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On Evolutionary Sequence Systematics: Taking Permian Waagenophylloid Coral Fauna as an Example
Authors:Xu Guirong  Gong Shuyun  Wang Yongbiao
Abstract:Evolutionary sequence systematics is a discipline in paleontology and biology that deals with phylogeny of organism, the main purpose of which is to understand relatives between species and taxa and to show their evolutionary sequence. The systematics is set up on the phenetic-cladistic systematics. A number of important theoretical concepts in the cladistics are added the concept of time in the systematics. For example, the time-span between the closest sister species is considered limited in the duration of their mother species. The foundational methodology of the systematics is to analyze sister-group and character mosaic distribution, and the key to study evolutionary sequence is to understand cbaracter mosaic distribution fully. These aualyses can be executed by using computer. In the paper, the Permian Waagenophylloid coral fauna is taken as an example to illustrate the analysis on evolutionary sequence. These coral fossils are widespread in Tethyan area.Seventeen characters and a number of cbaracter states for each character used in study result from features of Waagenophylloid corals. According to principles of tbe widest distribution of character states, fossil records and ontogenesis we discerned plesiomorph character states for each character. Characters are divided into 4 ranks on the basis of their important place in pkylogeny. Based on the theory and methopology of tbe evolutionary sequence systematics the paper analyzes the sister groups and evolutionary sequence. In the examination each other between the analyses of sister groups and evolutionary sequence tbe paper found many importunt phenomena of paleontology,such as character degeneration, parallel evolution, and polydirectional clades, and distinguished a number of abnormal order. Iu the discussion to test tke rirst occurrence event8 of biostratigraphy we infer the Possible stratigraphic levels of some fossils.
Keywords:evolutionary sequence systematics  character mosaic distribution  Permian Waagenophylloid corals
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