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1.
The effects of diel period and tow duration (5, 10 and 20 min) on samples of estuarine fauna in a beam trawl, were tested over bare sediment in Tuggerah Lake (New South Wales, Australia). Mean catch rates (numbers of fish caught 5 min−1) were significantly larger at night for the total numbers of individuals and abundant, economically important species of fish and invertebrates (e.g. Gerres subfasciatus, Metapenaeus macleayi, Penaeus plebejus). Greater proportions of larger fish were also caught at night for some species (e.g. G. subfasciatus, Acanthopagrus australis, Rhabdosargus sarba), but not across all tow durations. Multivariate analyses detected dissimilarities in the composition and structure of assemblages between diel periods, which were driven by species caught predominately, or in larger proportions, at night. Short tows (5 min) were more efficient than longer tows (10 or 20 min) for sampling the diversity of species (i.e. most species were caught in the first 5 min of a tow). There were, however, no clear or consistent patterns relating to the effect of tow duration on the catch rates of other variables, the size ranges of abundant species, or the structure and composition of assemblages. Our data confirm that at night, bare sediment is an important habitat for a wide size- and species-range of estuarine fish and invertebrates. In future, more cost-effective and reliable information concerning these taxa would be achieved by sampling with the beam trawl at night, using tow durations of 5 min. We also highlight a problem inherent in the design of many studies of diel variation of fauna (i.e. the potential non-independence of data among day and night periods) and discuss its solution.  相似文献   

2.
We examine the effect of a northward shift in the position of the southern hemisphere subpolar westerly winds (SWWs) on the vertical and horizontal distribution of temperature and salinity in the world ocean. A northward shift of the SWWs causes a latitudinal contraction of the subpolar gyres in the southern hemisphere (SH). In the Indian and Pacific, this leads to subsurface warming in the subtropical thermocline. As the southern margins of the gyres move into latitudes characterised by warmer surface air temperature (SAT), the layers at mid-depth below 400 m depth become ventilated by warmer water. We characterize the approximation of the ventilated thermocline in our coarse resolution model using a set of passive tracer experiments, and illustrate how the northward shift in the SWWs causes an equatorward shift in the latitude of origin of water ventilating layers deeper than 400 m in the Indian and Pacific, leaving the total surface ventilation of the upper 1200 m unchanged. In contrast, the latitudinal constraint on the Antarctic Circumpolar Current posed by the Drake Passage causes a cooling and freshening throughout the Atlantic thermocline; here, subsurface thermocline water originates from higher latitudes under the wind shift. On longer timescales Atlantic cooling and freshening is reinforced by a reduction in North Atlantic Deep Water (NADW) formation and surface salinification of the Indian and Pacific Oceans. In effect, the latitude of zero wind stress curl in the SWWs regulates the relative importance of the “cold water route” via the Drake Passage and the “warm water route” associated with thermocline water exchange via the Indian Ocean. Thus, a more northward location of the SWWs corresponds with a reduced salinity contrast between the Indian/ Pacific Oceans and the Atlantic. This results in reduced NADW formation. Also, a more northward location of the SWWs facilitates the injection of cool fresh Antarctic Intermediate Water into the South Atlantic subtropical gyre. Beyond these changes, on a millennial timescale, the deep ocean warms throughout the water column in response to the wind shift. Global salinity stratification also becomes less stable, as more saline water remains at the surface and accumulates in the Indian and Pacific thermocline. The freshening of the deep ocean reflects a reduced stirring of the global ocean due to reduced net circulation arising from a misalignment between the westerlies and the topographically constrained ACC. Our results lend support to the idea that a more equatorward location of the SWW maximum during glacial climates contributed to cooler and fresher conditions in the Atlantic, inhibiting NADW.  相似文献   

3.
The mid-domain effect was tested to evaluate the bathymetric patterns of the polychaete species richness in the Upper and Lower Gulf of California as a possible hypothesis to explain the species richness gradient, exploring the overlapping of species depth ranges towards the middle continental shelf. The bathymetric gradient of the number of species was estimated with the depth ranges of 554 polychaete species, and the mid-domain effect was tested using a Monte Carlo simulation program at bands of 10 m depth. The Upper (251 species) and Lower (491 species) Gulf regions showed clear differences in their faunal composition (Jaccard similarity index = 0.34); the species richness pattern was characterized by a highly significant presence of polychaetes with short depth ranges (< 10 m). The richness distribution could be described as a cubic polynomial curve, but the maximum values in both Gulf regions (141 and 317 species, respectively for Upper and Lower Gulf regions) are strongly biased to shallow waters (40 m). This is not consistent with the peak of diversity at 60–70 m predicted by the model. The observed patterns cannot be reproduced by the mid-domain effect, suggesting the existence of non-random factors affecting the species richness gradients in the Gulf.  相似文献   

4.
Using the hydroacoustic method with a 200 kHz scientific echo sounding system,the diel vertical migration(DVM) of the sound-scattering layer(SSL) in the Yellow Sea Bottom Cold Water(YSBCW) of the southeastern Yellow Sea was studied in April(spring) and August(summer) of 2010 and 2011.For each survey,13–27 hours of acoustic data were continuously collected at a stationary station.The acoustic volume scattering strength(Sv) data were analyzed with temperature profile data.In the spring of both 2010 and 2011,the SSL clearly showed the vertical migration throughout the entire water column,moving from the surface layer at night to near the bottom during the day.Conductivity,temperature,and depth data indicated that the entire water column was well mixed with low temperature of about 8 C.However,the SSL showed different patterns in the summers of 2010 and 2011.In the summer of 2010(28 C at the surface),the SSL migrated to near the bottom during the day,but there were two SSLs above and below the thermocline at depth of 10–30 m at night.In the summer of 2011(20 C at the surface),the SSL extended throughout the entire water column at night,possibly owing to an abrupt change in sea weather conditions caused by the passage of a Typhoon Muifa over the study area.It was concluded that the DVM patterns in summer in the YSBCW area may be greatly influenced by a strengthened or weakened thermocline.  相似文献   

5.
Meiofaunal standing stock and nematode community structure were investigated in the western continental shelf of India by collecting samples from every degree square of the shelf during two cruises of the FORV (Fishery and Oceanographic Research Vessel) Sagar Sampada, conducted in 1998 and 2001. Samples were collected from 30, 50, 100 and 200 m depths using a Smith Mc Intyre grab. Meiofaunal density ranged from 8 Ind. 10 cm−2 to 1208 Ind. 10 cm−2 and biomass from 0.07 mg 10 cm−2 to 6.11 mg 10 cm−2. Nematodes were the dominant meiofaunal group, contributing 88% of the density and 44% of the biomass. Harpacticoid copepods were the second important taxa, contributing 8% of both biomass and density. Altogether, 154 species of nematodes belonging to 28 families were recorded from the study area. Numerically, Desmodora spp., Dorylaimopsis sp., Tricoma spp., Theristus spp. and Halalaimus spp. were the dominant species. In general, there was a decrease in biomass and density of meiofauna and species diversity of nematodes with increase in depth. There was a 67% drop in species number from 51 to 100 m (106 species) to the shelf edge (35 species). Species richness and diversity indices showed consistent decrease with depth. The species dominance index was higher below 150 m depth. ANOSIM (from PRIMER) showed a significant difference between the nematodes of the near shore and shelf edge. Latitudinal variation was observed only in the number of nematode species. Biomass and abundance of nematodes were found to increase from coarse to fine sediment, while copepods showed an opposite trend. Multivariate analyses of nematode communities did not reveal any latitudinal or substratum differences. Variables such as depth, latitude, organic matter (OM) and amount of clay were the most relevant parameters influencing the biomass and density of meiofauna, while depth and temperature were the important parameters explaining the distribution of the nematode communities along the western Indian shelf.  相似文献   

6.
Ocean upwelling rates are difficult to measure because of the relatively small velocities involved, and therefore are typically inferred from indirect methods such as heat budget estimates or tracer observations. Here we present the first results using a novel technique, based on the isotope 7Be, to infer rates of upwelling along the equator. Beryllium-7 (half-life=53.3 d) is a cosmic-ray produced radioactive nuclide that is deposited by rainfall upon the ocean surface and subsequently enriched and homogenized within the mixed layer. Previous investigations have utilized the penetration of characteristically high mixed layer concentrations into the upper thermocline to trace ocean ventilation and subduction over seasonal timescales. Here, the tracer is used in a reverse sense; that is, the 7Be concentration in the usually 7Be-rich surface mixed layer will be diluted from penetration of 7Be “dead” water upwelled from below. This dilution provides a means to infer upwelling rates. Furthermore, with knowledge of upwelling rates, 7Be profiles can be used to constrain vertical diffusivity within the upper thermocline. These ideas were tested with samples collected during the Tropical Atlantic Climate Experiment (TACE) cruise (May 22-June 27, 2009). The observations indicated a nearly linear relationship between 7Be inventory and mixed layer temperature, as with increased upwelling, lower mixed layer temperatures correspond to greater 7Be dilution from depth. With this data, upwelling rates were estimated at a number of stations near the equator between 0°E and 30°W within and adjacent to the equatorial cold tongue. The derived upwelling rates ranged from 0 to 2.2 m/d, with maximum values found between the equator and 2°S. The corresponding Kz values derived for the upper thermocline were in the range 1-4×10−4 m2/s.  相似文献   

7.
Protistan community structure was examined from 6 depths (1.5, 20, 42, 150, 500, 880 m) at a coastal ocean site in the San Pedro Channel, California. A total of 856 partial length 18S rDNA protistan sequences from the six clone libraries were analyzed to characterize diversity present at each depth. The sequences were grouped into a total of 259 Operational Taxonomic Units (OTUs) that were inferred using an automated OTU calling program that formed OTUs with approximately species-level distinction (95% sequence similarity). Most OTUs (194 out of 259) were observed at only one specific depth, and only two were present in clone libraries from all depths. OTUs were obtained from 21 major protistan taxonomic groups determined by their closest BLAST matches to identified protists in the NCBI database. Approximately 74% of the detected OTUs belonged to the Chromalveolates, with Group II alveolates making up the largest single group. Protistan assemblages at euphotic depths (1.5, 20 and 42 m) were characterized by the presence of clades that contained phototrophic species (stramenopiles, chlorophytes and haptophytes) as well as consumers (especially ciliates). Assemblages in the lower water column (150, 500 and 800 m) were distinct from communities at shallow depths because of strong contributions from taxa belonging to euglenozoans, acantharians, polycystines and Taxopodida (Sticholonche spp. and close relatives). Species richness (Chao I estimate) and diversity (Shannon index) were highest within the euphotic zone and at 150 m, and lowest for protistan assemblages located in the oxygen minimum zone (500 and 880 m). Multivariate analyses (Bray-Curtis coefficient) confirmed that protistan assemblage composition differed significantly when samples were grouped into shallow (≤150 m) and deep water assemblages (≥150 m).  相似文献   

8.
We hypothesized that temporal variation in fish species composition and community structure in a low complexity habitat in the Pueblo Viejo Lagoon, Mexico, is influenced by diel light/dark cycles and tidal stage, and by seasonal changes in salinity and temperature. We collected a total of 17,661 individuals during 2‐h interval sampling over six bi‐monthly 24‐h sampling cycles representing 53 species, of which 11 (~20%) were previously unknown in the system. Diel variation indicated that significantly higher numbers of individuals and species were caught at night, whereas diversity and evenness were higher during the day. Species richness was significantly higher in July and January, whereas diversity and evenness peaked around May; both were correlated with temperature. Diel variation in species composition was influenced primarily by the light/dark cycle. Cluster analyses of each diel cycle separated fish assemblages from midday samples from those of nocturnal samples, separated by an extended wide transition period as fish moved at dawn and during the late afternoon/dusk. Significant shifts (as determined by MANOVA) in assemblage structure occurred between months. Canonical correspondence analysis showed that temperature and day/night effects were the most important environmental variables structuring the fish community. This constrained ordination also defined species with specific habitat preferences as follows: (i) diurnal, warm temperature species (mainly planktivores) (Brevoortia gunteri, Cetengraulis edentulus, Diapterus auratus, and Membras martinica); (ii) nocturnal, warm temperature species (mainly predators) (Citharichthys spilopterus, Cathorops melanopus, and Bairdiella spp.); and (iii) low temperature, diurnal species (Brevoortia patronus and Mugil curema) or those with twilight and nocturnal distributions (Anchoa mitchilli, the most numerically abundant species). Our results indicate that diel and seasonal changes in fish community structure were mainly related to day/night cycles and temperature regimes.  相似文献   

9.
徐亚军  赵亮  原野 《海洋学报》2016,38(8):124-131
浮游动物的昼夜迁移活动与其种群变动和摄食节律紧密联系,浮游动物昼夜移动的研究已经成为种群动力学研究的一个重要组成部分。2013年夏季在浙江东部近海,结合声学多普勒流速剖面仪(ADCP)和激光粒径分析仪(LISST-100)等仪器进行了一次定点周日连续观测。通过声学反演方法,得到后向散射强度剖面的时间变化,结合LISST-100得到的水体悬浮物粒径谱,研究了浮游动物垂直迁移及其习性。分析发现了可能是精致真刺水蚤的一次昼夜垂直迁移过程,其在夜间进入跃层附近进食,白天蛰伏于底层低温高盐的台湾暖流水中,垂向迁移速度达到了0.05 m/s。LISST-100观测还发现在夜间跃层边界处大粒径颗粒聚集和100~150 μm大小的颗粒物的减少,水体中不同粒径的悬浮颗粒物有明显的昼夜变化节律,推测水体中不同层次生物群落结构存在昼夜差异。  相似文献   

10.
Zooplankton in the coastal upwelling region off northern Chile may play a significant biogeochemical role by promoting carbon flux into the subsurface OMZ (oxygen minimum zone). This work identifies the dominant zooplankton species inhabiting the area influenced by the OMZ in March 2000 off Iquique (20°S, northern Chile). Abundance and vertical distribution studies revealed 17 copepod and 9 euphausiid species distributed between the surface and 600 m at four stations sampled both by day and by night. Some abundant species remained in the well-oxygenated upper layer (30 m), with no evidence of diel vertical migration, apparently restricted by a shallow (40–60 m) oxycline. Other species, however, were found closely associated with the OMZ. The large-sized copepod Eucalanus inermis was found below the oxycline and performed diel vertical migrations into the OMZ, whereas the very abundant Euphausia mucronata performed extensive diel vertical migrations between the surface waters and the core of the OMZ (200 m), even crossing it. A complete assessment of copepods and euphausiids revealed that the whole sampled water column (0–600 m) is occupied by distinct species having well-defined habitats, some of them within the OMZ. Ontogenetic migrations were evident in Eucalanidae and E. mucronata. Estimates of species biomass showed a substantial (>75% of total zooplankton biomass) daily exchange of C between the photic layer and the OMZ. Both E. inermis and E. mucronata can actively exchange about 37.8 g C m−2 d−1 between the upper well-oxygenated (0–60 m) layer and the deeper (60–600 m) OMZ layer. This migrant biomass may contribute about 7.2 g C m−2 d−1 to the OMZ system through respiration, mortality, and production of fecal pellets within the OMZ. This movement of zooplankton in and out of the OMZ, mainly as a result of the migratory behavior of E. mucronata, suggests a very efficient mechanism for introducing large amounts of freshly produced carbon into the OMZ system and should, therefore, be considered when establishing C budgets for coastal upwelling systems.  相似文献   

11.
In order to understand the mechanism by which zooplankters from different origins co-occur during the upwelling season within Golfo de Arauco, one of the most productive areas in central Chile, we assessed short term variations in the vertical distribution of the most abundant copepod species. Fine-scale, day and night vertical zooplankton sampling was done with a pump over 12 days in summer. The water column in the gulf consisted of three layers: Equatorial Subsurface Water of low dissolved oxygen content in the deeper part of the water column, strong temperature and oxygen gradients at mid-depth (15-25 m), and a layer of warmer, more oxygenated, less saline water at the surface. Copepods within the gulf originated from offshore, from the continental shelf, and from the coastal area. Most taxa showed distinctive vertical distributions. Three copepod groups were identified by their mean weighted depths of residence. One group included shallow residents found above the thermocline/oxycline (Acartia tonsa, Centropages brachiatus, Corycaeus sp., Paracalanus parvus, Oncaea sp.). A second group was comprised by species distributed at or below the thermocline/oxycline (Oithona sp., Oncaea conifera, Lucicutia sp., Metridia sp., Heterorhabdus papilliger). The third group was composed of vertical migrators that crossed the thermocline/oxycline (Calanus chilensis, Calanoides patagoniensis, Aetideus armatus, Pleuromamma piseki). In spite of their different vertical distribution ranges, the most abundant and frequent copepod species (P. parvus, C. chilensis, C. patagoniensis, C. brachiatus) share a common capacity to withstand wide ranges of oxygen concentration and temperature. This characteristic, along with the capacity to vary their life strategies under different environmental conditions, seems to facilitate the maintenance of large numbers of copepods in coastal waters along the Humboldt Current.  相似文献   

12.
Different estimates were used to assess the diversity of the total macrofauna and its major taxonomic groups separately from a broad bathymetric range at a site in the NE Atlantic. In the Goban Spur region, a transect was sampled from the shelf to the abyssal plain over a depth range from 200 to 4500 m and in the Porcupine Sea Bight two stations were sampled (at 3670 m and 4115 m). Species diversity (the number of species per number of individuals) increased with increasing water depth, both when expressed as Hurlbert's E(Sn) and as Shannon's H′log e. The expected number of species in a 100-individual sample E(S100) of total macrofauna increased from 30 on the shelf to 68 on the abyssal plain. Evenness (the proportional abundance of species), estimated with Shannon's J′, also increased with water depth from 0.66 to 0.91, whereas dominance (Simpson's D) decreased from 0.09 to 0.01. Species richness (the number of species per unit of area), however, showed a parabolic pattern with a peak at the upper slope. The largest number of species was found at the slope station at 1425 m (232 species within 0.66 m2). It is argued that species richness is not a synonym of species diversity, but that species richness depends both on species density (which decreases with increasing water depth) and on species diversity. Across the whole bathymetric range (200 to 4500 m) a total of 696 species within 8327 specimens in a total sampled area of 4.12 m2 were counted, yielding mean values of 12 individuals per species and 169 species per m2. Different communities were found to exist on the shelf, slope and abyss. It is suggested that this could have been caused by different selection processes. Differences in life-history strategies and organic-matter supply could (at least partly) explain the different community structures and diversity patterns found along the depth gradient.  相似文献   

13.
Few studies have been conducted to quantify and understand the role of melatonin in invertebrates, and particularly in crustaceans and in deep-sea animals. In this study, we examined day–night differences in haemolymph melatonin of the burrowing decapod crustacean Nephrops norvegicus (L.) during exposure to cycles of monochromatic blue light (480 nm) and darkness cycles of 10 and 0.1 lx. These differential intensity conditions simulate illumination at the depth of the shelf (80–100 m) and of the slope (300–400 m), where these lobster populations are chiefly found in the Western Mediterranean Sea. Our objectives were: (a) to verify the presence of melatonin in the haemolymph of this species using liquid chromatography/tandem mass spectrometry (LC–MS/MS) and fluorescence HPLC (HPLC); and (b) to study the relationship between diel variations in melatonin concentration and locomotor rhythms, in order to examine whether the former influences behaviour. Melatonin was identified in LC–MS/MS by Q1 and Q3 mass peaks at an elution time of 3.7 min, and it was also detected by HPLC. Melatonin concentration was found to be two orders of magnitude higher at 10 lx (4.8±5.3 ng ml−1) than at 0.1 lx (0.06±0.03 ng ml−1). Also, the increase at daytime in 10 lx was absent in 0.1 lx. When the locomotor rhythm of animals exposed to both photoperiod regimes was compared, the diel periodicity was found to be preserved, but the timing of activity shifted from night to day. Extrapolating these data to the field, we interpret our results to mean that locomotor activity preserves its diel character, but not its phase and amplitude, in a bathymetric range where haemolymph melatonin reduces its concentration and rhythmic fluctuation.  相似文献   

14.
Understanding temporal variation at the scale of weeks to months is critical to understanding broad temporal patterns in diversity in the same way as understanding diversity across landscapes relies on understanding variation at the scale of meters. However, whereas small-scale spatial variation in temperate reef algal assemblages has been extensively studied, fine-scale temporal changes have not been well addressed. By sampling the macroalgae of a subtidal reef near Perth (Australia), dominated by the small kelp Ecklonia radiata, every ∼40 days over a 2-year period, we were able to test whether temporal changes in species richness, assemblage structure and species turn-over were related to seasonal changes in surface temperature, solar radiation and wave height. A total of 93 macroalgal taxa were identified, and species richness per sampling time ranged from 25 to 64 taxa 1.25 m−2. Biomass of E. radiata was positively correlated with changes in sea surface temperature and light, and negatively correlated with wave height. Species richness, assemblage structure and turn-over of other macroalgae were more associated with seasonal changes in kelp biomass than environmental variables per se. We conclude that seasonal changes in environmental conditions drive changes in the kelp canopy, which in turn drive changes in species richness and assemblage structure. This suggests that habitat-formers such as kelps can exert a strong temporal influence on associated communities, analogous to well-described spatial influences. Thus, as kelp canopy biomass expands and retracts over time-scales of weeks to months, so does available space for colonization and growth, resulting in a high species turn-over. Species richness is therefore increased and maintained through time, in the same way as canopy-gap mosaics increase and maintain species richness across spatial landscapes.  相似文献   

15.
Siphonophores were sampled using stratified 1.67-m2-mouth 330-μm mesh nets during both day- and nighttime in Sagami Bay (35°0.50′N, 139°20′E), off south-eastern Japan during a 9-day period in March 2006, when the Kuroshio Current was in an offshore non-large meander phase. The samples were collected at 50-m intervals spanning from 1,000 m depth to the surface. Fifty-eight species and 11 eudoxid forms of unknown parentage were collected, of which 5 represent first-time records from Japanese waters, and Clausophyes laetmata Pugh and Pagès 1993, Eudoxia cf. galathea Moser 1925 and Lensia panikkari Daniel 1970 were recorded for the first time not only from Japan but from the entire Pacific Ocean. The highest abundance and diversity was found below the thermocline during the day, and above it at night. A second peak in abundance, around 400 m depth, associated with a decrease in diversity, could be linked to the increase in abundance of a single species—Dimophyes arctica (Chun 1897). The siphonophore communities could be related to the different water masses in the Bay, with an important influence of lateral transport of both tropical and subarctic species into the Bay by the different water masses.  相似文献   

16.
Successive measurements of the size distribution and abundance of marine snow in the upper 100 m of the Santa Barbara Channel, California, with an in situ still camera system following 11 tagged water masses revealed a consistent pattern of nighttime decreases in the abundance of large particles. A net nocturnal reduction in particulate flux from the upper 100 m as calculated from camera profiles occurred in 75% of the day–night comparisons, and nighttime aggregate carbon losses resulted in a 38% average reduction in camera-derived aggregate flux. Intensive investigation of three stations for 24–48 h each indicated that nighttime decreases in aggregate concentrations and derived aggregate flux could be registered throughout the observed water column. Nocturnal decreases in marine snow concentration are unlikely to result from diel variations in the production of marine snow either as feeding webs of zooplankton or through variations in aggregation rates of smaller particles. Moreover, measured diel variations in the intensity of surface mixing and convective overturn during one of the 24 h deployments were not intense enough to produce aggregate fragmentation and reduced aggregate flux. Nighttime increases in large crustacean zooplankton (i.e., euphausiids and the large copepod Calanus pacificus) could explain some or all of the reduction in aggregate abundance at most stations. Fragmentation and consumption of marine snow by migrating macrozooplankton could produce our observed synchronous diel cycles in marine snow concentration. This is the first empirical evidence of a diel pattern in the concentration and calculated particulate flux of large sinking particles in near-surface waters. The data presented here are consistent with the only other existing diel study, which also reported decreases in marine snow abundance at night at 270 m depths in the oceanic north Atlantic. Diel variations in the sizes and concentrations of marine snow may impact water column processes dependent upon particle availability and size, such as grazing and remineralization, and may generate a diel cycle of food availability to the benthos.  相似文献   

17.
Modelling trace metal concentration distributions in estuarine waters   总被引:2,自引:1,他引:2  
The concentration of dissolved organic carbon (DOC) was measured every few months from September 2000 through October 2001 at a coastal location in the center of Suruga Bay, Japan (34°51′N, 138°38′E). Water samples were collected three times per day (midday, night and predawn). DOC concentrations ranged from 91.3 to 45.2 μM C on the surface to 100 m depth. Diel variation in DOC concentrations, among the three sampling times, was greater in the upper 20 m, with a maximum difference of 21.7 μM C in July 2001, and reflected in diel DOC inventory variations from the surface to 50 m. Diel variations were controlled by both physical and biological factors. DOC concentrations were significantly correlated with potential density in the deeper layers (100–1000 m), indicating that the distribution of DOC concentrations in the deeper layer was mainly due to mixing. Most DOC concentrations in the upper layer (0–50 m) did not display the same relationship as in the deeper layer. Using the relationship with potential density at 100–1000 m, the DOC concentration in the upper layer, due simply to mixing, was calculated. The difference between the calculated and observed DOC was used to estimate biological contribution. The biological contributions to the DOC inventory in the upper layer (0–50 m) were found greatly in November 2000 and April 2001. This indicates that excess DOC accumulated, by biological processes, in the upper layer during these periods. In November 2000, the excess DOC in the inventory was constant throughout the sampling days (0.36–0.37 mol C m−2), whereas diel variations of DOC in the vertical profile were large and contrary to the variation between 10 and 20 m. This suggests that the excess DOC was contributed biologically during daytime in the uppermost layer and reached to the 50 m depth by deeper mixing. As a result, the inventory appeared to be stable over a day because of the compensating effects of DOC production and consumption throughout 50 m. In contrast, in spring and summer, there was a distinct diel inventory decrease in the nighttime, with apparent rates ranging from −0.61 to −0.35 μM C h−1. It is probable that the DOC, which accumulated during the daytime, was mostly labile, with a turnover time of a few hours. The results indicate that the dynamics of diel DOC variations varied seasonally, and suggest that these variations need to be considered when estimating seasonal DOC pools in the coastal ocean.  相似文献   

18.
北黄海典型水域秋冬季浮游植物群落的昼夜变化   总被引:1,自引:0,他引:1  
利用冬季(2007年1月)和秋季(2007年10月)在北黄海中部昼夜连续观测所获浮游植物样品,分析了浮游植物群落的昼夜变化.结果表明:浮游植物丰度在各层存在昼夜变化,冬季10~20 m层主要在白天出现高丰度,而20~30 m层和30~底层都是在夜间出现高丰度,冬季丰度垂直分布相对较均匀.秋季各层都主要在夜间出现高丰度,丰度垂直分布是下层水体明显比上层高.2个季节优势种昼夜组成都比较稳定.冬季优势种为具槽帕拉藻(Paralia sulcata),主要分布在下层水体.秋季罗氏角毛藻(Chaetoceros lauderi)和密连角毛藻(Chaetoceros densus)成为优势种,两者分别以下层和上层水体分布较多.冬季浮游植物群落多样性和均匀度昼夜变化都不明显,秋季0~10 m层和20~30 m层多样性和均匀度昼夜变化较明显.从季节变化来看,秋季多样性要高于冬季,均匀度的季节变化不明显.  相似文献   

19.
A time-series sediment trap was operated from July 2003 to July 2008 at a station located in the 10°N thermocline ridge of the northeastern equatorial Pacific (10°30′N, 131°20′W), with the aim of understanding variations in natural background sinking-particle flux and the influence on such fluxes of ENSO (El Niño-Southern Oscillation). Each one of weak El Niño, moderate El Niño and moderate La Niña were observed during the monitoring period. During non-ENSO periods, total mass fluxes varied from 4.1 to 36.9 mg m−2 d−1, with a distinct seasonal variation, ranging from an average flux of 14.0 mg m−2 d−1 in the warm season (June-November) to 25.3 mg m−2 d−1 in the cold season (December-May). This seasonal fluctuation was characterized by a distinct difference in CaCO3 flux between the two seasons. The enhanced particle fluxes during the cold season are attributed to the supply of nutrient-enriched subsurface water by wind-driven vertical mixing, supported by a simultaneous reduction in sea surface temperature and enhanced trade winds. The weak El Niño event occurred in the monitoring period had no recognizable effect on particle fluxes in the study area, but the moderate El Niño event was accompanied by a significant reduction in particle fluxes to 60% of the average background value in the warm season. In contrast, particle fluxes during the moderate La Niña increased to a maximum value of 129.9 mg m−2 d−1, almost three times the average background value. Organic carbon and biogenic silica fluxes were most sensitive to the El Niño and La Niña conditions. The observed variations of particle fluxes are synchronized with those of chlorophyll-a, suggesting primary productivity for the main cause of flux change. The present data indicate that marked seasonal variability in background fluxes commonly exceeds the variability associated with ENSO and post-ENSO signals, which should be taken into account when evaluating the influence of ENSO on sinking particle fluxes in the 10°N thermocline ridge area.  相似文献   

20.
We study the formation of the vertical distribution of temperature in the upper layer of the oceans (0– 300m) at low latitudes (10°N-10°S) by using the nonlinear dependence of the vertical heat flow on the vertical temperature gradient with regard for the influence of the bulk absorption of solar radiation and heat sink on the temperature of water. The thermocline is formed under the condition that the modulus of temperature gradient attains values for which their subsequent increase leads either to insignificant variations of the heat flow or even to its decrease. We consider the possibility of solution of inverse problems for the evaluation of the heat-exchange coefficient and the parameter of the heat sink. For the Equatorial Atlantic considered as an example, we compare the theoretical results with the data of instrumental measurements.__________Translated from Morskoi Gidrofizicheskii Zhurnal, No. 6, pp. 10–16, November–December, 2004.  相似文献   

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