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1.
The laminated marine mudstones of the Late Jurassic of Kimmeridge, southern England, yield two exceptionally well-preserved partial skeletons of a previously unrecognised species of early batoid. These are described as a new genus and species, Kimmerobatis etchesi gen. et sp. nov. which has a general “guitarfish” bauplan as in all other batoids known from the Jurassic. This species possesses a combination of primitive characters such as centra present within the majority of the synarcual and antorbital cartilages that fail to reach the pectoral skeleton along with more derived characters, such as the lack of fin spines. Until now, little study has been carried out on the affinities of Jurassic batoids, despite their key role in understanding batoid evolution. Results from parsimony and likelihood phylogenetic reconstruction indicates that the whole-bodied Jurassic batoids Spathobatis, Belemnobatis, and Kimmerobatis gen. nov. form their own clade, Spathobatidae, and do not lend support to a monophyletic “Rhinobatidae”. Among Jurassic batoids, Kimmerobatis gen. nov. is most derived, but with derived characters being independently acquired compared to modern batoids (e.g. presence of a postpelvic process). The inclusion of whole bodied Jurassic fossils have generated a more resolved hypothesis of batoid evolution throughout the Cretaceous and into the Cenozoic.  相似文献   

2.
Teeth of a new species of hybodont shark Vectiselachos (Chondrichthyes: Lonchidiidae) are described from the late Aptian (Early Cretaceous) of southern England. Vectiselachos gosslingi sp. nov. has very distinctive coarse striations that form raised ridges over the occlusal surfaces of the crown.  相似文献   

3.
A thin phosphate-granule conglomerate within the Upper Cretaceous (middle Campanian) Rattlesnake Mountain sandstone member of the Aguja Formation preserves a diverse chondrichthyan and osteichthyan fauna. This highly fossiliferous deposit (the ‘Ten Bits Microsite’) yielded about 5000 teeth, spines, and denticles in a small amount of matrix (c. 150 kg). About 30 identifiable species of sharks, rays, and bony fishes are recognized. Two of the three most abundant chondrichthyan species at Ten Bits (Scapanorhynchus texanus and Ischyrhiza mira) are also the most common species in other middle to late Campanian marine vertebrate faunas along the Gulf and Atlantic Coastal Plain. The myliobatiform rays Brachyrhizodus and Rhombodus that occur at Ten Bits also appear to be characteristic of the Gulf and Atlantic Coast, as are lamniform sharks such as Cretalamna and Serratolamna. These taxa are entirely absent or extremely rare in Western Interior Campanian faunas. In contrast, some small orectolobiform sharks (Cantioscyllium, Chiloscyllium, Columbusia) and small rays (Protoplatyrhina) found at Ten Bits are common in shallow water faunas of the Western Interior and Texas Coastal Plain, but rarely reported from the eastern Gulf or Atlantic Coast. The common Western Interior ray Myledaphus bipartitus does not occur at Ten Bits or in any Gulf or Atlantic Coast fauna. Ptychotrygon agujaensis is abundantly represented in the Ten Bits fauna, but unknown in correlative marine faunas. Although Ptychotrygon occurs in all Western Interior, Gulf and Atlantic Coastal Plain faunas, it is represented elsewhere by apparently endemic species at each collection site. The differences between Western Interior, Gulf, and Atlantic Coastal Plain faunas probably reflect latitudinal variation in water temperature or salinity, or different oceanic water circulation patterns between the Western Interior Seaway and the Gulf or Atlantic Coast that restricted the distributions of some marine fish species. The Ten Bits fauna shares typical species with both Western Interior and Gulf and Atlantic Coast faunas, reflecting its position at the border between these provinces; however, the dominant taxa found at Ten Bits are the same as those found on the Gulf and Atlantic Coast, and indicate that western Texas was more closely allied biogeographically with that province than with the Western Interior of North America. One species tentatively identified in the Ten Bits fauna on the basis of a single tooth, Igdabatis cf. I. indicus, is otherwise known only from southern Europe and Asia, although a similar large myliobatid ray also occurs rarely in Texas Coastal Plain faunas. These occurrences indicate that western Texas may have been near the northern limit of the range for some tropical Tethyan marine vertebrate species.  相似文献   

4.
One hundred and thirty-six species, representing 67 genera have been recorded from the late Jurassic-Maastrichtian marine sediments of South Africa. The faunas show a major dichotomy across a regionally-developed late Cenomanian-early Coniacian hiatus with the Portlandian-Cenomanian Cytheruridae/Progonocytheridae/Schizocytheridae dominated faunas being replaced in the Coniacian by Trachyleberididae/Brachycytheridae/Schizocytheridae dominated faunas. Comparison with other Gondwanide localities shows that the two South African basins from which ostracods have been described (Outeniqua and Natal/Zululand) formed part of a Callovian-Cenomanian South Gondwana ostracod province that stretched from the Neuquen Basin of Argentina to Madagascar/Tanzania/Kutch and west Australia. The most characteristic and cosmopolitan forms within this province belong to the Majungaella/Amicytheridea/Progonocythere group, along with Arculicythere in the Aptian-Cenomanian.In Tanzania, (the only locality of the old South Gondwana province where the succession is complete) these assemblages are replaced in the Turonian by the influx of Brachycythere, and Cythereis and various other trachyleberids. Changes of a similar nature are seen whenever marine sedimentation resumed after the local “mid” Cretaceous hiatus (South Africa, India, Argentina). Argentina differs in not having Brachycythere, whose rapid appearance in the West Indian Ocean basin soon after its earliest record in Brazil, is attributed to the destruction of the barrier at the eastern end of the Walvis Ridge/Rio Grande Rise in late Cenomanian or early Turonian times. Despite this common element with Brazil and West Africa, the South African Coniacian to Maastrichtian faunas are closer to those of Tanzania and Australia than they are to either Argentina or Brazil/West Africa. In Zululand they show evidence of a steady increase in water depth, leading to the establishment of progressively more diverse cytheracean populations, with a particularly large increase across the Santonian/Campanian boundary.  相似文献   

5.
New trilobite material and the first graptolites from outcrop are described from the Am5 member of the Amdeh Formation near Al Fleij in northeast Oman. The sediments in which these faunas occur are interpreted as distal-shelf deposits with storm beds packed with brachiopods and orthoconic nautiloids. The deposits and its faunas are considered of late Darriwilian age and younger than the shallower-water Am5 deposits known from other outcrops. No palynomorphs could be recovered to confirm this due to the increased burial temperature the Al Fleij area has experienced.The trilobites are of considerable palaeo-biogeographic interest as few faunas of this age are known from the Arabian Plate, though their preservation precludes the establishment of new species. They include Isabelinia aff. glabrata, Liomegalaspides sp., Neseuretus tristani, Neseuretinus sp. and the deeper-water forms, Cyclopyge cf. C. bohemica, Arthrorhachis sp. and Brachypleura sp. The graptolites are pendent Didymograptus spp. of later Darriwilian type.Rare elements of the conodonts Nordiora, Amorphognathus and Microzarkodina have been recovered from shell beds that occur interbedded with the faunas. They too indicate a late Darriwilian age and differ from richer, restricted, shallow-water faunas known from the Am5 at other locations, and the more cosmopolitan shelf fauna from the Ayim Member of the Rann Formation of the United Arab Emirates.A coarser, 80 m-thick, terrestrial sequence containing igneous pebbles, included in the Am5 in the 1980s, is shown from the occurrence of fossil plants to be of Permian age and probably equivalent to the Basal Saiq Clastics of Jabal al Akhdar.  相似文献   

6.
The Kocali Complex in SE Turkey includes pelagic sediments (pelagic limestones, cherts, etc.), basic volcanic rocks of oceanic crust origin together with platform-derived sediments. Its depositional age was previously assigned as Late Jurassic–Early Cretaceous. In order to study the radiolarian contents of volcano-sedimentary sequences in this complex, six stratigraphic sections have been measured mainly at the NE and NW Adiyaman city.Radiolarian faunas from these stratigraphic sections reveal that the age of these sequences ranges from middle Carnian to Rhaetian. Based on these data, the depositional age of the complex is older than the previously assigned. Lithological characteristics (widespread Triassic basic volcanic rocks associated with pelagic sediments) and radiolarian contents of these sequences have close similarities with the sequences of the Alakircay Nappe of the Antalya Nappes in western and central Taurides.Based on taxonomic studies, 99 taxa have been determined of which one genus (Adiyamanium) and four species (Monocapnuchosphaera kocaliensis, Paronaella speciosa, Ferresium okuyucui and Adiyamanium crassum) are described as new.  相似文献   

7.
Several Gigantopithecus faunas associated with taxonomically undetermined hominoid fossils and/or stone artifacts are known from southern China. These faunas are particularly important for the study of the evolution of humans and other mammals in Asia. However, the geochronology of the Gigantopithecus faunas remains uncertain. In order to solve this problem, a program of geochronological studies of Gigantopithecus faunas in Guangxi Province was recently initiated. Chuifeng Cave is the first studied site, which yielded 92 Gigantopithecus blacki teeth associated with numerous other mammalian fossils. We carried out combined ESR/U-series dating of fossil teeth and sediment paleomagnetic studies. Our ESR results suggest that the lower layers at this cave can be dated to 1.92 ± 0.14 Ma and the upper layers can be dated to older than 1.38 ± 0.17 Ma. Correlation of the recognized magnetozones to the geomagnetic polarity timescale was achieved by combining magnetostratigraphic, biostratigraphic and ESR data. The combined chronologies establish an Olduvai subchron (1.945–1.778 Ma) for the lowermost Chuifeng Cave sediments. We also analyzed the enamel δ13C values of the Gigantopithecus faunas. Our results show that southern China was dominated by C3 plants during the early Pleistocene and that the Gigantopithecus faunas lived in a woodland-forest ecosystem.  相似文献   

8.
The vertebrate remains from the early Cenomanian Lagerstätte of Puy-Puy (Tonnay-Charente, Charente-Maritime, France) are described. They consist of two hybodont shark egg capsules (Palaeoxyris sp.) and a single isolated body contour feather. The hybodont shark Tribodus is regarded as the most likely producer of the egg capsules, while the feather belonged to an indeterminate (avian or non-avian) theropod. These rare specimens are the first vertebrate fossils recovered from the plant-bearing clay of Puy-Puy and add to the short faunal list of the locality, thus providing important information for the palaeoecological reconstruction of this mid-Cretaceous paralic Lagerstätte. The fossils described here represent the first Cenomanian occurrence of Palaeoxyris and one of the very few records of Cenomanian feathers.  相似文献   

9.
Thirteen decapod crustacean species, eight of which are new, from the mid-Cretaceous (late Albian) limestones of the Koskobilo quarry in northern Spain are described, illustrated, and discussed. They include: Graptocarcinus texanus; Navarrara betsieae gen. nov., sp. nov.; Acareprosopon bouvieri n. comb.; Laeviprosopon hispanicum sp. nov.; L. planum sp. nov.; L. edoi sp. nov.; L. crassum sp. nov.; Viaia robusta; Cretamaja granulata gen. nov., sp. nov.; Koskobilius postangustus gen. nov., sp. nov.; Navarrahomola hispanica; Glytodynomene alsasuensis; and Albenizus minutus gen. nov., sp. nov. Cretamaja and Koskobilius represent the oldest known spider crabs. In total, 36 species are now known from the Koskobilo locality based on 1078 specimens. To discover the magnitude of the diversity of the decapod fauna of Koskobilo, a comprehensive overview of decapod-rich localities and formations from the Cretaceous worldwide was compiled. It appears that Koskobilo is the most diverse decapod fauna from a single locality currently known from the Cretaceous. A rarefaction analysis shows that the maximum number of species is nearly reached. The number of genera, 26, is also unsurpassed for the Cretaceous. Forty-two species are found from localities within the Eguino Formation to which sediments from the Koskobilo quarry are ascribed, which is also unprecedented for a single formation within the Cretaceous. Evidence suggests that the most diverse decapod faunas from the Cretaceous are found in coral-associated limestones. This is consistent with evidence from the Recent, where decapod diversity is high in coral reefs compared to other habitats. This also suggests that the decapod peak diversity in Koskobilo is largely ecological in nature and not caused by a preservational bias. This is one of the most comprehensive studies on Cretaceous decapod diversity so far. Field work in coral-associated strata is expected to yield more decapod-rich faunas.  相似文献   

10.
Late Albian ammonite faunas from the Aitamir Formation of the Koppeh Dagh Basin in northeast Iran are described and illustrated. These comprise 14 taxa, several of which are recorded from Iran for the first time, namely Anahoplites planus (formerly recorded from central Iran in open nomenclature only), Semenoviceras solidus, Epihoplites (Metaclavites) iphitus, Hysteroceras orbignyi and Pseudhelicoceras robertianum. New records of Placenticeras grossouvrei extend the stratigraphic range of this species downwards into the Late Albian; previously it was known from the Early and Middle Cenomanian only. The record of the rare E. (M.) iphitus fills a palaeobiogeographic gap between Crimea and Tajikistan, and the holotype of Spath is re-illustrated here. Additionally, Epihoplites trapezoidalis, from the Late Albian of Tajikistan, is relegated into the synonymy of Spath's species. A large number of taxa typical of the Late Albian (upper part of the Gault Clay Formation) of northwest Europe indicate close palaeobiogeographic affinities with the Koppeh Dagh Basin and faunal exchange across the Russian Platform and Transcaspia. The stratigraphic succession of the ammonite faunas is used for a biostratigraphic subdivision of the upper Aitamir Formation.  相似文献   

11.
Composition and abundance of benthic and planktonic foraminifera in surface sediments of the brine-filled Shaban and Kebrit Deeps and some bathyal-slope environments in the northern Red Sea were examined for correlation with environmental conditions (e.g., bathymetry, sediment grain-size, organic matter, and carbonates) of the brine-filled deeps and normal Red Sea water. About 67 benthic foraminiferal species were recorded in these sediments. The lowest faunal density and diversity were recorded in the Shaban and Kebrit Deeps, whereas the highest density and diversity were recorded in the bathyal-slope sediments. Cluster analysis divided the benthic foraminiferal species into three major faunal assemblages. Buccella granulataGyroidinoides soldaniiBolivina persiensis assemblage dominated the 650–1,300 m depth due to predominance of oligotrophic, highly oxygenated bottom waters. The Melonis novozealandicumSpirophthalmidium acutimargo assemblage was recorded in the deep and bathyal-slope sediments indicating its tolerance for wider ranges of environmental conditions. The deeps were only dominated by the Brizalina spathulata assemblage indicating existence of un-totally anoxic conditions. The deeps yielded also very low planktonic foraminiferal density that may be attributed to occurrence of the seawater–brine interface which not only minimized the deposition of high buoyancy, large-test species (Globigerinoides sacculifer, Globigerinella siphonifera, and Orbulina universa), but also overestimated the small-test species (Globigerinoides ruber, Globoturborotalita rubescens, and Globigerinita glutinata) in the sediments. These findings should be taken into consideration when reconstructing paleoceanographic conditions of the Red Sea using core sediments from the brine-filled deeps.  相似文献   

12.
Valanginian marine sediments are widespread in the eastern margin of Asia from China northward over the Sikhote-Alin region, as far west as the Amur river, and as far north as the Amgun river. This was an interior type sea with abundant islands and a boreal fauna. It is surmised that Kamchatka, Sakhalin, Hokkaido, and the other Japanese islands were not invaded by the sea. A paralic coal basin is known on southwestern Honshu. Post-Valanginian folding and erosion preceded a Barremian-Albian marine transgression over the same general area. In these younger Cretaceous rocks southern faunas appear and the boreal fauna is present north of 45° latitude. Aucella and Aucellina species are important elements of the faunas. This review of faunas and correlations, including western North America, suggests that Hauterivian folding was a major event in North Pacific tectonic history. -- C. G. Tillman.  相似文献   

13.
14.
Ashdown Brickworks, near Bexhill, East Sussex, has produced a large number of vertebrate fossils from the Wadhurst Clay Formation, part of the Wealden Supergroup (Hastings Group; Valanginian; Lower Cretaceous). Here we describe the microvertebrate fauna of the ‘conglomerate bed’, representing a rich sample of taxa. While most of the recovered teeth and bones are abraded, some heavily, most can be identified to species level. The taxa include four species of hybodont sharks (Egertonodus basanus, Planohybodus ensis, Polyacrodus parvidens, P. brevicostatus), three taxa of bony fishes (an unidentified Lepidotes-like semionotiform, the pycnodontiform Ocloedus, and an albuliform), three taxa of crocodyliforms (the goniopholid Hulkepholis, a bernissartiid, and the atoposaurid Theriosuchus), and the theropod dinosaurs Baryonyx and an allosauroid. Sediments of the Wadhurst Clay Formation as a whole indicate freshwater to very slightly brackish-water environments of deposition, and the mainly aquatic time-averaged mixture of fishes and tetrapods recovered from the ‘conglomerate bed’, together with isolated terrestrial species, confirms this interpretation.  相似文献   

15.
《Quaternary Science Reviews》2007,26(9-10):1236-1300
Multidisciplinary investigations of the sequence at Beeches Pit, West Stow (Suffolk, UK), have a direct bearing the age of the Hoxnian Interglacial and its correlation with the continental Holsteinian and with the global marine record. At this site, glacial deposits (till and outwash gravels) referable to the Anglian Lowestoft Formation fill a subglacial channel cut in Chalk bedrock. Above these glacial deposits a series of interglacial sediments occurs, consisting of limnic, tufaceous and colluvial silts, lacking pollen but rich in shells, ostracods and vertebrates. Lower Palaeolithic flint artefacts of Acheulian character have also been recovered, including refitting examples. Charred material is abundant at certain horizons and many of the bones have been burned. Several discrete areas of burnt sediment are interpreted as hearths. The molluscan fauna comprises some 78 taxa and includes species of considerable zoogeographical and biostratigraphical importance. The land snail assemblage from the tufa consists of woodland taxa with no modern analogue, including species that are either extinct (e.g. Zonitoides sepultus) or which no longer live in Britain (e.g. Platyla polita, P. similis, Neniatlanta pauli). This is also the type locality of Retinella (Lyrodiscus) skertchlyi, which belongs to a subgenus of zonitid land snail now living only on the Canary Islands. There are indications from this fauna (‘the Lyrodiscus biome’) that the climate was wetter and perhaps warmer than the present day. The vertebrate fauna is also noteworthy with species of open habitats, such as rabbit (Oryctolagus cf. cuniculus), and of closed forest, such as squirrel (Sciurus sp.) and garden dormouse (Eliomys quercinus) present at different times. The occurrence of southern thermophiles, such as Aesculapian snake (Zamenis longissimus), indicates temperatures warmer than those of eastern England today. The upper levels include much material reworked from the interglacial sediments, although there is clear faunal evidence for climatic deterioration. Both the molluscan and vertebrate faunas suggest correlation of the interglacial sediments with the Hoxnian. Uranium series dates from the tufa (∼455 ka BP), TL dates from burnt flints (414±30 ka BP) and a range of amino acid racemization data all support correlation of this interglacial with MIS 11. However, four OSL dates from sand beneath the interglacial sequence yield a mean age of 261±31 ka BP, far younger than all other age determinations and far younger than implied by the biostratigraphy. Archaeologically the site is unusual in showing prolonged human occupation within closed deciduous forest and evidence for controlled use of fire in a Lower Palaeolithic context. Biostratigraphical correlations with other Lower Palaeolithic sites support the suggestion that Acheulian and Clactonian industries both occurred in southern Britain during the same substage of the Hoxnian, although not necessarily at precisely the same time. The characteristics of the MIS 11 interglacial in Britain are discussed in the light of evidence from Beeches Pit and elsewhere.  相似文献   

16.
Ostracod from the upper Lower to Middle Devonian rocks of the Argentine Precordillera Basin (Talacasto and Punta Negra formations) are studied. One new genus Pircawayra nov. gen., and five species (including three new: Pircawayra gigantea nov. gen. and sp., Lapazites trinodis nov. sp. and Keslingiella? teresae nov. sp.) are defined. The recorded ostracod fauna closely resembles that coeval from Bolivia and South Africa, exhibiting a remarkable endemism, not only at the genus level, but also at the species level. In addition to its low-diversity, the Malvinokaffric ostracod association is also characterized by having large, thick, coarsely ornamented and swollen valves. The similar ostracod composition from the Andean and South African basins suggests faunal exchange between these two areas. Based on the ostracod faunas, the Malvinokaffric Realm is clearly recognizable at least up to the Middle Devonian.  相似文献   

17.
Assemblages of foraminifers, ostracods and molluscs from temperate Ipswichian Stage (last temperate stage) sediments and overlying cold Devensian Stage (last cold stage) sediments at Somersham in the southern Fenland of Cambridgeshire have been analysed. The Ipswichian sediments contain faunas consistent with temperate brackish water conditions under tidal influence. The Devensian assemblages were recovered from a series of sands and gravels laterally accreting in a channel cutting into Ipswichian sediments. In contrast to the Ipswichian faunas, the faunas of particular Devensian samples show a complex mixture of temperate freshwater, brackish and marine taxa. The molluscs are mainly freshwater, with few land snails; they occur together with foraminifers and ostracods. Freshwater, brackish water and marine ostracods are present with foraminifers. A sample of Devensian fine laminated sediment in the channel was analysed for pollen; only abundant pre-Quaternary spores were present, with abundant foraminifers in the same sample. The taphonomy of the assemblages and the difficulties of their interpretation in environmental terms are discussed. The importance of taphonomy in assessing environments, climate, range of taxa and dating is stressed.  相似文献   

18.
In the lower part of sections at Skilvika and Linneelva, western Svalbard, marine silts and sands characterized by infinite radiocarbon ages (<40,000 BP) on shells are found. These sediments are covered by at least one basal till of Late Weichselian age. The till is overlain by marine sediments from the last deglaciation (12,800-10,000 BP) which contain shallow-water, subarctic foraminiferal assemblages, similar to modern near-glacial faunas from western Svalbard. The most common foraminifera in all zones in the sub-till sediments are Cassidulina reniforme, Astrononion gallowayi and/or Elphidium excavatum . The richest zones at both localities are found in the sub-till units and contain more than 20 foraminiferal species, including some boreal-arctic species. These faunal assemblages are similar to the living faunas on the west coast of Svalbard. Faunas from the postglacial climatic optimum are not yet described. We suggest that the foraminiferal assemblages in the sub-till sediment reflect Early or Middle Weichselian interstadial environments, although an Eemian interglacial cannot be excluded.  相似文献   

19.
We sampled 150 sites in fine-grained Plio-Pleistocene sediments of the Palm Springs and Imperial Formations. Sampling was confined to 3000 m of stratigraphically continuous section containing abundant vertebrate fossil remains of the Vallecito Creek, Arroyo Seco, and Layer Cake local faunas of the Irvingtonian and Blancan Land Mammal Ages. The magnetic stability of these sediments was sufficient to delineate the magnetic stratigraphy, which ranges from below the Cochiti event at the base, to the Matuyama reversed magnetic epoch at the top of the section. Eight faunal events are placed relative to the magnetic polarity sequence. They are cf. Pliohippus extinction, Geomys appearance, cf. Equus appearance, Tremarctos appearance, Hypolagus extinction, cf. Odocoileus appearance, Smilodon and ? Euceratherium appearance. The latter two faunal events characterize Irvingtonian Land Mammal Age. The transition from the Blancan to Irvingtonian Land Mammal Age occurs in the lower Matuyama magnetic epoch close to the Pliocene-Pleistocene boundary. The appearance of European migrants during the lower Matuyama magnetic epoch indicates that the Bering land bridge was exposed for animal migration between Europe and North America at this time.  相似文献   

20.
《Gondwana Research》2007,11(3-4):316-327
Mesozoic brackish-water bivalve faunas in Japan diversified in three steps: at the beginning of the Early Jurassic, Early and Late Cretaceous. The Hettangian Niranohama Fauna in northeastern Honshu represents the establishment of a heterodont-dominated brackish-water fauna that persisted until the early Late Cretaceous. No similar composition is known from the Triassic. The infauna consists mostly of non-siphonate and some short-siphonate heterodonts, while the epifauna is represented by diverse pteriomorphian families. In the Early Cretaceous Tetori Group in central Honshu, the long-siphonate heterodonts Tetoria (Corbiculidae) and the semi-infaunal soft-bottom oyster Crassostrea appeared. The evolutionary diversification of the latter, known as the most important element of modern brackish-water faunas, may thus originate at that time. In the early Late Cretaceous (Cenomanian) of the Goshoura and Mifune Groups in west Kyushu, several euryhaline deep-burrowing heterodont families, such as Veneridae and Tellinidae, further diversified in the brackish and marine environments. The Late Cretaceous is characterized by massive shell biolithic beds in which large Crassostrea species are common, a feature common for Cenozoic brackish-water faunas. The long-term changes in the composition of the brackish-water faunas in Japan represents thus an evolutionary record, irrespective of the severe physiological and environmental conditions imposed on the highly conservative nature of the fauna.  相似文献   

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