Modelling phytoplankton succession on the Bering Sea shelf: role of climate influences and trophic interactions in generating Emiliania huxleyi blooms 1997–2000     

Agostino Merico  Toby Tyrrell  Evelyn J. Lessard  Temel Oguz  Phyllis J. Stabeno  Stephan I. Zeeman  Terry E. Whitledge 《Deep Sea Research Part I: Oceanographic Research Papers》2004,51(12):1929

Several years of continuous physical and biological anomalies have been affecting the Bering Sea shelf ecosystem starting from 1997. Such anomalies reached their peak in a striking visual phenomenon: the first appearance in the area of bright waters caused by massive blooms of the coccolithophore Emiliania huxleyi (E. huxleyi). This study is intended to provide an insight into the mechanisms of phytoplankton succession in the south-eastern part of the shelf during such years and addresses the causes of E. huxleyi success by means of a 2-layer ecosystem model, field data and satellite-derived information. A number of potential hypotheses are delineated based on observations conducted in the area and on previous knowledge of E. huxleyi general ecology. Some of these hypotheses are then considered as causative factors and explored with the model. The unusual climatic conditions of 1997 resulted most notably in a particularly shallow mixed layer depth and high sea surface temperature (about 4 °C above climatological mean). Despite the fact that the model could not reproduce for E. huxleyi a clear non-bloom to bloom transition (pre- vs. post-1997), several tests suggest that this species was favoured by the shallow mixed layer depth in conjunction with a lack of photoinhibition. A top-down control by microzooplankton selectively grazing phytoplankton other than E. huxleyi appears to be responsible for the long persistence of the blooms. Interestingly, observations reveal that the high N:P ratio hypothesis, regarded as crucial in the formation of blooms of this species in previous studies, does not hold on the Bering Sea shelf.  相似文献   

Respiratory Rate and Assessment of Secondary Production in the Holothuroid Holothuria tubulosa (Echinodermata) from Mediterranean Seagrass Beds     

Pierre  Coulon  Michel  Jangoux Patrick  Bulteel 《Marine Ecology》1992,13(1):63-68

Abstract. Respiration in Holothuria tubulosa was investigated in individuals from the Posidonia oceanica meadow off Lacco Ameno (Ischia Island, Italy). Respiratory rates increase with increasing body weight and increasing sea water temperature. Oxygen consumption of an average individual (7g dw body wall) ranges from 0.409 (14 °C) to 1.300 (26 °C) mg O₂· h^-1. Data on population density, mean size of individuals, and annual sea water temperature variations allow an assessment of holothuroid production. Values of 45.65 and 13.75 kJ · m^-2· y^-1 were calculated for shallow (3 to 10 m) and deep (25 to 33 m) areas of the Posidonia meadow, respectively. Holothuroid production shows a bathymetric pattern similar to primary production of the Posidonia -epiphytes complex and the production of Posidonia litter.  相似文献   

Population Structure and Secondary Production of the Mediterranean Octocoral Lophogorgia ceratophyta (L. 1758)     

M. Mistri 《Marine Ecology》1995,16(3):181-188

Abstract. The gorgonian Lophogorgia ceratophyta thrives in turbid water at Tinetto Rock, La Spezia Gulf, Ligurian Sea. Its age structure suggests that the population is in a steady state; nevertheless, it is difficult to discern whether the main factors controlling this population are biological or event-related. Secondary production was estimated by means of an increment-summation method and yielded 0.54 g ash-free dry weight (AFDW)*m^-1 a^-1. This estimate does not take into account biomass losses due to reproductive output and non-lethal predation. Standing stock biomass was 5.44 g AFDW e m^-2. The annual P/B ratio was 0.10 and turnover time about 10a  相似文献   

胶州湾女姑山水域夏季赤潮高发期浮游植物群落结构特征   总被引：7，自引：0，他引：7  

张利永  刘东艳  孙军  邹景忠 《中国海洋大学学报(自然科学版)》2004,34(6):997-1002

根据 1998年 5～ 8月夏季赤潮高发期间 ,胶州湾女姑山水域浮游植物群落结构的种群组成特征和演替变化对赤潮的发生和种群变动特点进行初步研究。结果表明 ,夏季女姑山水域的浮游植物主要由硅、甲藻组成 ,硅藻在细胞和物种丰度上都占优势。浮游植物优势种在调查期间出现明显的演替现象 ,水温的升高和营养条件的变化是引起物种演替的主要原因。浮游植物细胞丰度在夏季呈明显的增长趋势 ,从 5月份 6.3× 10 5个cell/m3 增至 8月份的 6.7× 10 6个cell/m3 ;其中 ,由于赤潮的爆发 ,细胞丰度的最高峰出现在 7月份 ,为 1.5× 10 7个 /m3 。在赤潮爆发期间 ,赤潮原因种高齿状藻和中肋骨条藻发生了明显的演替现象 ,不同细胞大小的优势种和营养条件的变化是发生物种演替的主要原因。叶绿素a的浓度和细胞丰度的变化共同反映了赤潮发生的全部过程。  相似文献   

Theoretical aspects of cap-rock and fault seals for single- and two-phase hydrocarbon columns   总被引：6，自引：0，他引：6  

N.L. Watts   《Marine and Petroleum Geology》1987,4(4)

Cap-rock seals can be divided genetically into those that fail by capillary leakage (membrane seals) and those whose capillary entry pressures are so high that seal failure preferentially occurs by fracturing and/or wedging open of faults (hydraulic seals). A given membrane seal can trap a larger oil column than gas column at shallow depths, but below a critical depth (interval), gas is more easily sealed than oil. This critical depth increases with lower API gravity, lower oil GOR and overpressured conditions (for the gas phase). These observations arise from a series of modelling studies of membrane sealing and can be conveniently represented using pressure/ depth (P/D) profiles through sealed hydrocarbon columns. P/D diagrams have been applied to the more complex situation of the membrane sealing of a gas cap underlain by an oil rim; at seal capacity, such a two-phase column will be always greater than if only oil or gas occurs below the seal.These conclusions contrast with those for hydraulic seals where the seal capacity to oil always exceeds that for gas. Moreover, a trapped two-phase column, at hydraulic seal capacity will be less than the maximum-allowed oil-only column, but more than the maximum gas-only column. Unlike membrane seals, hydraulic seal capacity should be directly related to cap-rock thickness, in addition to the magnitude of the minimum effective stress in the sealing layer and the degree of overpressure development in the sequence as a whole.Fault-related seals are effectively analogous to membrane cap-rocks which have been tilted to the angle of the fault plane. Consequently, all of the above conclusions derived for membrane cap-rocks apply to both sealing faults sensu stricto (fault plane itself seals) and juxtaposition faults (hydrocarbon trapped laterally against a juxtaposed sealing unit). The maximum-allowed two-phase column trapped by a sealing fault is greater than for equivalent oil-only and gas-only columns, but less than that predicted for a horizontal membrane cap-rock under similar conditions. Where a two-phase column is present on both sides of a sealing fault (which is at two-phase seal capacity), a deeper oil/water contact (OWC) in one fault block is associated with a deeper gas/oil contact (GOC) compared with the adjacent fault block. If the fault seal is discontinuous in the gas leg, however, the deeper OWC is accompanied by a shallower GOC, whereas a break in the fault seal in the oil leg results in a common OWC in both fault blocks, even though separate GOC's exist. Schematic P/D profiles are provided for each of the above situations from which a series of fundamental equations governing single- and two-phase cap-rock and fault seal capacities can be derived. These relationships may have significant implications for exploration prospect appraisal exercises where more meaningful estimates of differential seal capacities can be made.The membrane sealing theory developed herein assumes that all reservoirs and seals are water-wet and no hydrodynamic flow exists. The conclusions on membrane seal capacity place constraints on the migration efficiency of gas along low-permeabiligy paths at depth where fracturing, wedging open of faults and/or diffusion process may be more important. Contrary to previous assertions, it is speculated that leakage of hydrocarbons through membrane seals occurs in distinct pulses such that the seal is at or near the theoretically calculated seal capacity, once this has been initially attained.Finally, the developed seal theory and P/D profile concepts are applied to a series of development geological problems including the effects of differential depletion, and degree of aquifer support, on sealing fault leakage, and the evaluation of barriers to vertical cross-flow using RFT profiles through depleted reservoirs. It is shown that imbibition processes and dynamic effects related to active cross-flow across such barriers often preclude quantitative analysis and solution of these problems for which simulation studies are usually required.  相似文献   

Correspondence between the distribution of hydrodynamic time parameters and the distribution of biological and chemical variables in a semi-enclosed coral reef lagoon     

Jean-Pascal Torrton  Emma Rochelle-Newall  Aymeric Jouon  Vincent Faure  Sverine Jacquet  Pascal Douillet 《Estuarine, Coastal and Shelf Science》2007,74(4):766

Hydrodynamic modeling can be used to spatially characterize water renewal rates in coastal ecosystems. Using a hydrodynamic model implemented over the semi-enclosed Southwest coral lagoon of New Caledonia, a recent study computed the flushing lag as the minimum time required for a particle coming from outside the lagoon (open ocean) to reach a specific station [Jouon, A., Douillet, P., Ouillon, S., Fraunié, P., 2006. Calculations of hydrodynamic time parameters in a semi-opened coastal zone using a 3D hydrodynamic model. Continental Shelf Research 26, 1395–1415]. Local e-flushing time was calculated as the time requested to reach a local grid mesh concentration of 1/e from the precedent step. Here we present an attempt to connect physical forcing to biogeochemical functioning of this coastal ecosystem. An array of stations, located in the lagoonal channel as well as in several bays under anthropogenic influence, was sampled during three cruises. We then tested the statistical relationships between the distribution of flushing indices and those of biological and chemical variables. Among the variables tested, silicate, chlorophyll a and bacterial biomass production present the highest correlations with flushing indices. Correlations are higher with local e-flushing times than with flushing lags or the sum of these two indices. In the bays, these variables often deviate from the relationships determined in the main lagoon channel. In the three bays receiving significant riverine inputs, silicate is well above the regression line, whereas data from the bay receiving almost insignificant freshwater inputs generally fit the lagoon channel regressions. Moreover, in the three bays receiving important urban and industrial effluents, chlorophyll a and bacterial production of biomass generally display values exceeding the lagoon channel regression trends whereas in the bay under moderate anthropogenic influence values follow the regressions obtained in the lagoon channel. The South West lagoon of New Caledonia can hence be viewed as a coastal mesotrophic ecosystem that is flushed by oligotrophic oceanic waters which subsequently replace the lagoonal waters with water considerably impoverished in resources for microbial growth. This flushing was high enough during the periods of study to influence the distribution of phytoplankton biomass, bacterial production of biomass and silicate concentrations in the lagoon channel as well as in some of the bay areas.  相似文献   

渤海大型底栖动物次级生产力的初步研究   总被引：12，自引：2，他引：12  

于子山  张志南  韩洁 《中国海洋大学学报(自然科学版)》2001,31(6):867-871

1998年 9月和 1999年 4月分别对渤海 2 0个站进行了大型底栖动物两个航次的调查取样。整个研究海域大型底栖动物年次级生产力平均值为 6 .4 9g(AFDW) /(m2 · a) ,其中渤海海峡以东海域较高 ,为 12 .5 9g(AFDW) /(m2· a) ;渤海中部为 4 .4 6 g(AFDW) /(m2· a) ,渤海大型底栖动物年平均 P/B为 0 .82  相似文献   

Mechanism of Secondary Pore Formation and Prediction of Favorable Reservoir of Paleogene in Jiyang Sag,Eastern China     

Lü Xixue 《中国地质大学学报(英文版)》2008,19(6):675-684

Jiyang (济阳) sag is an oil rich basin,consisting of Huimin (惠民),Dongying (东营),Zhanhua (沾化),and Chezhen (车镇) depressions.The elastic rock of Paleogene has undergone early and middle diagenetic stages and now the main clastic reservoir is in the middle diagenetic stage.Primary and secondary pores are developed in Paleogene sandstone,the latter is generated from the dissolution of feldspar and calcite cement in rocks owing to the organic acid from the maturated source rock,but the materials dissolved are different in different depressions.The reservoir secondary pores of Dongying depression are generated from the dissolution of calcite cement,the ones of Zhanhua and Huimin depressions from the dissolution of feldspar,the secondary pores of Chezhen depression from the dissolution of feldspar in upper section,and the dissolution of calcite cement in the lower section of Paleogene,respectively.The secondary pores are developed in two depths and the depth goes down from west to east,from south to north in Jiyang sag.The major controlling factors for secondary pore development are maturity and location of source rock.Lastly,the favorable reservoirs are evaluated according to reservoir buried depth,sedimentation,and diagenesis.The reservoir with high quality is located in the northern and central parts in Dongying depression; there are some good reservoirs in Gndao (孤岛),Gudong (孤东),and Gunan (孤南) areas in Zhanhua depression,and the favorable reservoirs are located in the north steep slope and the south gentle slope of Chezhen depression and central uplift,south gentle slope of Huimin depression.  相似文献   

井中激电地-井方式井旁球体正反演   总被引：2，自引：0，他引：2  

周峰  潘和平  吴国平  徐健  陈滢 《物探与化探》2008,32(3):321-325

以激发极化法中的地-井方式中的体极化球形体作为研究对象,采用解析法对地下半空间的电位场进行求解,利用提取出的井中二次场异常电位差进行反演,得出极化球体的球心离井的距离和球心的埋深。通过计算机进行正反演模拟,验证了进行正反演计算的解析式的正确性,并从物理意义上对正反演过程中出现的现象进行分析。最后给出该方法的适用条件以及注意事项。研究对激发极化法地下勘探目标的定量分析具有明确意义。  相似文献   

天津市地下水开采对地面沉降影响的多元回归分析   总被引：1，自引：0，他引：1  

马锋  杨发俊  陈润桥  李亚平 《中国地质灾害与防治学报》2008,19(2):63-66

地下水的过量开采是天津市引起地面沉降的主要原因。因此天津市提出了“压缩地下水开采量”、“地下水人工回灌”、“调整地下水开采层次”等控制地面沉降的3大技术措施。经过多年的努力,控制地面沉降效果明显。如何解决地下水开发与控制地面沉降的关系,更好的贯彻这3大技术措施,是该文编写的初衷。即在开采同样地下水量的情况下,如何使地面沉降量最小;或在地面沉降量容许的情况下,如何开采最大量的地下水。压缩地下水开采量是治理地面沉降的根本措施,亦即如何压缩采水量或调整开采层次会达到最好效果。论文对天津市某区各个地下水开采层的多年累计开采量、累计沉降量进行数据统计分析,建立了该地区累计沉降量及各个地下水开采层的多元相关方程。在此基础上,分析了各个地下水开采层对地面沉降影响的相关程度。以此为该区控制地面沉降的提供依据。  相似文献