Extinctions, Morphological Gaps, Major Transitions, Stem Groups, and the Origin of Major Clades, with a Focus on Early Animals     

XIAO Shuhai 《《地质学报》英文版》2022,96(6):1821-1829

Systematic extinctions can leave major morphological gaps between living crown-group clades. Such morphological gaps would be perceived, from a neontological point of view, as major evolutionary transitions. In order to fill these morphological gaps and to map the evolutionary steps toward major evolutionary transitions, we need to integrate extinct stem-group taxa in phylogenetic studies. However, the recognition of stem group has not been widely adopted in the study of early animal fossils, despite that all fossils are stem groups at one level or another. Part of the difficulty is that stem groups may not have all features that collectively diagnose the respective crown group, and they can have unique (autapomorphic) features, making them tantalizingly similar to and frustratingly different from the crown group (e.g., stem-group eukaryotes can be prokaryotic and stem-group animals can be protistan). The need to embrace stem groups and to implement the PhyloCode, in order to achieve phylogenetic clarity and to offer key paleontological insights into the origin and early animal evolution, is illustrated in debates on several controversial Ediacaran and Cambrian fossils.  相似文献   

四川省大地构造单元划分及其基本特征          下载免费PDF全文

罗改  王全伟  秦宇龙  曾宜君  李云泉  杨学俊  李有波 《沉积与特提斯地质》2021,41(4):633-647

四川省大地构造演化研究已有百余年的历史,不同时期、不同学派在不同的尺度上提出了不同的大地构造单元划分方案,其中仍存在诸多争议。本文结合近30年来四川省特别是造山带地区开展的蛇绿岩和洋板块地层等调查和研究成果,以李廷栋所提出的洋板块地质学理念为指导思想,以区域主构造事件形成的优势大地构造相的时空结构组合和存在状态为基本原则,划分出秦-祁-昆造山系、勉县-略阳对接带、北羌塘-三江造山系以及扬子克拉通4个一级构造单元,包括11个二级构造单元和24个三级构造单元。  相似文献   

Cascades of sub-decadal,channel-floodplain changes in low-gradient,non-vegetated reaches near a dryland river terminus: Salar de Uyuni,Bolivia     

Jiaguang Li  Stephen Tooth  Guangqing Yao 《地球表面变化过程与地形》2019,44(2):490-506

The terminus of the ephemeral Río Colorado is located at the margins of Salar de Uyuni, Bolivia, the world's largest salt lake. The low-gradient (<0.0006 m m^-1), non-vegetated reaches approaching the terminus provide an excellent natural laboratory for investigating cascades of channel-floodplain changes that occur in response to quasi-regular flows (at least once annually) and fine-grained sediment supply (dominantly silt and clay). High-resolution satellite imagery (<0.65 m, various dates from 2004 onwards) and field data reveal widespread, pronounced and rapid morphodynamics on sub-decadal timescales, including channel erosion and chute cutoff formation, and development of crevasse channels and splays, floodouts (unchannelled surfaces at channel termini), and erosion cells (floodplain scour-transport-fill features). In particular, following high annual precipitation (>400 mm) in 2004–2005 and two subsequent high magnitude daily precipitation events (~40 mm), all of which led to widespread flooding, numerous crevasse splays formed between 2004 and 2016, avulsions occurred at nearby floodouts, and erosion cells downstream of the splays and floodouts underwent striking morphological changes. High-precision GPS data reveal two preferential localities for erosion cell development: partially or fully abandoned channels with crevasse splay remnants, and topographic lows between channels. In this overall low-gradient setting, comparatively high gradients (up to ~0.0006 m m^-1) at the edge of splay deposits and topography created by crevasses and abandoned channels may initiate knickpoint retreat and thereafter erosion cell development. Abandoned channels with splays tend to give rise to narrow, deep erosion cells, while topographic lows promote relatively shallow, wide erosion cells. In both situations, erosion cells may extend upslope and downslope, and eventually connect to form straight channels. The channel-floodplain morphodynamics near the Río Colorado terminus extend previous analyses of low-gradient, dryland river systems, particularly because the lack of vegetation and quasi-regular floods drive cascades of rapid changes on sub-decadal timescales. © 2018 John Wiley & Sons, Ltd.  相似文献   

一种基于交叉剖面的地质模型构建方法     

陈国良  刘修国  盛谦  张勇慧 《岩土力学》2011,32(8):2409-2415

针对基于近似平行断面数据的建模方法的缺点,利用交叉地质剖面数据,提出了一种通过划分空间单元格的思路来构建地质结构模型的方法。该方法基于"分而自治"的思想,将复杂的整体的建模区域划分为单个单元格内的建模,待每个格子内的建模完成之后,采用相反的思路,将所有格子内的模型拼接起来形成最终的建模区域内的整体模型。  相似文献   

驼背鲈外周血细胞的形态学研究   总被引：1，自引：0，他引：1       下载免费PDF全文

王静香  李加儿  区又君 《海洋科学》2011,35(4):77-82

利用Wright's和Giemsa双重染色法和改良的血涂片染色法两种方式,通过光镜对驼背鲈(Cromileptes altivelis)的血涂片进行了观察.结果表明:驼背鲈的血涂片中可区分出6种细胞--红细胞、嗜中性粒细胞、嗜酸性粒细胞、单核细胞、淋巴细胞、血栓细胞,偶尔可见到分裂的红细胞、分裂的血栓细胞、核影红细胞和...  相似文献   

Strenulagus (Mammalia: Lagomorpha) from the Middle Eocene Irdin Manha Formation of the Erlian Basin, Nei Mongol, China     

?ucja FOSTOWICZ-FRELIK  LI Chuankui  LI Qian  MENG Jin  WANG Yuanqing 《《地质学报》英文版》2015,89(1):12-26

New material of Strenulagus solaris, a stem lagomorph, from the lower part of the Middle Eocene Irdin Manha Formation at the Irdin Manha and Huheboerhe localities, Erlian Basin, Nei Mongol, China includes remains of isolated cheek teeth, fragmentary upper incisors(d I2) and postcranial elements, reported from the species for the first time. The tarsal bones display advanced features, such as a wide calcaneofibular articulation surface and, along with an increase in hypsodonty, the new data document the early stage of lagomorph evolution in the Middle Eocene of the Erlian Basin. This finding is stratigraphically well constrained and probably predates that of the type material of S. solaris from the Khaychin Formation, Bugin-Tsav Basin, Mongolia.  相似文献   

繁茂膜海绵清除大肠杆菌的过程     

张喜昌  张卫  褚亚东  赵宗宝  谭海东  梁玉波  许淑芬  冷晓飞  李丹 《海洋科学》2015,39(5):21-27

通过分析海绵清除大肠杆菌的过程,研究海绵净化细菌的机理。作者利用荧光显微镜和激光共聚焦显微镜观测等手段,监测和分析了绿色荧光大肠杆菌(Escherichia coli)在繁茂膜海绵(Hymeniacidon perlevis)体内、体外水环境中数量变化过程。在1 L含有3×107个/m L绿色荧光大肠杆菌的海水中放入鲜重(1.02±0.11)g的繁茂膜海绵24块,处理7 h,海水中的荧光大肠杆菌数量逐渐降低;而海绵体内荧光大肠杆菌数量在2 h时内逐渐增多,之后的2 h趋于稳定,4 h以后开始逐渐减少。水体中大肠杆菌不仅进入海绵体内,而且进入海绵细胞内。含有荧光大肠杆菌的海绵块转入无菌海水中后,海绵体内及细胞中大肠杆菌逐渐消失,而且大肠杆菌没有被释放到环境海水中。分析表明,繁茂膜海绵能够以摄食的方式净化水环境中的大肠杆菌。  相似文献   

免疫细胞化学法测定鱼类淋巴囊肿病毒(LCDV)滴度     

宋佳蕾  绳秀珍  战文斌 《中国海洋大学学报(自然科学版)》2012,42(5):55-59

利用免疫细胞化学法测定鱼类淋巴囊肿病毒(LCDV)滴度.以牙鲆鳃细胞系(FG)作为感染细胞,将生长旺盛的FG细胞接种于48孔培养板中培养至形成细胞单层,用2倍连续稀释的LCDV粗提液分别接种FG细胞.固定各稀释度LCDV感染后的FG细胞,孵育抗牙鲆LCDV单克隆抗体,其后再运用生物素-亲合素反应系统,以碱性磷酸酶底物APRed试剂盒发色.倒置显微镜观察,被病毒感染的FG细胞的细胞质呈现红色,未被感染细胞的细胞质呈无色.记录各稀释度病毒感染的阳性细胞孔数,按Reed-Muench法计算组织细胞培养半数感染量(TCID0).结果显示,免疫细胞化学法测得LCDV在FG的滴度为1.77×210 TCID50/mL.该法可以用来有效测定LCDV滴度,且结果直观、准确性较好,灵敏度较高.  相似文献   

日本七鳃鳗外周血细胞显微结构及类淋巴细胞体外培养   总被引：1，自引：0，他引：1  

逄越  李庆伟 《海洋科学》2012,36(1):23-29

以日本七鳃鳗(Lampetra japonica)为研究对象,观察其外周血细胞显微结构和探讨类淋巴细胞原代培养条件。采用Wright氏和Giemsa氏染色法对日本七鳃鳗外周血液有形成分进行显微观察,可鉴别出红细胞、淋巴细胞、中性粒细胞、单核细胞、嗜酸性粒细胞;未发现嗜碱性粒细胞。通过Ficoll密度梯度离心法和流式细胞仪分选获得高纯度的七鳃鳗类淋巴细胞。经条件优化后确定最适培养液L15+20%FBS(5%七鳃鳗血清)+0.46%NaCl,在18℃,pH 6.8-7.0的条件下对日本七鳃鳗类淋巴细胞进行体外培养,大多数类淋巴细胞半贴壁生长,细胞状态较好,最长可以存活近一个月。类淋巴细胞的原代培养为日本七鳃鳗细胞体外培养的深入研究提供了实验依据。  相似文献   

长蛸的盐度耐受性及盐度胁迫对其血细胞和体内酶活力的影响   总被引：1，自引：0，他引：1  

张玉玉  王春琳  李来国 《台湾海峡》2010,29(4):452-459

通过控制暂养水体盐度,用盐度突变与盐度渐变2种方法测定长蛸的盐度耐受性.测定了长蛸血细胞密度、各类血细胞数量比例以及各种酶的活力几组数据,并分析其血细胞与体内酶活力的变化.结果表明：长蛸的生存盐度范围为7.0~30.3,适宜盐度范围为16.3~27.3,最适盐度范围为18.3~24.3,对盐度的适应范围较广,有利于长蛸大规模养殖的开展.在盐度胁迫下,长蛸血细胞密度,各种血细胞数量比例均发生显著改变,长蛸个体也随着胁迫加大变得越来越不适应,出现喷墨、休克、甚至死亡,这表明长蛸的免疫机能降低.无论在低盐度还是高盐度胁迫情况下,受渗透压的影响,长蛸肌体大量吸水和失水,呼吸作用减弱,体内供氧降低,导致长蛸体内有氧呼吸降低和无氧呼吸提高,于是催化有氧呼吸的LDH酶活力降低.同时,细胞内与免疫和消化相关的细胞器由于细胞吸水或失水作用功能受到影响,保护酶（SOD、POD、CAT）、磷酸酶（ACP、AKP）和消化酶（蛋白酶和脂肪酶）的活力均呈下降趋势.由此可见,盐度胁迫对长蛸的影响是显著的.  相似文献