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Irregular satellites—moons that occupy large orbits of significant eccentricity e and/or inclination I—circle each of the giant planets. The irregulars often extend close to the orbital stability limit, about 1/3-1/2 of the way to the edge of their planet's Hill sphere. The distant, elongated, and inclined orbits suggest capture, which presumably would give a random distribution of inclinations. Yet, no known irregulars have inclinations (relative to the ecliptic) between 47 and 141°.This paper shows that many high-I orbits are unstable due to secular solar perturbations. High-inclination orbits suffer appreciable periodic changes in eccentricity; large eccentricities can either drive particles with ∼70°<I<110° deep into the realm of the regular satellites (where collisions and scatterings are likely to remove them from planetocentric orbits on a timescale of 107-109 years) or expel them from the Hill sphere of the planet.By carrying out long-term (109 years) orbital integrations for a variety of hypothetical satellites, we demonstrate that solar and planetary perturbations, by causing particles to strike (or to escape) their planet, considerably broaden this zone of avoidance. It grows to at least 55°<I<130° for orbits whose pericenters freely oscillate from 0 to 360°, while particles whose pericenters are locked at ±90° (Kozai mechanism) can remain for longer times.We estimate that the stable phase space (over 10 Myr) for satellites trapped in the Kozai resonance contains ∼10% of all stable orbits, suggesting the possible existence of a family of undiscovered objects at higher inclinations than those currently known.  相似文献   
154.
Measurement of acetylcholinesterase (AChE) is widely used as a biomarker of exposure to neurotoxic compounds in aquatic environments. In the present study, AChE activity was monitored in the clam Tapes philippinarum in the Lagoon of Venice, with the aim of evaluating the possible exposure of animals to neurotoxic compounds. Two sampling strategies were followed. In the first case, temporal and spatial variations of AChE activity were measured in clams collected seasonally (from October 2003 to June 2004) in 3 sites of the Lagoon: Campalto (site 1), near a sewage treatment plant, Marghera (site 2), a highly polluted area, and Poveglia (site 3), a reference site. The condition index (CI) of clams was also calculated. In the second case, AChE activity was measured in clams collected at the end of seasonal samplings in 12 sites located throughout the Lagoon: areas licensed for clam farming (sites 5 to 11), nearby canals characterised by waste waters from agricultural areas (sites 4, 12, 13, 14), and a reference site outside the Lagoon (site 15). Results revealed a significant reduction in AChE activity in clams from Marghera in October, January and April, indicating their probable exposure to neurotoxic compounds originating in the industrial zone of Porto Marghera. A seasonal trend in AChE activity was also observed, enzyme activity being higher in January and lower in June. Clams from Campalto generally had higher CI values than those from Marghera and Poveglia. However, no particular correlation between AChE activity and CI was found. Lastly, marked differences in AChE activity were recorded when comparing enzyme activity of clams from various sites in the Lagoon of Venice with those of clams collected at site 15. Significant reductions in AChE activity were observed in animals collected in both nearby canals and licensed areas, indicating the homogeneous spatial distribution of potentially neurotoxic compounds throughout the Lagoon. It is probable that sediment mobilisation caused by clam harvesting and the passage of shipping plays an important role in resuspension of persistent contaminants having neurotoxic activity.  相似文献   
155.
Tethyan ophiolites and Pangea break-up   总被引:6,自引:0,他引:6  
Abstract The break‐up of Pangea began during the Triassic and was preceded by a generalized Permo‐Triassic formation of continental rifts along the future margins between Africa and Europe, between Africa and North America, and between North and South America. During the Middle–Late Triassic, an ocean basin cutting the eastern equatorial portion of the Pangea opened as a prograding branch of the Paleotethys or as a new ocean (the Eastern Tethys); westwards, continental rift basins developed. The Western Tethys and Central Atlantic began to open only during the Middle Jurassic. The timing of the break‐up can be hypothesized from data from the oceanic remnants of the peri‐Mediterranean and peri‐Caribbean regions (the Mesozoic ophiolites) and from the Atlantic ocean crust. In the Eastern Tethys, Middle–Late Triassic mid‐oceanic ridge basalt (MORB) ophiolites, Middle–Upper Jurassic MORB, island arc tholeiite (IAT) supra‐subduction ophiolites and Middle–Upper Jurassic metamorphic soles occur, suggesting that the ocean drifting was active from the Triassic to the Middle Jurassic. The compressive phases, as early as during the Middle Jurassic, were when the drifting was still active and caused the ocean closure at the Jurassic–Cretaceous boundary and, successively, the formation of the orogenic belts. The present scattering of the ophiolites is a consequence of the orogenesis: once the tectonic disturbances are removed, the Eastern Tethys ophiolites constitute a single alignment. In the Western Tethys only Middle–Upper Jurassic MORB ophiolites are present – this was the drifting time. The closure began during the Late Cretaceous and was completed during the Eocene. Along the area linking the Western Tethys to the Central Atlantic, the break‐up was realized through left lateral wrench movements. In the Central Atlantic – the link between the Western Tethys and the Caribbean Tethys – the drifting began at the same time and is still continuing. The Caribbean Tethys opened probably during the Late Jurassic–Early Cretaceous. The general picture rising from the previous data suggest a Pangea break‐up rejuvenating from east to west, from the Middle–Late Triassic to the Late Jurassic–Early Cretaceous.  相似文献   
156.
Possible xenoestrogenic effects were investigated, for the first time, in two bivalve species from the Lagoon of Venice (Italy): the clam Tapes philippinarum and the cockle Cerastoderma glaucum. Bivalves were collected far from their reproductive phase at the very early stage of gametogenesis (January), and in the pre-spawning period (June) in six sites. Vitellogenin (Vg)-like proteins (a biomarker of exposure to estrogenic compounds) were measured by the alkali-labile phosphate method (ALP), in both haemolymph and digestive gland from males and females (when it was possible to distinguish sex by microscopic observation of gonadal tissue). Haemolymph calcium concentrations (a parameter considered closely related to the presence of Vg-like proteins) and the bivalve condition index (CI) were also measured. In both seasons, bivalves collected at Campalto (near a sewage treatment plant) and Marghera (a highly contaminated area) had higher Vg-like protein levels, particularly in haemolymph, than animals from the other sampling sites. Interestingly, CI had high values in these polluted sites. In June only, Vg-like proteins and Ca2+ levels in haemolymph exhibited similar trends in both bivalve species at most sampling sites. The responsiveness of bivalves to environmental xenoestrogens was higher in June, allowing better discrimination among sites. The present study demonstrates that animals from highly polluted areas have increased Vg-like protein levels. As endocrine disruption due to exposure to estrogenic compounds may cause fertility reduction, alterations in the sex ratio, and a decrease in reproductive rate, a condition of potential risk for bivalve populations in estuarine areas is highlighted.  相似文献   
157.
Volatile organic compounds (VOCs) may play the role of infochemicals and trigger chemotaxis and ecologically relevant responses in freshwater and marine invertebrates. Aquatic grazers use these signals as chemical cues to trace the presence of their food or to detect their predators. However, detailed data are still needed to fully understand the role of these relationships in marine plankton. We investigated the ability of the copepod Centropages typicus to perceive the odour of three planktonic diatoms (Skeletonema marinoi, Pseudonitzschia delicatissima and Chaetoceros affinis) and a dinoflagellate (Prorocentrum minimum). This information is ecologically relevant for orientation, habitat selection, predator avoidance and communication. In addition, as the pH of the medium influences the perception of chemical cues in aquatic environments, we tested the effect of seawater acidification resulting from increasing levels of CO2, and its influences on the olfactory reactions of copepods. For this reason, our tests were repeated in normal (pH 8.10) and acidified (pH 7.76) seawater in order to simulate future ocean acidification scenarios. Using replicated chemokinetic assays we demonstrated that VOCs produced by Ps. delicatissima and Pr. minimum attract copepods at normal pH, but this effect is lowered in acidified water. By contrast, the odour of S. marinoi mainly induces a reaction of repulsion, but in acidified water and at higher concentrations this toxic diatom becomes attractive for copepods. Our experiments demonstrate, for the first time, that copepods are sensitive to the volatile compounds contained in various microalgae; VOCs prompt chemokinesis according to algal species and odour concentrations. However, seawater acidification induces changes in copepods' perception of odours. These findings highlight the sensitivity of chemically mediated interactions to global changes  相似文献   
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