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31.
Ancient microbialites reflect interactions between microbial communities and environmental conditions. However, evaluating the relative roles of microbial community processes and environmental influences on microbialite morphology and internal fabric in the rock record can be challenging. The Neoproterozoic Beck Spring Dolomite preserves diverse microbialites, and thus provides an opportunity to explore the factors that influenced microbialite development locally. Stromatolitic, thrombolitic and composite microbialites are abundant in subtidal to upper intertidal carbonates in the Beck Spring Dolomite. Thrombolitic and composite microbialites have not been recognized previously in this unit, but compose much of the newly defined thrombolitic member. Stratigraphic relationships demonstrate that these three types of microbialites formed in close spatial and temporal association in subtidal to intertidal environments. The relative proportions and distributions of stromatolitic and thrombolitic microbialites vary with depositional environment; stromatolitic microbialites dominate in deeper intertidal to subtidal facies, whereas thrombolitic textures are more abundant in upper intertidal facies. Composite microbialites, composed of intermingled clotted and laminated textures, formed in all environments but are most abundant in intertidal facies. The broad environmental distribution of stromatolitic, thrombolitic and composite microbialites and the intermingling of textures suggest that laminated and clotted textures reflect diverse microbial community morphologies rather than environmental variations. Furthermore, the ca 750 Ma age of thrombolitic microbialites in the Beck Spring Dolomite requires that they formed without the influence of calcimicrobes or metazoans colonizing and grazing the microbial mat surface. Thus, these thrombolites provide further evidence that the biostratigraphic distribution of thrombolites cannot be uniquely attributed to evolution of calcifying and grazing organisms in the earliest Cambrian, and that older microbial communities were capable of producing clotted textures. 相似文献
32.
由于在川西北地区二叠纪-三叠纪界线地层的白云岩中一直没有找到可靠的生物化石,长期以来,白云岩就成为划分该地区二叠纪-三叠纪界线的标志。严格来说,这是一条岩石地层界线,而不是真正意义上的生物地层界线。在对四川江油渔洞子剖面进行详细的生物地层和沉积相研究时,在二叠纪长兴组珊瑚层以上的灰质白云岩中发现了微生物岩层并在其中首次找到了牙形刺H.parvus和灾难种有孔虫化石,从而为我们精确划分川北地区二叠纪-三叠纪的界线生物地层及确定该地区白云岩与微生物岩的时代及生态环境变化提供了可靠的生物化石依据。沉积特征的分析有利于了解从晚二叠世长兴组顶部至早三叠世飞仙关组中下部的沉积历史。 相似文献
33.
Bioturbation has long been considered an antagonist of microbialite development and preservation, because metazoan grazing and burrowing destroy benthic microbial communities. However, metazoan bioturbation, in conjunction with microbial accretion, may have had a significant role in the morphogenesis of some columnar microbialites, as suggested by the case study presented and by some Phanerozoic and Upper Proterozoic analogues discussed here. Late Miocene in age, the studied microbial biostrome developed in a western Mediterranean restricted shallow-water platform dominated by grainy sediments and with a notable influence of bioturbation. This study is focused on the complex accretionary history of the columnar microbialite biostrome and on its striking dark grey colour, which is attributed to Mn-oxyhydroxides precipitated during meteoric diagenesis linked to subaerial exposure. The characteristic columnar structure of the microbialite biostrome has features consistent with an accretionary origin of the columns, but also has features suggesting metazoan disruption. Therefore, a new morphogenetic model for columnar microbialites is presented, highlighting the concomitant roles of microbial accretion, bioturbation and grainy sediment infill of the intercolumn space. Whether this model is an exception or a rule, should be tested on other examples of Phanerozoic and Upper Proterozoic columnar microbialites. Nevertheless, this model is a step forward in understanding the complex microbe–metazoan interactions as constructive coexistence rather than just as destructive competition. 相似文献
34.
Difference of mound-bank complex reservoir under different palaeogeographic environment of the Sinian Dengying Formation in Sichuan Basin 
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下载免费PDF全文 Xu Zhe-Hang Lan Cai-Jun Hao Fang Chen Hao-Ru Yang Wei-Qiang Ma Xiao-Lin Zou Hua-Yao 《古地理学报》1999,22(2):235-250
The mound-bank deposits of the Dengying Formation of the Sinian System are well developed in Sichuan Basin,with good reservoir value. Based on outcrop and core observation,thin section observation,porosity and permeability of samples test,we analyzed the reservoir quality of different mound-bank complexes. During the terminal of Sinian Period,development of the Mianyang-Changning rift affected distribution of sedimentary facies of the 2nd and the 4th Members of Dengying Formation,forming different palaeogeographic units. The high-energy environment,transition environment and low-energy environment of the mound-bank complex rock types are identified,and their reservoir quality is different. The keep-up platform margin mound-bank complex,which is represented by Hujiaba section in the 4th Member of the Dengying Formation mainly deposited high-energy zone rock types. The catch-up platform margin mound-bank complex,which is represented by Yangba section in the 2nd Member of the Dengying Formation,mainly deposited transform zone rock types. The give-up platform interior mound-bank complex,which is represented by Fucheng section in the 4th Member of the Dengying Formation mainly deposited low-energy zone rock types. By comparing reservoir characteristics of three types of mound-bank complex,it is found that the keep-up platform margin mound-bank complex is the‘Sweet spots’ of the Dengying Formation in the Sichuan Basin. The above studies are conducive to find the‘Sweet spots’ of the Sinian Dengying Formation in the Sichuan Basin. 相似文献
35.
Microbialites (benthic microbial carbonate deposits) were discovered in a hypersaline alkaline lake on Eleuthera Island (Bahamas). From the edge towards the centre of the lake, four main zones of precipitation could be distinguished: (1) millimetre‐sized clumps of Mg‐calcite on a thin microbial mat; (2) thicker and continuous carbonate crusts with columnar morphologies; (3) isolated patches of carbonate crust separated by a dark non‐calcified gelatinous mat; and (4) a dark microbial mat without precipitation. In thin section, the precipitate displayed a micropeloidal structure characterized by micritic micropeloids (strong autofluorescence) surrounded by microspar and spar cement (no fluorescence). Observations using scanning electron microscopy (SEM) equipped with a cryotransfer system indicate that micrite nucleation is initiated within a polymer biofilm that embeds microbial communities. These extracellular polymeric substances (EPS) are progressively replaced with high‐Mg calcite. Discontinuous EPS calcification generates a micropeloidal structure of the micrite, possibly resulting from the presence of clusters of coccoid or remnants of filamentous bacteria. At high magnification, the microstructure of the initial precipitate consists of 200–500 nm spheres. No precipitation is observed in or on the sheaths of cyanobacteria, and only a negligible amount of precipitation is directly associated with the well‐organized and active filamentous cyanobacteria (in deeper layers of the mat), indicating that carbonate precipitation is not associated with CO2 uptake during photosynthesis. Instead, the precipitation occurs at the uppermost layer of the mat, which is composed of EPS, empty filamentous bacteria and coccoids (Gloeocapsa spp.). Two‐dimensional mapping of sulphate reduction shows high activity in close association with the carbonate precipitate at the top of the microbial mat. In combination, these findings suggest that net precipitation of calcium carbonate results from a temporal and spatial decoupling of the various microbial metabolic processes responsible for CaCO3 precipitation and dissolution. Theoretically, partial degradation of EPS by aerobic heterotrophs or UV fuels sulphate‐reducing activity, which increases alkalinity in microdomains, inducing CaCO3 precipitation. This degradation could also be responsible for EPS decarboxylation, which eliminates Ca2+‐binding capacity of the EPS and releases Ca2+ ions that were originally bound by carboxyl groups. At the end of these processes, the EPS biofilm is calcified and exhibits a micritic micropeloidal structure. The EPS‐free precipitate subsequently serves as a substrate for physico‐chemical precipitation of spar cement from the alkaline water of the lake. The micropeloidal structure has an intimate mixture of micrite and microspar comparable to microstructures of some fossil microbialites. 相似文献
36.
The Neoarchean Carawine Formation, Hamersley Group, Western Australia is a carbonate ramp that preserves diverse microbial structures which are characteristic of specific depositional environments. These distinctive structures are distributed in five shallow subtidal and two deeper‐water facies in the Oakover area of the Carawine Formation. The shallow subtidal facies are composed of biohermal and bedded stromatolites, centimetre‐scale ridge‐shaped microbialites and wavy‐laminated microbialites. The deeper‐water facies are composed of fenestrate microbialites, planar laminated dolostone and dolostone with rolled‐up microbial laminae. Microbialites in the Carawine Formation lie within a continuum of Archean to Proterozoic microbial facies. Some shallow‐water microbial facies in the Carawine Formation are similar to Proterozoic facies, such as large bioherms internally composed of a variety of stromatolite morphologies. In contrast, fenestrate microbialites grew in quiet subtidal environments and are common in Archean rocks but have not been documented in similar Proterozoic environments. The similarity of shallow‐water facies across the Archean–Proterozoic transition, before and after the oxidation of the atmosphere and surface oceans, indicates that stromatolite growth in shallow subtidal environments was not strongly affected by the chemical changes associated with oxidation of the oceans or by biological responses to those chemical changes. Rather, stromatolite morphology was controlled mostly by the physical environment and the corresponding biological responses to that environment. In contrast, the absence of fenestrate microbialites from Proterozoic deep subtidal environments suggests that the morphology of deep subtidal microbial structures was influenced by chemical or biological changes that occurred in association with oxidation of the surface oceans. 相似文献
37.
MARÍA E. SANZ-MONTERO† JUAN P. RODRÍGUEZ-ARANDA† MARÍA A. GARCÍA DEL CURA†‡ 《Sedimentology》2008,55(4):729-750
This research provides an ancient analogue for biologically mediated dolomite precipitation in microbial mats and biofilms, and describes the involvement of highly structured extracellular polymeric secretion (EPS) templates in dolomite nucleation. The structure of EPS is shown to match the hexagonal–trigonal lattice geometry of dolomite, which favoured the epitaxial crystallization of dolomite on the organic substrate. This structure of EPS also matches the arrangement of silica nanospheres in opal, which further accounts for the organically‐templated formation of opal enabling the non‐replacive co‐existence of dolomite and silica. The study is focused on a 50 m thick dolomite succession that is exposed in central areas of the Tertiary Duero Basin and was deposited in a mudflat‐saline lake sedimentary complex during the Middle to Late Miocene (9 to 15 Ma). In the intermediate intervals of the succession, poorly indurated dolomite beds pass gradually into silica beds. On the basis of sedimentological, compositional, geochemical and petrographic data, silica and dolomite beds have been interpreted as mineralized microbial mats. The silica beds formed in marginal areas of the lake in response to intense evaporative concentrations; this resulted in the rapid and early precipitation of opal. Silicification accounted for the exceptional preservation of the microbial mat structure, including biofilms, filamentous and coccoid microbes, and EPS. Extracellular polymeric secretions have a layered structure, each layer being composed of fibres which are arranged in accordance with a reticular pattern, with frequent intersection angles at 120° and 60°. Therefore, the structure of EPS matches the lattice geometry of dolomite and the arrangement of silica nanospheres in opal. Additionally, EPS binds different elements, with preference to Si and Mg. The concurrence of suitable composition and surface lattice morphologies in the EPS favoured the crystallization of dolomite on the substrate. In some cases, dolomite nucleation took place epicellularly on coccoid micro‐organisms, which gave way to spheroid crystals. Organic surfaces enable the inorganic mineral precipitation by lowering the free energy barrier to nucleation. Most of the microbial mats probably developed on the lake floor, under sub‐aqueous conditions, where the decomposition of organic matter took place. The subsequent formation of openly packed dolomite crystals, with inter‐related Si‐enriched fibrils throughout, is evidence for the pre‐existence of fibrillar structures in the mats. Miocene dolomite crystals are poorly ordered and non‐stoichiometric, with a slight Ca‐excess (up to 5%), which is indicative of the low diagenetic potential the microbial dolomite has towards a more ordered and stoichiometric structure; this confirms that microbial imprints can be preserved in the geological record, and validates their use as biosignatures. 相似文献
38.
华南二叠纪—三叠纪之交微生物岩的古地理背景及沉积微相特征 总被引：5，自引：1，他引：4
二叠纪末生物大灭绝后,在华南地区碳酸盐岩台地上普遍形成一套微生物岩。微生物岩所代表的是一种以蓝细菌和其他底栖生物为主体的独特的生态系。蓝细菌作为生产者是该生态系的基础,种类单调的小型腹足类和介形虫等广盐性多细胞生物和灾后残存的有孔虫则是该生态系中的主要消费者。根据所处的古地理位置不同,华南微生物岩的分布可分为3个主要地区,即上扬子地区、中下扬子地区及南盘江地区。上述3个地区微生物岩分别产出在生物礁顶、一般碳酸盐岩台地边缘和孤立碳酸盐岩台地环境,且不同地区微生物岩所在剖面的沉积序列也存在差异。根据不同地区微生物岩在宏观沉积构造和碳酸盐岩微相组合上的差异,可进一步划分为层纹状微生物岩、叠层石微生物岩、花斑状微生物岩、树枝状微生物岩及穹窿状微生物岩。不同微相类型的微生物岩在沉积水深、蓝细菌化石、底栖群落组合上仍存在差异,其中树枝状和穹窿状微生物岩沉积时所处水深最小,花斑状微生物岩和层纹状微生物岩沉积时所处水深可能相对较大。对不同地区微生物岩的研究为认识二叠纪末全球事件期间海洋环境的多样性和演化过程提供了重要的信息。 相似文献