New discoveries of vertebrate remains from the Triassic of Riba de Santiuste,Guadalajara (Spain) |
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| Authors: | Mélani Berrocal-Casero Julia Audije-Gil Rui Alexandre Castanhinha Juan Alberto Pérez-Valera Vanda Faria dos Santos Manuel Segura |
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| Institution: | 1. Grupo de Investigación Procesos Bióticos Mesozoicos, Departamento de Paleontología, Universidad Complutense de Madrid, 28040 Madrid, Spain;2. Grupo de Investigación IberCreta, Departamento de Geología y Geografía, Universidad de Alcalá, 28871 Alcalá de Henares, Spain;3. Departamento de Biología, Laboratorio de Poblaciones del Pasado (LAPP), Universidad Autónoma de Madrid, 28049 Madrid, Spain;4. Department of Biology and CESAM, University of Aveiro, 3810-193 Aveiro, Portugal;5. Museu da Lourinhã ? GEAL, 2530-158 Lourinhã, Portugal;6. Departamento de Ciencias de la Tierra y Medio Ambiente, Universidad de Alicante, 03690 San Vicente del Raspeig, Spain;g. Departamento de Geologia, Faculdade de Ciências da Universidade de Lisboa, Edifício C6, Campo Grande, 1749-016 Lisboa, Portugal |
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| Abstract: | The palaeontological sites of Riba de Santiuste and Sienes (Riba de Santiuste area) are located in the province of Guadalajara, Spain. They include a stratigraphic interval in Muschelkalk facies belonging to the “Cuesta del Castillo Sandstones” Formation and “Royuela Dolostones, Marls and Mudstones” Formation. These sites include numerous fossil plants, direct vertebrate remains, and vertebrate swim traces. The vertebrate remains correspond to a multitude of anatomical elements of Sauropterygia (Nothosauroidea, Placodontia) and possible Archosauria (Rauisuchia) remains. The fossil material attributed to nothosaurs includes teeth, coracoids, a thoracic vertebra, some isolated vertebral centra, humerus, rib fragments, and some dorsal and caudal vertebrae. The remains attributed to placodonts correspond to fragments of skull, quadrate, teeth and osteoderms. Other undetermined sauropterygian remains, such as ulnas, fragments of long bones, fragments of ribs, and articular facets of ribs have been also recovered. Additionally, a fragment of mandible and an intervertebral disk of indeterminate reptiles whose size could be compatible with archosaurs are also described. These bones are exceptionally well-preserved because the fossilization processes have preserved the microstructure of the tissues. The sites also show vertebrate traces, with parallel scratch impressions interpreted as swim traces. The relative stratigraphic position and the palaeontological content of these sites suggest a Ladinian age (Middle Triassic). The interpretation of the sedimentary facies here described also suggests that the sites could correspond to detrital-carbonate mixed deposits of coastal intertidal to supratidal environments. |
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| Keywords: | Nothosaurs Placodonts Rauisuchia Swim traces Ladinian Spain |
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