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Investigation of Uncertainties of Establishment Schemes in Dynamic Global Vegetation Models
作者姓名:SONG Xiang  ZENG Xiaodong
基金项目:supported by the Strategic Priority Research Program of the Chinese Academy of Sciences(Grant No.XDA05110103);the State Key Project for Basic Research Program of China(Grant No.2010CB951801);the National High Technology Research and Development Program of China(863 Program)(Grant No.2009AA122105)
摘    要:In Dynamic Global Vegetation Models (DGVMs), the establishment of woody vegetation refers to flowering, fertiliza- tion, seed production, germination, and the growth of tree seedlings. It determines not only the population densities but also other important ecosystem structural variables. In current DGVMs, establishments of woody plant functional types (PFTs) are assumed to be either the same in the same grid cell, or largely stochastic. We investigated the uncertainties in the competition of establishment among coexisting woody PFTs from three aspects: the dependence of PFT establishments on vegetation states; background establishment; and relative establishment potentials of different PFTs. Sensitivity experi- ments showed that the dependence of establishment rate on the fractional coverage of a PFT favored the dominant PFT by increasing its share in establishment. While a small background establishment rate had little impact on equilibrium states of the ecosystem, it did change the timescale required for the establishment of alien species in pre-existing forest due to their disadvantage in seed competition during the early stage of invasion. Meanwhile, establishment purely fiom background (the scheme commonly used in current DGVMs) led to inconsistent behavior in response to the change in PFT specification (e.g., number of PFTs and their specification). Furthermore, the results also indicated that trade-off between irtdividual growth and reproduction/colonization has significant influences on the competition of establishment. Hence, further development of es- tablishment parameterization in DGVMs is essential in reducing the uncertainties in simulations of both ecosystem structures and successions.

关 键 词:植被模型  不确定性  植物功能型  生态系统  功能检查  种子生产  结构变量  平衡状态

Investigation of uncertainties of establishment schemes in dynamic global vegetation models
SONG Xiang,ZENG Xiaodong.Investigation of uncertainties of establishment schemes in dynamic global vegetation models[J].Advances in Atmospheric Sciences,2014,31(1):85-94.
Authors:Xiang Song  Xiaodong Zeng
Institution:1. International Center for Climate and Environment Sciences, Institute of Atmospheric Physics, Chinese Academy of Sciences, Beijing, 100029, China
Abstract:In Dynamic Global Vegetation Models (DGVMs), the establishment of woody vegetation refers to flowering, fertilization, seed production, germination, and the growth of tree seedlings. It determines not only the population densities but also other important ecosystem structural variables. In current DGVMs, establishments of woody plant functional types (PFTs) are assumed to be either the same in the same grid cell, or largely stochastic. We investigated the uncertainties in the competition of establishment among coexisting woody PFTs from three aspects: the dependence of PFT establishments on vegetation states; background establishment; and relative establishment potentials of different PFTs. Sensitivity experiments showed that the dependence of establishment rate on the fractional coverage of a PFT favored the dominant PFT by increasing its share in establishment. While a small background establishment rate had little impact on equilibrium states of the ecosystem, it did change the timescale required for the establishment of alien species in pre-existing forest due to their disadvantage in seed competition during the early stage of invasion. Meanwhile, establishment purely from background (the scheme commonly used in current DGVMs) led to inconsistent behavior in response to the change in PFT specification (e.g., number of PFTs and their specification). Furthermore, the results also indicated that trade-off between individual growth and reproduction/colonization has significant influences on the competition of establishment. Hence, further development of establishment parameterization in DGVMs is essential in reducing the uncertainties in simulations of both ecosystem structures and successions.
Keywords:Dynamic Global Vegetation Model  uncertainty  establishment scheme  PFT classification  fractional coverage
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