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At hatching the larvae of flatfish closely resemble the bilateral symmetric larvae of other teleosts, especially perciforms. Literature data show that transformation to asymmetric benthic juveniles normally occurs at body lengths between 10 and 25 mm. Unexpectedly, minimal size at its completion (including eye migration) can be 4.1 mm SL and maximal size is over 72 mm. In this paper we consider the functional requirements for a successful switch from a symmetric pelagic larva to a typical asymmetric juvenile benthic flatfish partly based on evidence from other teleosts. The unfavourable period of eye migration and transition to a benthic habitat requires some food reserves and rewiring and/or recalibration of vision and gravity-associated structures utilised previously by the still symmetric larvae for e.g. food detection. Binocular fixation of the prey probably occurs in that stage. Critical or sensitive periods occurring during development of fish larvae suggest that a completely functional symmetric stage of development must precede transformation. The normal size range in flatfish larvae at transformation seems to confirm our considerations. Recent data on temperature effects during development provide an explanation for metamorphosis at the minimal size. Some evidence for paedomorphic heterochrony in flatfish larvae is presented. 相似文献
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This paper presents a potential based boundary element method for solving a nonlinear free surface flow problem for a ship moving with a uniform speed in finite depth of water. The free surface boundary condition is linearized by the systematic method of perturbation in terms of a small parameter up to third order. The surfaces are discretized into flat quadrilateral elements and the influence coefficients are calculated by Morino's analytical formula. Dawson's upstream finite difference operator is used in order to satisfy the radiation condition. The second order solution gives better result than the first or third order solution. So the present method with the second order solution can be adopted as a powerful tool for the hydrodynamic analysis of the thin ship in finite depth of water. 相似文献
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The steady response of the ventilated thermocline to an increase in Ekman pumping is investigated, focusing on the effect
of the mixed layer depth distribution on the subsurface density anomaly. We consider only the subtropical gyre, and the mixed
layer is assumed to be deep in the northwest and shallow elsewhere with a narrow transition zone separating the deep and shallow
mixed layer regions. At the intersection of this narrow transition zone and the outcrop line, low potential vorticity fluid
is subducted into and ventilates the thermocline. In such a situation, an enhancement of the Ekman pumping confined to the
northern subtropical gyre leads to pronounced subsurface cold anomalies in the southern subtropics, which is free of anomalous
forcing. These density anomalies are much greater than those that occur when either the mixed layer depth is zonally uniform
or the Ekman pumping is enhanced in the whole subtropical gyre. They are caused by anomalous changes in the trajectory of
the low potential vorticity fluid in response to anomalous Sverdrup flow.
This revised version was published online in July 2006 with corrections to the Cover Date. 相似文献
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- The spectral form of wind waves is investigated based on the ocean wave data observed at three nearshore stations of Taiwan. In this study, the generalized forms of Pierson-Moskowitz spectrum and JONSWAP spectrum are used to describe the local wave spectrum by selecting suitable spectral form parameters. It is shown that, at a specific site, the similarity of wave spectral form exists. Thus it is possible to use a representative spectral form for a given nearshore region to describe the wave spectrum at this nearshore. On the other hand, the effects of relative water depth on spectral form are examined. The feasibility of two spectral models in finite water depth is evaluated by using the same field wave data. 相似文献
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山东三山岛金矿床流体包裹体特征及其地质意义 总被引：5，自引：0，他引：5
三山岛金矿床位于莱州市三山岛—仓上断裂带内,矿石主要为蚀变岩和含金石英脉2种类型。随着胶东金矿床研究的深入和三山岛金矿资源的不断减少,对三山岛金矿床的成矿作用及深部开发前景问题研究具有更重要的意义。针对上述问题,对三山岛金矿床2种类型矿石的成矿流体特征进行对比研究,认为代表成矿早—中期的蚀变岩型矿石形成于中温(均一温度为325～240℃)、低盐度(2.07%～6.88%)、低密度(0.720～0.868g/cm3)、酸碱性不均匀(pH=3.27～10.43)、以氟化物和氯化物为成矿物质载体的还原性流体;代表成矿中—晚期的含金石英脉型矿石形成于中—低温(均一温度为306～160℃)、低盐度(1.05%～9.73%)、低密度(0.739～0.962g/cm3)、碱性(pH=9.25～9.85)、以氯化物和硫化物为成矿物质载体的还原性流体。成矿流体性质的转变反映了成矿流体处于由比较封闭到比较开放的构造环境的转变期,且成矿流体早期以原生岩浆水为主,后期有变质水、大气降水以及海水的参与作用。三山岛金矿成矿深度为2.5～5km,根据目前的开采深度推算其深部还有一定的资源量可供开发。 相似文献