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Scydmobisetia gen. nov., the first definite Glandulariini from Upper Cretaceous Burmese amber (Coleoptera: Staphylinidae: Scydmaeninae)
Institution:1. Museum of Natural History, University of Wroc?aw, Sienkiewicza 21, 50-335 Wroc?aw, Poland;2. Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Hakozaki 6-10-1, Higashi-ku, Fukuoka 812-8581, Japan;3. Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tenoudai, Tsukuba, Ibaraki 305-0001, Japan;4. Japan Society for the Promotion of Science Research Fellow (DC), Japan;1. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;2. Department of Biology, Shanghai Normal University, Shanghai 200234, China;3. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China;4. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;1. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;1. Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai, 200234, China;2. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;1. Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605-2496, USA;2. The Kyushu University Museum, 6-10-1 Hakozaki, Fukuoka 812-8581, Japan;3. Canadian Food Inspection Agency, 960 Carling Ave., Ottawa, ON K1A 0C6, Canada;4. Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tenoudai, Tsukuba, Ibaraki 305-0001, Japan;1. Tokushima Prefectural Museum, Mukoterayama, Hachiman-cho, Tokushima 770-8070, Japan;2. Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, 6-10-1 Hakozaki, Fukuoka 812-8581, Japan;3. Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennoudai, Tsukuba City, Ibaraki 305-0001, Japan;1. Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Hakozaki 6-10-1, Higashi-ku, Fukuoka 812-8581, Japan;2. Japan Society for the Promotion of Science, Tokyo, Japan
Abstract:The Glandulariini (=Cyrtoscydmini) includes about 60 genera comprising over 70% of the extant species of Scydmaeninae. This successful group is today very common and abundant in forests of all continents. However, while extinct representatives of much smaller extant tribes, as Clidicini, Eutheiini and Cephenniini, were described from Cretaceous deposits, named genera and species of Glandulariini were so far known only from much younger Baltic, Sicilian and Dominican ambers. Here we report the first genus and species of definite Glandulariini from the Upper Cretaceous Burmese amber, Scydmobisetia vetutissima Ja?oszyński and Yamamoto, gen. and sp. nov. Scydmobisetia is the first known glandulariine beetle with a pair of very long vertexal sensilla chaetica, previously not found in this tribe, but common among unrelated Staphylinidae. The new genus shows similarities to the extant genera Anthicimimus, Sciacharis, Spinosciacharis and Horaeomorphus, but S. vetutissima has protibial modifications typical of males of some extant species of Euconnus. We discuss the apparent disproportion in the oldest fossil record of various tribes of scydmaenines as a result of identification problems, and not the lack of known fossils.
Keywords:Fossil  Beetle  Laurasia  Phylogeny  Evolution  Cenomanian  AMNH"}  {"#name":"keyword"  "$":{"id":"kwrd0045"}  "$$":[{"#name":"text"  "_":"American Museum of Natural History  New York  USA
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