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1.
The Glandulariini (=Cyrtoscydmini) is today the largest, most species-rich and most diverse tribe of Scydmaeninae comprising over 70% of the extant species of this subfamily. Named genera and species of Glandulariini are known mostly from Miocene to Eocene ambers, with only one, recently described Mesozoic taxon. Here we report the second genus of Glandulariini from Upper Cretaceous Burmese amber, Cenomaniola Jałoszyński and Yamamoto, gen. nov., with two species, C. carinata Jałoszyński and Yamamoto, sp. nov. and C. macrophthalma Jałoszyński and Yamamoto, sp. nov. Cenomaniola shows the general body plan typical of the ‘Euconnus complex’ within Glandulariini, with thick bristles on the sides of head and pronotum, a character conserved for nearly a hundred million years. With the previous discovery of Scydmobisetia Jałoszyński and Yamamoto, two major body forms typical of the extant Glandulariini are already proved to have differentiated in or before the Late Cretaceous. This demonstrates early origins and a long conservation of ‘Euconnus-like’ and ‘Sciacharis/Horaeomorphus-like’ body plan in the currently largest group among Scydmaeninae.  相似文献   

2.
The Eutheiini includes over 90 extant species classified in seven genera and distributed predominantly in the Northern Hemisphere. So far only one extinct genus and species unambiguously placed in this tribe has been known, Archeutheia, from Albian of Spain. We report the discovery of Eutheia, a member of the largest extant genus of Eutheiini, in Santonian of northern Siberia. Extant species of Eutheia are primarily defined on the basis of male genital characters, and the specimen discovered in Taimyr amber is a female; consequently it is described as Eutheia sp. The new finding remarkably extends the known range of Eutheiini during Cretaceous over the area of about 6 thousand kilometers, from the Iberian Plate to northern Siberia. A long morphological stasis in Eutheia suggests that this genus was associated with stable mesic microhabitats of the upper soil layers or rotten wood for at least 83 my.  相似文献   

3.
The taxonomic status of previously misplaced species of an ant-like stone beetle from Spanish amber (Albian) is clarified. Specimens of Kachinus magnificus (originally placed as incertae sedis within Scydmaenitae) were re-examined and their characters were found different from those of the type species of Kachinus from Myanmar amber. Consequently, Archeutheia gen. nov. is proposed to accommodate the Spanish species, resulting in Archeutheia magnifica comb. nov. Moreover, in a comprehensive phylogenetic analysis Archeutheia was placed as a sister group to extant Eutheia, within the tribe Eutheiini of Cephenniitae. A specimen of an undetermined genus from Myanmar amber clearly belonging in Cephenniini is also recorded. Both taxa represent the first definite Cretaceous Cephenniitae, a supertribe whose recent members are distributed on all continents but are especially diverse in the Oriental and East Palaearctic regions. This finding demonstrates a diversification of two presently most species-rich tribes of Cephenniitae already in Early Cretaceous. Archeutheia is strikingly similar to the extant species of Eutheia and Veraphis, showing male dimorphic characters (modified protrochanters) and antennal cavities characteristic of Palaearctic Eutheiini. This fact suggests a long morphological stability in the supertribe. A previously proposed hypothesis of an early presence of the Cephenniitae in the Northern Hemisphere is for the first time supported by the fossil record, and the palaeolocalities (western Europe and Southeast Asia) demonstrate a wide distribution of Cephenniitae in the Cretaceous and an early split of its ancestral lineage.  相似文献   

4.
The Mastigitae is a small supertribe of ant-like stone beetles that currently includes nine extant and five extinct genera. Extinct taxa are known within tribes Clidicini and Mastigini; the latter with one genus discovered in Baltic amber. For the first time, a Mesozoic genus of the tribe Mastigini is described, Clidicostigus arachnipes Jałoszyński, Brunke and Bai, gen. et sp. nov., from Cenomanian Burmese amber. The new taxon shares an enlarged and spiny scape and pedicel with its extant relatives but has deep elytral grooves and a strongly elongate and asymmetrical maxillary palpomere IV; the beetle has also enormously elongate maxillary palps and legs. The elongation of appendages in combination with only moderately large eyes is postulated to have evolved as an adaptation to running quickly, with tactile and chemical senses predominating over sight. The mode of life of Clidicostigus might have been similar to that of extant Mastigini, especially of particularly slender South African species that run on the ground and climb bushes and trees in search of prey.  相似文献   

5.
The new tribe Mediumiugamiini (Coleoptera: Polyphaga: Tenebrionoidea: Mordellidae) is described based on Mediumiuga sinespinis gen. et sp. nov. It is a fossil beetle from Albian (Early Cretaceous) amber from the Peñacerrada I outcrop (Spain). It is the first Spanish beetle described in amber. The mesotibiae and mesotarsi bearing multiple dorsal–lateral ridges, running oblique, metatibiae without any dorsal or dorsal–lateral ridge, only showing a subapical ridge, and metatibiae without apical spurs, define the new tribe. A key for worldwide tribes of Mordellinae, including Mediumiugamiini, is provided. Evolution of some characters of Mordellidae along Cretaceous is discussed.  相似文献   

6.
7.
A recently discovered articulated partial skeleton of Ornithomimus from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada is remarkable in the extent and quality of preservation of integumentary structures including feathers. It is the first ornithomimid to preserve a tail bearing extensive plumaceous feathers that are slightly more elongate in comparison to those present on the remainder of the body. However, the underside of the tail and the hind limb distal to the middle of the femur appear devoid of plumage. Overall, the plumage pattern in Ornithomimus is similar to that of Struthio camelus (ostrich) and other large palaeognaths, indicating a probable function in thermoregulation. The specimen also preserves the body outline around the legs, including a skin contour anterior to the femur, analogous to skin webs in extant birds. Whereas the knee web of birds bridges the knee to the abdomen, in Ornithomimus it spans from the mid-femoral shaft to the abdomen, and is herein referred to as an anterior femoral web. This is the first report of such soft tissue structures in non-avian theropods. It may indicate that the resting position of the femur was positioned more anteroventrally in ornithomimids than in most theropods, and in that sense may have been transitional to the situation in modern birds.  相似文献   

8.
The enigmatic staphylinid subfamily Dasycerinae consists of only 17 species within a single extant genus Dasycerus, but it is easily distinguished from other rove beetles by overall, latridiid beetle-like body and extremely slender, verticillate antennae. Direct fossil evidence is lacking for this group. Here, I describe the first fossil of this peculiar subfamily, Protodasycerus aenigmaticus gen. and sp. n., from the Upper Cretaceous Burmese amber. Despite considerable external similarities to extant dasycerine species, this new genus possesses several important morphological differences: a much smaller body size (ca. 1.1 mm), narrowly elongate antennomeres 1 and 2, not strongly transverse metaventrite, elytra striate but not tricostate, and truncate posterior margin of the elytra exposing four abdominal tergites dorsally. The new finding of a Cretaceous representative of the Dasycerinae helps elucidate the origin and early evolution of the omaliine group subfamilies, implying rather close similarities between the Dasycerinae and the monobasic subfamily Neophoninae. Discovery of P. aenigmaticus suggests a rather ancient origin of the subfamily by the Upper Cretaceous.  相似文献   

9.
Mesozoic whip-scorpions are very rare, with only two Cretaceous species known to date. Here we describe a new genus and species of Thelyphonidae, Mesothelyphonus parvus gen. & sp. nov., based on a very well-preserved male in Upper Cretaceous amber from Myanmar. Mesothelyphonus is firmly placed in the extant subfamily Thelyphoninae as supported by the abdominal tergites with a median longitudinal suture. Mesothelyphonus differs from other fossil and recent genera primarily by its very small body size, the absence of ommatoids on abdominal segment XII, and the elongate, slender and toothed patellar apophysis of the male pedipalp. The new discovery represents the oldest definitive fossil record for Thelyphoninae, highlighting the antiquity of the whip-scorpion group.  相似文献   

10.
Rapid radiation of the Bittacidae during the Jurassic resulted in high diversity within this family of Mecoptera. More than 40 species within 23 genera have been described from this period. However, around the end of the Jurassic the abundance and diversity of Bittacidae decreased and only seven species in five genera are known to date from the Cretaceous. A new specimen from the basal Cretaceous of England, Tytthobittacus jarzembowski sp. nov., described here, represents the eighth species and the second fossil representative of this family from the European Cretaceous. This hangingfly belongs to an extant and relictual genus previously known only from Australia. The paper also includes a review of all known Cretaceous bittacids and a re-examination of European species Antiquanabittacus nanus Petrulevičius and Jarzembowski, 2004.  相似文献   

11.
Here we describe an isolated tooth of a metriorhynchid crocodylomorph from the Hybla Formation (Aptian, Lower Cretaceous) of Rocca Chi Parra quarry (Montagna Grande, Calatafimi, Trapani Province), Sicily, Italy. The specimen shares with the Upper Jurassic taxon Plesiosuchus manselii a mediolaterally compressed conical tooth crown, noticeable lingual curvature, mesial and distal carinae with microscopic, rectangular contiguous denticles, strong distal curvature of the mesial margin, and the presence of weak 'carinal flanges' on the labial and lingual surfaces (which are preeminent at the mid-crown). This suite of morphologies is also present in an unnamed Valanginian (Lower Cretaceous) plesiosuchinan from France. However, the Sicilian tooth differs from these taxa in having more pronounced carinae, and faint apicobasally aligned enamel ridges. It also differs from P. manselii in having more extensive 'carinal flanges' on the labial surface. The specimen extends the known geological range of Metriorhynchidae and Thalattosuchia by approximately 7–8 million years. This overturns previous hypotheses of Metriorhynchidae becoming extinct early in the Early Cretaceous.  相似文献   

12.
An inclusion in Cretaceous amber from Myanmar is described as a new fossil species in the extant liverwort genus Frullania. The name Frullania pinnata is proposed for the taxon that is characterized by entire underleaves paired with a conspicuous pinnate branching pattern, two distinctive and stable morphological features. Entire underleaves are known in several extant Frullania species but had not previously been documented in Frullania in amber. The combination of morphological characters in this new fossil species is unknown in any crown group lineage of Frullania, and may very well represent a stem lineage element of the genus. This discovery is important because it expands our understanding of the diversity of Frullaniaceae in the Burmese amber forest as well as important ramifications for the phylogenetic reconstruction of extant Frullania lineages.  相似文献   

13.
A new genus and species, Prajna tianmiaoae gen. & sp. nov., is described and figured based on a well-preserved individual in Cretaceous amber from northern Myanmar. Prajna is definitely placed in the extant tribe Thinobiini Sahlberg of the staphylinid subfamily Oxytelinae Fleming based on its mostly convergent gular sutures, distinctly delimited and constricted neck, subulate maxillary palpi 4, stylus-bearing sternite IX in females, closely situated mesocoxae, spineless tibiae, 3-3-3 tarsal formula with strongly compressed basal two tarsomeres and with lobes on tarsomere 2, and typical Thinobiini-like pubescence. Prajna is separated from other allied genera by the depressions on the head, complete epistomal suture, slender maxillary palpomere 3, subulate palpomere 4, and tri-sulcate pronotum. The new genus combines some key characters of both Oxytelini and Thinobiini, indicating the close relatedness of the two tribes.  相似文献   

14.
A new species belonging to the extant dermestid genus Attagenus, Attagenus burmiticus sp. nov., is described and illustrated based on a well-preserved specimen from the Upper Cretaceous Burmese amber. The discovery suggests that Attagenus is an ancient group, originating as early as in the mid-Cretaceous. Along with another species of Attagenus known from the Upper Cretaceous New Jersey amber, it implies that Attagenus were widespread in the Mesoozic.  相似文献   

15.
A new fossil soldier beetle Myamalycocerus vitalii gen. et sp. nov., is described and illustrated from an inclusion in Upper Cretaceous Burmese amber (Myanmar). It differs from all extant genera that have elytra adorned with small striae by possessing raised punctation, very probably an ancient character as it is absent among current species. It differs from the fossil Ornatomalthinus Poinar et Fanti, by the long elytra and relief points which are less raised and more numerous. This new genus appears vaguely related (not necessarily phylogenetically) to the current genus Lycocerus Gorham.  相似文献   

16.
The Cretaceous new species and genus Albocryptophagus cantabricus gen. et sp. n. is described based on a fossil specimen from the El Soplao amber deposit (Spain). The new genus is similar to the extant genera Cryptophagus and Micrambe, but differs from them in the transverse pedicel, anterior angles of pronotum, pronotal margin unmodified, pronotal pits absent. Albocryptophagus gen. n. is undoubtedly the most ancient representative for the subfamily Cryptophaginae described up today. Because the saproxylic habits in recent species of the family, a similar behavior is inferred for this new fossil. It is a new example that agrees with the idea that fossil beetles from Mesozoic ambers are characterized by a saproxylic lifestyle.  相似文献   

17.
A new polypore fungus beetle is described and figured from an individual preserved in Upper Cretaceous (Cenomanian, ca. 99 Ma) amber from northern Myanmar. Cretosynstrophus archaicus gen. et sp. nov. is confidently placed in the extant subfamily Eustrophinae based on its elongate oval body and pronotum with two basal, sublinear impressions. It shares several characters belonging to two Recent tribes (Eustrophini and Holostrophini), but it cannot be attributed to either of them. Together with other tetratomid genera from the Cretaceous, the new discovery implies that the Recent small family Tetratomidae is much more diverse and more widespread than previously documented. In addition, a morphological similarity between Cretosynstrophus and extant Synstrophus suggests a similar fungi feeding habit for Cretosynstrophus, highlighting an ancient association between tetratomid beetles and fungi in the Mesozoic.  相似文献   

18.
Sclerogibbid wasps are obligate parasitoids of webspinners (Embiodea). Both groups have a particularly scarce geological record and are known since the Cretaceous: there are only four species of webspinners known from Burmese amber, and only two sclerogibbids were described from Barremian Lebanese and Cenomanian Burmese ambers. Here we report transferred genus from Aptian Choshi (Japan) amber and new sclerogibbids from Cenomanian Burmese and Charentese (France) ambers. The taxa described from Burmese amber are: Burmasclerogibba aptera gen. et sp. nov., Cretosclerogibba gen. nov. (with C. antennalis sp. nov., C. contractocollis sp. nov., C. neli sp. nov. and C. rasnitsyni sp. nov.) and Edrossia vetusta gen. et sp. nov. The first European fossil sclerogibbid Gallosclerogibba alnensis gen. et sp. nov. is described from Charentese amber. The holotype of Chosia yamadai Fujiyama, from Choshi amber, is re-described; it appears to be the oldest Laurasian sclerogibbid. The significant abundance and variety of Burmese sclerogibbid wasps (60% of fossil species known worldwide), as proxy of their hosts, were probably caused by the protection granted to them by the silk webs and possibly by the limited predation from ornithuromorph birds or crown-group ants. While all three extant sclerogibbid genera have apterous females, genera with winged females (Cretosclerogibba and Edrossia) dominated in Burmese amber. Small silk galleries from hosts may have favored the preservation of wings in females of Cretaceous sclerogibbids. Most new species described in the present paper, in addition to C. yamadai, are characterized by a very slender neck and a very long frontal process concealing the antennal toruli. These characters disappeared in extant species. We suggest that this loss was caused by a change in the fauna of predators, penalizing species with long neck and rostrum.  相似文献   

19.
A new genus, Cretaproscolia, and three new species of scoliid wasps, Archaeoscolia hispanica, Cretoscolia montsecana and Cretaproscolia josai, are described from the Lower Cretaceous of Spain and Brazil. The species representing the new genus is attributed to the plesiomorphic extant subfamily Proscoliinae, while the two other new species are assigned to two previously described genera in the extinct, archaic subfamily Archaeoscoliinae. The Brazilian species is the first Mesozoic scoliidid to have been described from the New World.  相似文献   

20.
New material of pterasterid asteroids from the UK chalk is described on the basis of ossicles recovered from washed residues. A new species, Pteraster lyddenensis sp. nov., is erected for oral and adambulacral ossicles and a primary radial ossicle from the Cenomanian Grey Chalk Subgroup of Dover (Kent), and the first UK record of Pteraster kutscheri Gale, 2022 is described from the upper Campanian Chalk of Norwich (Norfolk); both taxa belong to extant groups of Pteraster. Pteraster lyddenensis sp. nov. is the oldest known representative of the genus. The benthic invertebrate fauna of the Cretaceous chalk facies includes a number of extant genera which at the present day dwell in the deep sea. However, their presence was probably due to the low-productivity oceanic palaeoenvironment of the Chalk Sea, simulating deep-ocean conditions, rather than its depth.  相似文献   

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