with a summer maximum of 16 μg chl a l^?1 and a winter minimum of 1.4 μg chl a l^?1. The phytoplankton community consisted mainly of diatoms, prasinophytes, cryptophytes and haptophytes, with diatoms and prasinophytes accounting for up to 67 % of total chl a. Changes in phytoplankton community composition showed strongest correlations with temperature. Light-saturated chl a-specific rates of photosynthesis and daily primary productivity varied with season and ranged from 1.6 to 14 mg C (mg chl a) ^?1?h^?1 (32–803 mg C m^?3?day^?1). Calculated daily rates added up to an annual carbon fixation rate of 84 g C m^?3?year^?1. Overall, changes in phytoplankton community composition and primary productivity in North Inlet showed a strong dependence on temperature, with PAR and spectral irradiance playing a relatively minor role due to short residence times, strong tidal forcing and vertical mixing.  相似文献   

Using Statistical Multivariable Models to Understand the Relationship Between Interplanetary Coronal Mass Ejecta and Magnetic Flux Ropes     

P. Riley  I. G. Richardson 《Solar physics》2013,284(1):217-233

In-situ measurements of interplanetary coronal mass ejections (ICMEs) display a wide range of properties. A distinct subset, “magnetic clouds” (MCs), are readily identifiable by a smooth rotation in an enhanced magnetic field, together with an unusually low solar wind proton temperature. In this study, we analyze Ulysses spacecraft measurements to systematically investigate five possible explanations for why some ICMEs are observed to be MCs and others are not: i) An observational selection effect; that is, all ICMEs do in fact contain MCs, but the trajectory of the spacecraft through the ICME determines whether the MC is actually encountered; ii) interactions of an erupting flux rope (FR) with itself or between neighboring FRs, which produce complex structures in which the coherent magnetic structure has been destroyed; iii) an evolutionary process, such as relaxation to a low plasma-β state that leads to the formation of an MC; iv) the existence of two (or more) intrinsic initiation mechanisms, some of which produce MCs and some that do not; or v) MCs are just an easily identifiable limit in an otherwise continuous spectrum of structures. We apply quantitative statistical models to assess these ideas. In particular, we use the Akaike information criterion (AIC) to rank the candidate models and a Gaussian mixture model (GMM) to uncover any intrinsic clustering of the data. Using a logistic regression, we find that plasma-β, CME width, and the ratio O ⁷/O ⁶ are the most significant predictor variables for the presence of an MC. Moreover, the propensity for an event to be identified as an MC decreases with heliocentric distance. These results tend to refute ideas ii) and iii). GMM clustering analysis further identifies three distinct groups of ICMEs; two of which match (at the 86 % level) with events independently identified as MCs, and a third that matches with non-MCs (68 % overlap). Thus, idea v) is not supported. Choosing between ideas i) and iv) is more challenging, since they may effectively be indistinguishable from one another by a single in-situ spacecraft. We offer some suggestions on how future studies may address this.  相似文献   

A tribute to John Stuart Gray (1941-2007)     

Richardson B  Wu R  Shin P  Lam P  Fleddum A  Elliott M  Sheppard C 《Marine pollution bulletin》2008,56(1):1-4

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Estimation of new production in the North Sea: consequences for temporal and spatial variability of phytoplankton   总被引：2，自引：0，他引：2  

Richardson  Katherine; Pedersen  Flemming Bo 《ICES Journal of Marine Science》1998,55(4):574-580

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