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A new ornithomimid dinosaur from the Upper Cretaceous Packard Shale formation (Cabullona Group) Sonora,México
Institution:1. Royal Ontario Museum, 100 Queen''s Park, Toronto, Ontario M5S 2C6, Canada;2. Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks Street, Toronto, Ontario M5S 3B2, Canada;3. Department of Biosphere-Geosphere Science, Okayama University of Science, 1-1 Ridai-cho, Kita-ku, Okayama-shi, Okayama 700-0005, Japan;4. School of International Liberal Studies, Waseda University, 1-6-1 Nishiwaseda, Shinjuku-ku, Tokyo 169–8050, Japan;5. Department of Vertebrate Paleontology, Cleveland Museum of Natural History, 1 Wade Oval Drive, Cleveland, OH 44106, USA;6. Department of Earth Sciences, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada;7. School of Earth and Environmental Sciences, Seoul National University, 1 Gwanak-ro, Gwanak-gu, Seoul 08826, South Korea;8. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada;9. Institute of Paleontology and Geology, Mongolian Academy of Sciences, S.Danzan Street 3/1, Ulaanbaatar 15160, Mongolia
Abstract:New ornithomimid material discovered from the Upper Cretaceous Packard Shale Formation, (Cabullona Group) of Sonora, Mexico is described. The material includes a partial skeleton, which is assigned to a new genus and species, Tototlmimus packardensis. This new taxon differs from other ornithomimids in having five unique characteristics that separate it from other Northamerican ornithomimids: (1) a distinctively articulation between metatarsals, where the distal ends of metatarsals II and IV contact directly with the distal facet of metatarsal III; (2) a metatarsal III with a weakly ginglymoid distal articular face; (3) the medial and lateral sides of metatarsal III are shaped into the form of metatarsals II and IV, so all distal ends fit together when they articulate; (4) an asymmetrical and narrow pedal ungual with shallow grooves in both medial and lateral sides; and finally, (5) the presence of a deep sulcus on the ventromedial edge, close to the articular end. Phylogenetic analysis suggests that Tototlmimus packardensis corresponds to a derived ornithomimid included in the Northamerican clade, forming a monophyly with Ornithomimus. Tototlmimus packardensis is the first definitive ornithomimid described for Mexico, and represents one of the southernmost occurrences in the Western Interior Basin of North America.
Keywords:Dinosaurs  Theropods  Ornithomimosauria  Ornithomimidae  Packard Shale Formation  Late Cretaceous  Sonora  Mexico
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