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1.
A third genus and species of fossil pygidicranid earwigs from Burmese amber is described and figured as Stonychopygia leptocerca Engel, Huang, Thomas, and Cai, gen. et sp. nov. (type species of Stonychopygia Engel and Huang, gen. nov.; Stonychopygiinae Engel and Huang, subfam. nov.). Stonychopygia have features similar to the subfamily Echinosomatinae such as the combination of shorter, subequal second and third flagellomeres and femora that are neither compressed or keeled. However, the new fossil species differs from echinosomatines in many details, particularly the slender form, head longer than wide, slender scape, slender and elongate forceps, and absence of stout, short bristles over the integument. The new fossil is compared with its contemporaneous and modern relatives, and comments are provided regarding the classification of Pygidicranoidea, with the genus Haplodiplatys Hincks removed from Diplatyidae to Haplodiplatyidae Engel, fam. nov. 相似文献
2.
Two new genera and species of Cretaceous webspinners (Embiodea) are described and figured, both preserved in amber from northern Myanmar. Atmetoclothoda orthotenes Engel and Huang, gen. et sp. nov., is the first fossil representative of the putatively primitive family Clothodidae, and is segregated into its own subfamily, Atmetoclothodinae Engel and Huang, subfam. nov., owing to its primitive retention of a distinct mentum, a quadrate submentum with straight borders, a postocciptal suture that meets the hypostomal sulcus at the posterior tentorial pit, and subgenae that do not meet medially (thus a ventral bridge or gula is lacking), completely symmetrical terminalia, and unsegmented cerci that apically bear a small inner lamellar projection. Litoclostes delicatus Engel and Huang, gen. et sp. nov., is the first definitive fossil of Oligotomidae (not including a subfossil species in Pleistocene copal and which is likely synonymous with an extant taxon), and differs from modern genera in a combination of head, tarsal, and terminalic morphology. The new fossil species double the known Cretaceous fossils and add a further two families to the Mesozoic record for the order. 相似文献
3.
A new fossil stick-insect (Phasmatodea) is described and figured from a male preserved in Cretaceous amber from Myanmar. Echinosomiscus primoticus Engel and Wang, gen. et sp. nov., is a robust, somewhat-compressed stick-insect bearing abundant acanthae over the head and body, and remarkably lacks an area apicalis on the tibiae. The species is described and assigned to a new, extinct subfamily of Phasmatidae s.l., as Echinosomiscinae Engel, subfam. nov. Brief remarks are made regarding the dating of phasmatodean lineages, with E. primoticus providing the first reliable evidence for Euphasmatodea and even Neophasmatodea in the Cenomanian. 相似文献
4.
A second Mesozoic twisted-wing parasite (Strepsiptera) is described and figured based on an exceptionally well-preserved male in mid-Cretaceous (Cenomanian) amber from northern Myanmar. Phthanoxenos nervosus Engel and Huang, gen. et sp. nov., is distinguished from other strepsipteran lineages, particularly the contemporaneous Cretostylops engeli Grimaldi and Kathirithamby, also in Burmese amber, and Protoxenos janzeni Pohl et al. in mid-Eocene Baltic amber, and assigned to a new family, Phthanoxenidae Engel and Huang, fam. nov. Phthanoxenos exhibit features indicative of a more primitive phylogenetic position than Cretostylopidae but still more derived than Protoxenidae. Brief remarks are made on the geological history of the Strepsiptera. 相似文献
5.
A new species of Meropeidae (earwigfly) is described and figured based on an exceptionally well-preserved individual in mid-Cretaceous amber from Myanmar. Burmomerope clara Zhao and Wang, sp. nov. is distinguished from the type species B. eureka Grimaldi and Engel, 2013 by presence of broader wings with six longitudinal veins in radial sector and seven in medial field, CuA with two terminal branches, and long setae on the anterior margin of the wing. A detailed comparison of the forewings venation in all fossil and extant species is given. The new find is the third fossil species of Meropeidae and also the first fossil female to be described. The female genital structure of B. clara sp. nov. is remarkably similar to that of extant species, revealing 100 million years of morphological conservatism, and thus highlighting the antiquity of the group. 相似文献
6.
Mites are relatively common and diverse in fossiliferous ambers, but remain essentially unstudied. Here, we report on five new oribatid fossil species from Lower Cretaceous Spanish amber, including representatives of three superfamilies, and five families of the Oribatida. Hypovertex hispanicus sp. nov. and Tenuelamellarea estefaniae sp. nov. are described from amber pieces discovered in the San Just outcrop (Teruel Province). This is the first time fossil oribatid mites have been discovered in the El Soplao outcrop (Cantabria Province) and, here, we describe the following new species: Afronothrus ornosae sp. nov., Nothrus vazquezae sp. nov., and Platyliodes sellnicki sp. nov. The taxa are discussed in relation to other fossil lineages of Oribatida as well as in relation to their modern counterparts. Some of the inclusions were imaged using confocal laser scanning microscopy, demonstrating the potential of this technique for studying fossil mites in amber. A table, including all the known Mesozoic oribatid mites preserved in amber, is included. 相似文献
7.
《Cretaceous Research》2012,33(6):794-805
Three new species within the stigmaphronid genus Tagsmiphron Engel and Grimaldi, 2009, and one new species within the megaspilid genus Conostigmus Dahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage. 相似文献
8.
A new species of Glaesoconis Meinander (Neuroptera: Coniopterygidae) from the Santonian Taimyr amber
Glaesoconis popovi sp. nov. (Neuroptera: Coniopterygidae) is described from Upper Cretaceous (Santonian) Taimyr amber of northern Siberia (Yantardakh locality). The new species may be distinguished from others in the genus by much smaller eyes and the shape of the terminal segment of the maxillary palpus. The generic affinity of Glaesoconis baliopteryx Engel, 2004 from the mid-Cretaceous Burmese amber needs confirmation. 相似文献
9.
Sclerogibbid wasps are obligate parasitoids of webspinners (Embiodea). Both groups have a particularly scarce geological record and are known since the Cretaceous: there are only four species of webspinners known from Burmese amber, and only two sclerogibbids were described from Barremian Lebanese and Cenomanian Burmese ambers. Here we report transferred genus from Aptian Choshi (Japan) amber and new sclerogibbids from Cenomanian Burmese and Charentese (France) ambers. The taxa described from Burmese amber are: Burmasclerogibba aptera gen. et sp. nov., Cretosclerogibba gen. nov. (with C. antennalis sp. nov., C. contractocollis sp. nov., C. neli sp. nov. and C. rasnitsyni sp. nov.) and Edrossia vetusta gen. et sp. nov. The first European fossil sclerogibbid Gallosclerogibba alnensis gen. et sp. nov. is described from Charentese amber. The holotype of Chosia yamadai Fujiyama, from Choshi amber, is re-described; it appears to be the oldest Laurasian sclerogibbid. The significant abundance and variety of Burmese sclerogibbid wasps (60% of fossil species known worldwide), as proxy of their hosts, were probably caused by the protection granted to them by the silk webs and possibly by the limited predation from ornithuromorph birds or crown-group ants. While all three extant sclerogibbid genera have apterous females, genera with winged females (Cretosclerogibba and Edrossia) dominated in Burmese amber. Small silk galleries from hosts may have favored the preservation of wings in females of Cretaceous sclerogibbids. Most new species described in the present paper, in addition to C. yamadai, are characterized by a very slender neck and a very long frontal process concealing the antennal toruli. These characters disappeared in extant species. We suggest that this loss was caused by a change in the fauna of predators, penalizing species with long neck and rostrum. 相似文献
10.
Palaeomanicapsocus margoae gen. et sp. nov. and Palaeomanicapsocus fouadi gen. et sp. nov. are characterized, described, illustrated, from the Cretaceous Burmese amber. Their phylogenetic position is discussed. These fossil taxa are the first manicapsocid barklice to be described from the Burmese amber. 相似文献
11.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight. 相似文献
12.
Millipedes (Diplopoda) are an important fossil group of land arthropods in the Palaeozoic. However, there is a gap in the Mesozoic, with only slightly more than a dozen fossils being known, until the much more recent fossil records mainly from Cenozoic Dominican and Baltic ambers become available. Burmese amber, precisely dated to the Late Cretaceous, is known for an apparently rich but still undescribed millipede fauna, and might represent the ideal bridge to close this gap in the millipede fossil record. In addition, modern micro-computed tomography (μCT) technology allows complex 3D reconstructions of objects fossilized in amber. Here we utilize μCT technology to describe the first millipedes from Burmese amber, which are also the first two fossil representatives of the order Siphoniulida. Siphoniulida, with two known species and just nine recorded specimens, are the rarest and least known of the 16 extant orders of the Diplopoda. Records are known from Sumatra, Guatemala and Mexico. The two new fossils described here represent two distinct species, Siphoniulus muelleri sp. nov. and S. preciosus sp. nov., and indicate a wider distribution of this order in the Cretaceous. The holotype of S. muelleri sp. nov. was well-enough preserved so that characters of the head, such as the incisura lateralis, not investigated before in extant representatives of the order, could be studied. This study highlights the possibilities and challenges provided by μCT technology in investigating Diplopoda fossilized in amber. 相似文献
13.
A remarkable new wasp of the superfamily Evanioidea (Apocrita: Evaniomorpha) is described and figured from a female preserved in mid-Cretaceous amber from the Hukawng Valley of northern Myanmar. Othniodellitha mantichora Engel and Huang, gen. et sp. nov., is characterized by its unique combination of primitive and derived features in mesosomal and metasomal construction and wing venation, along with its peculiarly blocky head with a prominent facial horn, clypeal projection, and massive, squared mandibles, among other traits. The genus is placed in the new family, Othniodellithidae Engel and Huang, fam. nov., and is among the lower Evanioidea and distinguished from the Praeaulacidae and other Mesozoic evanioid families. 相似文献
14.
Two new genera and two new species of fossil Throscidae: Potergosoma gratiosa gen. et sp. nov. and Rhomboaspis laticollis gen. et sp. nov. are described from the Lower Cretaceous Lebanese amber and are compared with extant and extinct genera. The described amber inclusions are the oldest known representatives of the family Throscidae. Some hypotheses on the phylogeny of the family Throscidae and the position of it in the superfamily Elateroidea are discussed. 相似文献
15.
A new species of Dicranoptycha Osten Sacken, 1860, Dicranoptycha plicativa sp. nov., and a unassigned species are illustrated and described from Myanmar (Burmese) amber (lowermost Cenomanian, Upper Cretaceous). A complete list of fossil species of the genus Dicranoptycha is summarized. In addition, an updated key to the known fossil species of the Dicranoptycha is provided. 相似文献
16.
Three new species within the stigmaphronid genus TagsmiphronEngel and Grimaldi, 2009, and one new species within the megaspilid genus ConostigmusDahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage. 相似文献
17.
18.
Cretocrenis burmanicus, gen. et sp. nov. is described from Burmese amber and represents the oldest known amber inclusion of the family Hydrophilidae (Coleoptera: Polyphaga). The new genus resembles the small-bodied modern genera of the tribe Anacaenini and the subfamily Acidocerinae, and the morphology of the venter and male genitalia suggest that it belongs to the Horelophus+Crenitis clade of the tribe Anacaenini. The fossil is described in detail, illustrated, and compared with extant hydrophilid genera. 相似文献
19.
The new tribe Taimyralticini trib. nov., new genus Taimyraltica gen. nov. and new species Taimyraltica calcarata sp. nov. (Coleoptera: Chrysomelidae: Galerucinae) are described from Upper Cretaceous (Santonian) Taimyr amber of northern Siberia (Yantardakh locality). The new genus shares characters with genera of the tribe Alticini (small size, transverse antebasal groove on pronotum, regular punctate-striate elytra) and the Galerucini (narrow, non-swollen metafemora). A possible apomorphy of the new genus and tribe is a large and acute metatibial spur, erect and oriented transversely to the longitudinal axis of tibia. This character state is unknown in recent and fossil genera of Galerucinae. Early evolution and divergence of leaf beetles are discussed, and a Jurassic and Early Cretaceous subfamily radiation is proposed. Fossil galerucines are reviewed. The oldest Phyllotreta is recorded from the lower Miocene of Izarra (Spain). Our findings show an extreme rarity of Phytophaga in Taimyr amber from Yantardakh. Absence of Phytophaga was recorded earlier at Obeschayuschiy (Santonian-Campanian of the Magadan Region) and compared with their much larger share in Khetana (north of Khabarovsk Krai); we find nearly the same differences between Taimyr amber from Yantardakh and Canadian amber. In Obeschayuschiy this appears due to the Mesophytic character of the flora. It is assumed that the Santonian amber gymnosperm forest of the Khatanga Basin had the same character. 相似文献
20.
Mesozoic leiodids are poorly known, and only one definitive leiodid is formally described from Burmese amber. Here we describe and illustrate the second definitive Mesozoic leiodid, Cretagyrtodes glabratus gen. et sp. nov., based on a single specimen from the Upper Cretaceous Burmese amber. The fossil is placed in Agyrtodini (subfamily Camiarinae) after maxillary palpomere 4 as wide as palpomere 3, and procoxal cavities closed behind. Cretagyrtodes is tentatively attributed to the extant “Eupelates group”. The discovery of Cretagyrtodes in Burmese amber suggests that the south hemisphere endemic tribe Agyrtodini is probably an ancient group, which has showed its first appearance before the breakup of Pangaea. 相似文献