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The shark-beds of the Eyam Limestone Formation (Lower Carboniferous,Viséan) of Steeplehouse Quarry,Wirksworth, Derbyshire,UK
Institution:1. University of Portsmouth, United Kingdom;2. The Natural History Museum, London, United Kingdom;1. Department of Paleontology and Geology, Hungarian Natural History Museum, Budapest, Hungary;2. Department of Paleontology, Eötvös University, Budapest, Hungary;1. Department of Earth and Planetary Sciences, Graduate School of Sciences, Kyushu University, Fukuoka 812-8581, Japan;2. Department of Earth and Planetary Sciences, Faculty of Sciences, Kyushu University, Fukuoka 812-8581, Japan;3. Department of Earth Science, Graduate School of Science, Tohoku University, Sendai 980-8571, Japan;4. Department of Earth and Planetary Materials Science, Tohoku University, Sendai 980-8571, Japan;5. Photon Factory, High Energy Accelerator Research Organization, Tsukuba 305-0801, Japan;1. Department of Earth Sciences, Natural History Museum, London, UK;2. Faculty of Earth Sciences, University of Silesia, B?dzińska 60, 41-200 Sosnowiec, Poland;1. Institut für Geologie, Universität Hamburg, Bundesstr. 55, 20146 Hamburg, Germany;2. Kuban State University, Kazan Federal University, Aphipskij, Post box demand, 353235 Krasnodar, Russia;3. Département de géologie, université de Liège, bâtiment B18, Sart-Tilman, 4000 Liège, Belgium;4. Institut royal des sciences naturelles de Belgique, DO terre et histoire de la vie, 29, rue Vautier, 1000 Bruxelles, Belgium;1. Research Institute for Fundamental Sciences, University of Tabriz, Tabriz, Iran;2. Earth Sciences Department, University of Tabriz, Tabriz, Iran;3. Geologic Survey of Iran-NW branch, Tabriz, Iran
Abstract:The Eyam Limestone Formation of Steeplehouse Quarry, Wirksworth, Derbyshire, UK yields a diverse assemblage of Lower Carboniferous vertebrate remains. The assemblage is dominated by dermal denticles of the enigmatic selachian Petrodus patelliformis M’Coy, 1848, but also contains teeth of petalodonts, hybodonts and neoselachians. Actinopterygian remains also occur. The assemblage has yielded the earliest Neoselachian, Cooleyella fordi (Duffin and Ward, 1983) and the earliest British lonchidiid, Reesodus wirksworthensis (Duffin 1985). The first occurrence of the enigmatic spiny shark Acanthorhachis (Listracanthidae) is reported from the Viséan, extending its range back some 10 million years. Associated invertebrate remains and sedimentological data indicates a thriving fore-reef environment, deposited in a low energy off-reef setting. The vertebrate remains are well preserved with little abrasion, indicating short transport distances. Conodont elements indicating a late Brigantian age (Early Carboniferous, Viséan) have unusual and extensive euhedral apatite overgrowths.
Keywords:Early Carboniferous  Viséan  Vertebrates  Elasmobranchs  Conodonts  England
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