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Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana
Institution:1. State Key Laboratory of Lithospheric Evolution, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing 100029, China;2. Department of Earth and Environmental Sciences, Università di Milano-Bicocca, 20126 Milan, Italy;3. State Key Laboratory of Mineral Deposits Research, School of Earth Sciences and Engineering, Nanjing University, Nanjing 210023, China;4. School of Earth Science and Geological Engineering, Sun Yat-sen University, Guangzhou 510275, China;2. Geological Survey of Western Australia, Department of Mines and Petroleum, 100 Plain Street, East Perth, WA 6004, Australia;3. Earth and Planetary Sciences, Western Australian Museum, Locked Bag 49, Welshpool DC, WA 6986, Australia;4. Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, S-104 05 Stockholm, Sweden;5. Department of Applied Geology, Curtin University, Kent Street, Bentley, WA 6102, Australia;6. Palaeobiology Programme, Department of Earth Sciences, Uppsala University, Villavägen 16, SE-752 36 Uppsala, Sweden;7. Geoscience Australia, GPO Box 383, Canberra, ACT 2601, Australia;8. School of Life and Environmental Sciences, Deakin University, 221 Burwood Highway, Burwood, VIC 3125, Australia;1. The Institute for Geoscience Research (TIGeR), School of Earth and Planetary Sciences, Curtin University, GPO Box U1987, Perth, WA 6845, Australia;2. Centre for Exploration and Targeting (CET), Curtin University, GPO Box U1987, Perth, WA 6845, Australia;3. School of Molecular and Life Sciences, Curtin University, GPO Box U1987, Perth, WA 6845, Australia;4. John de Laeter Center, Curtin University, GPO Box U1987, Perth, WA 6845, Australia;5. Geological Survey of Western Australia, 100 Plain St., East Perth, WA 6004, Australia
Abstract:Beetles belonging to the extinct tribe Notocupedini (Coleoptera: Cupedidae: Ommatinae) are widespread in Mesozoic deposits of eastern Pangaea. Four new species are proposed in the genus/collective group Zygadenia: Z. liui sp. nov. (Aptian: Yixian Formation, China), Z. martinae sp. nov. (Sinemurian–Toarcian: Cattamarra Coal Measures, Australia), Z. floodpagei sp. nov. and Z. simpsoni sp. nov. (Hauterivian–Barremian: Weald Clay Group, UK). The Purbeck species Z. tuberculata and Z. angliae are figured from the Weald Clay Group for the first time. It is also proposed that Notocupes, the largest genus in this presumably xylophagous, warm climate tribe, be divided into two subgenera: Notocupes subgen. n. and Amblomma stat. nov. In addition, the palaeoecological significance of colour markings in these Mesozoic beetles is also discussed for the first time. These discoveries extend the geographic distribution/stratigraphic range of Zygadenia and shed new light on the palaeoecology and diversity of so-called 'primitive' beetles in pre-angiospermous woodlands.
Keywords:colour behaviour  new species
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