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New gadiform fishes (Teleostei,Gadiformes) from the Miocene of Algeria
Institution:1. U.S. Geological Survey, MS 980, Denver Federal Center, Lakewood, CO 80225, USA;2. U.S. Geological Survey, MS 974, Denver Federal Center, Lakewood, CO 80225, USA;3. Ponderosa Associates, 130 Miners Dr., Lafayette, CO 80026, USA;4. ADC Management Services, Inc., 355 S. Teller St., Suite 200, Lakewood, CO 80226, USA;1. Dipartimento di Scienze della Terra, Università degli Studi di Torino, Via Valperga Caluso, 35, I-10125 Torino, Italy;2. Borisyak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya 123, Moscow 117997, Russia;3. National Museum of Natural History, Smithsonian Institution (MRC-159), Washington, D.C. 20560, USA;4. Sezione di Geologia e Paleontologia, Museo Civico di Storia Naturale di Verona, Lungadige Porta Vittoria, 9, I-37129 Verona, Italy
Abstract:As part of the completion of studies on the Miocene fishes of the Chelif Basin (north-western Algeria), this paper represents a contribution to the knowledge of the Messinian gadiform diversity of this western Mediterranean, semi-enclosed, Neogene basin. A new genus and species of the family Macrouridae is erected (Razelainia paradoxa n. gen. et sp.), two specimens are tentatively referred to already existing taxa (Gadiculus cf. jonas; Merluccius cf. merluccius), and a species formerly assigned to the gadid genus Brosme is transferred to the genus Gaidropsarus (Gaidropsarus murdjadjensis). The macrourid Razelainia paradoxa n. gen. et sp. is characterized by an unusual combination of: plesiomorphic gadiform features, such as low vertebral number (presumed), well-developed caudal-fin rays, presence of a single continuous dorsal fin originating just posterior to the neurocranium, anal-fin rays slightly longer than dorsal-fin rays; and derived, typically macrourid features, such as the presence of spinoid scales and the anterior anal-fin pterygiophores extending forward over the abdominal wall. A paleoecological analysis reveals that the Messinian gadiform assemblage of the Chelif Basin had a subtropical/warm temperate affinity, with a marked north-eastern Atlantic-Mediterranean biogeographic character.
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