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A new genus of whip-scorpions in Upper Cretaceous Burmese amber: Earliest fossil record of the extant subfamily Thelyphoninae (Arachnida: Thelyphonida: Thelyphonidae)
Institution:1. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;1. Department of Geosciences, University of Padova, Via G. Gradenigo 6, 35131, Padova, Italy;2. University of Vienna, Department of Paleontology, Geozentrum, Althanstrasse 14, 1090, Vienna, Austria;3. Soprintendenza per i Beni Archeologici del Friuli Venezia Giulia, Nucleo Operativo di Udine, Via Zanon, 22 I, 33100, Udine, Italy;1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China;2. University of Chinese Academy of Sciences, Beijing 100049, China;3. University of Science and Technology of China, 96 Jinzhai Road, Hefei 230026, China;4. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany;1. Department of Earth Sciences, Montana State University, Bozeman, MT, 59717, USA;2. Museum of the Rockies, Bozeman, MT, 59717, USA;1. Institute of Geology and Paleontology, Linyi University, Shuangling Road, Linyi, 276000, China;2. School of Agronomy and Environment, Weifang University of Science and Technology, Jinguang Road, Shouguang, 262700, China;3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, China;4. School of Geography and Tourism, Qufu Normal University, Rizhao, 276826, China;5. Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia;6. Natural History Museum, London, SW7 5BD, UK;1. Universidad Nacional de San Luis, CONICET, Departamento de Geología, Ejército de Los Andes 950, CP: 5700, San Luis, Argentina;2. Boston College, Charcas 3949, CP: 1425, Ciudad Autónoma de Buenos Aires, Argentina;3. Museo de La Plata, CONICET, División Paleontología Vertebrados, Paseo del Bosque S/n, B1900FWA, La Plata, Argentina;4. Biodiversity Institute, The University of Kansas, Lawrence, KS, USA;1. Museu Nacional/Universidade Federal do Rio de Janeiro, Departamento de Geologia e Paleontologia, 20940-040, Rio de Janeiro, RJ, Brazil;2. Museu de Zoologia da Universidade de São Paulo, Laboratório de Paleontologia, Avenida Nazaré, 48, Ipiranga, 04263-000, São Paulo, SP, Brazil
Abstract:Mesozoic whip-scorpions are very rare, with only two Cretaceous species known to date. Here we describe a new genus and species of Thelyphonidae, Mesothelyphonus parvus gen. & sp. nov., based on a very well-preserved male in Upper Cretaceous amber from Myanmar. Mesothelyphonus is firmly placed in the extant subfamily Thelyphoninae as supported by the abdominal tergites with a median longitudinal suture. Mesothelyphonus differs from other fossil and recent genera primarily by its very small body size, the absence of ommatoids on abdominal segment XII, and the elongate, slender and toothed patellar apophysis of the male pedipalp. The new discovery represents the oldest definitive fossil record for Thelyphoninae, highlighting the antiquity of the whip-scorpion group.
Keywords:Myanmar  Thelyphonida  Thelyphonidae  Thelyphoninae  Cretaceous
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