Carbon and nitrogen utilization in two species of Red Sea corals along a depth gradient: Insights from stable isotope analysis of total organic material and lipids     

Ada Alamaru  Yossi Loya  Eran Brokovich  Ruth Yam  Aldo Shemesh 《Geochimica et cosmochimica acta》2009,73(18):5333-5342

We examined the utilization of carbon and nitrogen in two common Red Sea coral species (Stylophora pistillata and Favia favus), differing in colony morphology and polyp size, along a depth gradient down to 60 m. We describe the changes in C/N ratios and in the stable isotope composition of carbon and nitrogen of coral’s tissue and algal symbionts. We also measured the carbon isotopic composition of the lipid fraction extracted from both coral tissue and algal symbionts in order to reveal the changes in the carbon source utilized by the host coral for lipid synthesis.The results show that for both species, δ¹³C decreases by 7–8‰ in animal tissue, algal symbionts and in the lipid fractions as depth increases. However, in contrast to previous reports, the difference between δ¹³C values of coral tissue and algal symbionts does not increase with depth. δ¹⁵N values of coral tissue and algal symbionts in both species do not correlate with depth suggesting that the heterotrophic capacity of these corals does not increase with depth. δ¹³C values of tissue lipids were depleted by an average of 3.5‰ compared to δ¹³C of the entire tissue at all depths. δ¹³C values of algal lipids were depleted by an average of 2‰ compared to δ¹³C of the entire zooxanthellae at all depths, indicating high efficiency of carbon recycling between the two symbiotic partners along the entire gradient. The depletion of lipids is attributed to the fractionation mechanism during lipid synthesis. In addition, for both species, δ¹³C values of algal lipids were enriched compared with δ¹³C of tissue lipids. In S. pistillata, the difference between δ¹³C values of tissue lipids and algal lipids increased linearly with depth, indicating a change in the sources of carbon utilized by the coral for lipid synthesis below 20 m from an autotrophic to a heterotrophic source. However, in F. favus, this average difference was 4 times larger compared to shallow S. pistillata and was constant along the entire depth gradient, suggesting that F. favus uses heterotrophically-acquired carbon for lipid synthesis regardless of depth. Overall, F. favus exhibited enriched δ¹³C and δ¹⁵N values compared to S. pistillata along the entire gradient. We attribute these differences to both morphological differences (i.e. colony morphology, tissue thickness and polyp size) between the two species and to a higher heterotrophy/autotrophy ratio in F. favus at all depths. The C/N ratio in S. pistillata tissue decreased with increasing water depth whereas in F. favus it remained constant. This reflects a higher heterotrophic capacity in the large polyped F. favus, at all depths.  相似文献   

Origins of subcalcic garnets and their relation to diamond forming fluids—Case studies from Ekati (NWT-Canada) and Murowa (Zimbabwe)     

Ofra Klein-BenDavid  D. Graham Pearson 《Geochimica et cosmochimica acta》2009,73(3):837-347

Subcalcic, high-Cr (G10) garnets are found as inclusions within diamonds and in peridotitic xenoliths. The strong spatial associations between G10 garnets and diamond make them an important tool in the investigation of diamond genesis. We present an integrated study of the major and trace element composition and oxygen-Sr-Nd-Hf isotopic ratios of eight G10 garnets from the Ekati mine (NWT-Canada) and four from the Murowa mine (Zimbabwe) in an attempt to determine their petrogenetic evolution and to further examine a possible relationship between the metasomatic agents responsible for G10 garnet signatures and diamond forming fluids.All garnets display sinusoidal to mildly sinusoidal REE patterns and have negative Ti, Sr and positive U anomalies. They have variably radiogenic ⁸⁷Sr/⁸⁶Sr (0.703261-0.731191) and non-radiogenic ε_Nd values (−8.1 to −27.1), except for one sample from Murowa that has a positive ε_Nd of 2.5. One Ekati sample has an extremely low ε_Hf value of −61.6. The Ekati garnets we have studied all appear to come from a single depth in the Slave lithospheric mantle. On the base of Cr-Ca relations they have crystallized at 4.9 GPa and display dunitic Ca intercept values. Their δ¹⁸O values range between +5.23‰ and +5.42‰.The Ekati G10 garnets record a complex, multi-stage metasomatic history involving the interaction of several components during their genesis. One metasomatic agent was enriched in HFSE, LREE, Sr, and depleted in Nb. This agent had the least radiogenic Sr. Another metasomatic agent had highly radiogenic Sr, and was enriched in LREE, Sr, Nb, Th and U.The G10 garnets have very low ε_Nd and ε_Hf values combined with radiogenic Sr, thus, they require an early lithospheric mantle enrichment event at some stage during their genesis or during the evolution of any precursor material that they formed from. The only Hf isotope composition measurable from the Ekati suite is so unradiogenic (ε_Hf = −61) that it yields a Lu/Hf model age of 3521 Ma. This indicates that the lithospheric enrichment event seen by the Ekati garnets or their precursors may have occurred in the early stages of the craton stabilization, during the diamond forming event [Westerlund K., Shirey S., Richardson S., Carlson R., Gurney J. and Harris J. (2006) A subduction wedge origin for Paleoarchean peridotitic diamonds and harzburgites from the Panda kimberlite, Slave craton: evidence from Re-Os isotope systematics. Contrib. Mineral. Petrol.152(3), 275-294]. Although our data cannot unequivocally discriminate between a variety of models for the genesis of subcalcic garnets it is clear that the host peridotite originated via melting at shallow depths followed by subduction and that the observed geochemical fingerprint of the garnets is strongly influenced by diamond forming fluids. Diamond forming fluids sampled from fibrous diamonds, have steep REE patterns, negative Ti and Sr anomalies and very low Sm/Nd ratios that are very similar to G10 garnet characteristics. These diamond forming fluids have been recently shown to have extreme Sr and Nd isotopic compositions [Klein-BenDavid O., Pearson D. G., Nowell G. M. and Cantigny P. (2008) Origins of diamond forming fluids—constraints from a coupled Sr-Nd isotope and trace element approach. Extended abstracts to the 9th International Kimberlite Conference, Frankfurt, Germany, 9IKC-A-00118.] that are closely concordant with G10 garnets. The fluids are also rich in LREE, P, K and water, sharing these features with mica-rich metasomes. These similarities suggest that ancient lithospheric metasomes could either provide a source region for, or be a product of diamond forming fluids. Diamond forming fluids appear to be intimately involved in the evolution of G10 garnets in the lithospheric mantle, either acting as a metasomatic agent, or being integral to triggering or enhancing garnet growth in a Cr-rich protolith. Such a link explains the strong association between G10 garnets and diamonds.  相似文献   

U-series and oxygen isotope chronology of the mid-Pleistocene Lake Amora (Dead Sea basin)     

Adi Torfstein  Alexandra Haase-Schramm  Nicolas Waldmann  Mordechai Stein 《Geochimica et cosmochimica acta》2009,73(9):2603-2630

This study establishes for the first time the chronology and limnological history of Lake Amora (Dead Sea basin, Israel), whose deposits (the Amora Formation) comprise one of the longest exposed lacustrine records of the Pleistocene time. The Amora Formation consists of sequences of laminated primary aragonite and silty-detritus, Ca-sulfate minerals, halite and clastic units. This sedimentary sequence was uplifted and tilted by the rising Sedom salt diapir, exposing ∼320 m of sediments on the eastern flanks of Mt. Sedom (the Arubotaim Cave (AC) section).The chronology of the AC section is based on U-disequilibrium dating (²³⁰Th-²³⁴U and ²³⁴U-²³⁸U ages) combined with floating δ¹⁸O stratigraphy and paleomagnetic constraints. The determination of the ²³⁰Th-²³⁴U ages required significant corrections to account for detrital Th and U. These corrections were performed on individual samples and on suites of samples from several stratigraphic horizons. The most reliable corrected ages were used to construct an age-elevation model that was further tuned to the oxygen isotope record of east Mediterranean foraminifers (based on the long-term similarity between the sea and lake oxygen isotope archives).The combined U-series-δ¹⁸O age-elevation model indicates that the (exposed) Amora sequence was deposited between ∼740 and 70 ka, covering seven glacial-interglacial cycles (Marine Isotope Stages (MIS) 18 to 5).Taking the last glacial Lake Lisan and the Holocene Dead Sea lacustrine systems as analogs of the depositional-limnological environment of Lake Amora, the latter oscillated between wet (glacial) and more arid (interglacial) conditions, represented by sequences of primary evaporites (aragonite and gypsum that require enhanced supply of freshwater to the lakes) and clastic sediments, respectively. The lake evolved from a stage of rapid shifts between high and low-stand conditions during ∼740 to 550 ka to a sabkha-like environment that existed (at the AC site) between 550 and 420 ka. This stage was terminated by a dry spell represented by massive halite deposition at 420 ka (MIS12-11). During MIS10-6 the lake fluctuated between lower and higher stands reaching its highest stand conditions at the late glacial MIS6, after which a significant lake level decline corresponds to the transition to the last interglacial (MIS5) low-stand lake, represented by the uppermost part of the Formation.δ¹⁸O values in the primary aragonite range between 6.0 and −1.3‰, shifting cyclically between glacial and interglacial intervals. The lowest δ¹⁸O values are observed during interglacial stages and may reflect short and intense humid episodes that intermittently interrupted the overall arid conditions. These humid episodes, expressed also by enhanced deposition of travertines and speleothems, seem to characterize the Negev Desert, and in contrast to the overall dominance of the Atlantic-Mediterranean system of rain patterns in the Dead Sea basin, some humid episodes during interglacials may be traced to southern sources.  相似文献   

Stratigraphy, depositional environments and level reconstruction of the last interglacial Lake Samra in the Dead Sea basin   总被引：1，自引：0，他引：1  

N. Waldmann  M. Stein  A. Starinsky 《Quaternary Research》2009,72(1):1-185

In this paper we describe the stratigraphy and sediments deposited in Lake Samra that occupied the Dead Sea basin between ∼ 135 and 75 ka. This information is combined with U/Th dating of primary aragonites in order to estimate a relative lake-level curve that serves as a regional paleohydrological monitor. The lake stood at an elevation of ∼ 340 m below mean sea level (MSL) during most of the last interglacial. This level is relatively higher than the average Holocene Dead Sea (∼ 400 ± 30 m below MSL). At ∼ 120 and ∼ 85 ka, Lake Samra rose to ∼ 320 m below MSL while it dropped to levels lower than ∼ 380 m below MSL at ∼ 135 and ∼ 75 ka, reflecting arid conditions in the drainage area. Lowstands are correlated with warm intervals in the Northern Hemisphere, while minor lake rises are probably related to cold episodes during MIS 5b and MIS 5d. Similar climate relationships are documented for the last glacial highstand Lake Lisan and the lowstand Holocene Dead Sea. Yet, the dominance of detrital calcites and precipitation of travertines in the Dead Sea basin during the last interglacial interval suggest intense pluvial conditions and possible contribution of southern sources of wetness to the region.  相似文献   

High-energy afterglow emission from gamma-ray bursts     

Yi-Zhong Fan †  Tsvi Piran †  Ramesh Narayan †  Da-Ming Wei † 《Monthly notices of the Royal Astronomical Society》2008,384(4):1483-1501

We calculate the very high-energy (sub-GeV to TeV) inverse Compton emission of GRB afterglows. We argue that this emission provides a powerful test of the currently accepted afterglow model. We focus on two processes: synchrotron self-Compton emission within the afterglow blast wave, and external inverse Compton emission which occurs when flare photons (produced by an internal process) pass through the blast wave. We show that if our current interpretations of the Swift X-ray telescope (XRT) data are correct, there should be a canonical high-energy afterglow emission light curve. Our predictions can be tested with high-energy observatories such as GLAST , Whipple, HESS and MAGIC. Under favourable conditions we expect afterglow detections in all these detectors.  相似文献   

Sedimentary and drainage evolution of the Condamine Valley Transition Zone (eastern Australia)     

P. Manchee  U. Shaanan  J. Esterle 《Australian Journal of Earth Sciences》2019,66(4):531-545

The basal succession of the Condamine Valley, which overlays the boundary between the Surat and Clarence-Moreton basins (eastern Australia), contains a clay-rich horizon ‘the Transition Zone’ that marks a pronounced unconformity between the Jurassic Walloon Coal Measures and the Quaternary Condamine Alluvium. This paper provides insights into the tectonic and drainage evolution of the Condamine Valley through integrated analysis of U–Pb ages of detrital zircon from three samples (494 concordant analyses), stable oxygen isotope analysis on eight authigenic clay samples, X-ray fluorescence of primary and trace elements, and hyperspectral mineral analysis from two drill cores (Lone Pine 17 and Daandine 164). The Transition Zone is interpreted to consist of both weathered Jurassic sediments and Cenozoic clay deposits. Two sequential cycles of erosion, deposition and pedogenesis, related to geomorphological and climatic conditions are recognised. Distinctive oxygen isotope signatures of the two weathering fronts demonstrated an initial Early Cretaceous phase (δ¹⁸O?=?11.9–15.7‰ VSMOW) associated with laterisation and possible uplift, followed by Paleogene (δ¹⁸O?=?16.4–17.3‰ VSMOW) silcretisation of reworked Jurassic sediments. Detrital zircon geochronology yielded Jurassic maximum constraints for the age of deposition of three samples that are indistinguishable within error, the oldest of which (163?±?8?Ma from the lateritic profile) corresponds to the age of the underlying Walloon Coal Measures. The two overlaying samples from a silcrete profile and granular alluvium yielded overlapping yet younger ages of 150?±?6?Ma and 156?±?9?Ma, respectively. Vitrinite reflectance used as a proxy for the thermal condition of the coal strata enabled an estimated 2–3?km of burial and subsequent (likely Early Cretaceous) uplift. Geochemical insights from the Condamine Valley correspond to broad-scale climatic and tectonic conditions, suggesting that ‘transition zone equivalents’ and corresponding groundwater dynamics may occur in similar sedimentary settings throughout eastern Australia.  相似文献   

Climate and surface properties: hydrological response of small arid and semi-arid watersheds     

《Geomorphology》2002,42(1-2):43-57

A positive relationship between annual rainfall and geomorphic processes (runoff and erosion rates) and environmental factors (water regime, soil and vegetation cover) is often assumed for arid and semi-arid areas with an annual rainfall in the range of 100–300 mm. This assumption disregards the fact that changes along a climatic gradient, at desert margins, are not limited to purely climatic factors. They are often accompanied by a parallel change in surface properties; especially the relative extent of rocky or soil covered surfaces that differ greatly in their response to rainfall. This raises the issue whether the change in surface properties along a climatic gradient enhances the assumed positive effects of rainfall increase or limits it. The hypothesis advanced in this paper is that runoff generation and rate in arid and semi-arid areas are primarily controlled by surface properties rather than by the absolute amounts of storm and annual rain amounts. Hydrological data collected at two instrumented watersheds, located one in an arid rocky area, and the second in a semi-arid soil covered area, support this hypothesis. The implications of data obtained for runoff generation and flow continuity under changing climatic conditions are analyzed. They point to the fact that the same regional climatic change may have different, and even opposite effects, on the hydrological response of different adjoining surface units. This response is expected to be strongly controlled by the specific local surface conditions that prevailed in the area prior to the climatic change.  相似文献   

Halite oolites and ripples in the Dead Sea, Israel     

Y. WEILER  E. SASS  I. ZAK 《Sedimentology》1974,21(4):623-632

Clastic features in recent halite deposits are observed along the beaches of an artificially dammed part of the Dead Sea. These features include halite oolites (termed halolites in this paper) and ripples. Halite precipitates initially either at the brine surface or on the floor. It is suggested that moderate increase of wave agitation shifts the balance towards brine-surface crystallization, and keeps the growing halite grains in constant motion. In this way rippled structures are formed. A further increase of wave energy leads to the growth of coated halite grains. The accumulation of the various halite grains along the beach, to form soft rippled floor and oolitic beach ridge is brought about during shoreward winds. During calm periods the bulk of the halite crystallizes directly on the floor. It develops into a hard crust which assumes the morphology of the substrate, including the ripple forms.  相似文献   

Growth pattern and 13C12C isotope fractionation of Cyanidium caldarium and hot spring algal mats     

J. Seckbach  I.R. Kaplan 《Chemical Geology》1973,12(3):161-169

A study was undertaken with the thermophilic green alga Cyanidium caldarium which grows optimally at low pH and high concentrations of CO₂. Carbon-isotope fractionation was not found to be a simple linear function of temperature. Maximum enrichment of ¹²C in cellular material occurred under optimum growth conditions (at approximately pH 2 and at temperatures between 40–50°C in a CO₂ atmosphere). A maximum measured fractionation of ?24‰ may account for low values ($\text{δ}{}^{13}\text{C}\text{< ?30‰}{}_{\mathrm{<mtext>PDB</mtext>}}$) in Precambrian kerogen presumably derived from algal mats.  相似文献   

On an effect of the integral scale of turbulence on one-dimensional spectra     

J. Neumann 《Meteorology and Atmospheric Physics》1961,12(3):389-395

Summary Assuming that in isotropic and stationary turbulence the Lagrangian correlation functions of the velocity components are of a simple exponential form, it is shown that the greater the integral scale of turbulence, the lower is the one-dimensional wave number to which a minus five-thirds law will appear to hold. It is pointed out, following a paper byGifford, that the apparent extension to low one-dimensional wave numbers of a minus-fivethirds law, does not necessarily mean thatKolmogoroff's inertial subrange of the three-dimensional turbulence extends to low (three-dimensional) wave numbers.

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Zusammenfassung Unter der Annahme, daß die Korrelationsfunktionen vonLagrange für isotrope und stationäre Turbulenz eine einfache exponentielle Form haben, wird bewiesen, daß, je größer das Integralmaß der Turbulenz ist, desto niedriger die eindimensionale Wellenzahl ist, bis zu der ein –5/3-Gesetz zu gelten scheint. Anschließend an eine Arbeit vonGifford wird betont, daß die scheinbare Erweiterung in Richtung auf niedrige eindimensionale Wellenzahlen nicht notwendigerweise bedeutet, daßKolmogoroffs Inertialteilbereich der dreidimensionalen Turbulenz sich bis zu niedrigen (dreidimensionalen) Wellenzahlen erstreckt.

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Résumé En supposant que les fonctions de corrélation lagrangiennes des composantes de vitesse de la turbulence isotropique et stationnaire sont d'une forme exponentielle simple, il est demontré que l'échelle intégrale de turbulence est d'autant plus grande que le nombre de l'onde unidimensionelle est petit, nombre, auquel une loi de –5/3 apparaît convenir. On pourra remarquer que, suivant un ouvrage deGifford, l'étendue apparente des petits nombres de l'onde unidimensionelle, ne signifie pas nécessairement que le rang d'inertie deKolmogoroff à propos de la turbulence en trois dimensions s'étend aux petits nombres de l'onde.

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