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1.
The Gurpi section in western Shiraz, faulted Zagros range of southwestern Iran, contains one of the most complete Early Santonian to Late Maastrichtian sequences. The lack of a good fundamental paleontological study is a strong motivation for investigating calcareous nannofossils in southwestern Iran. The Gurpi Formation is mainly made up of grey shale. As a result of this study, 23 genera and 47 species of nannofossils have been identified for the first time. This confirms the existence of biozones CC14–CC26, which suggests the age of Early Santonian to Late Maastrichtian. All Early Santonian to Late Maastrichtian calcareous nannofossil biozones from CC14 (equivalent to the Micula decussate Zone) to CC26 (equivalent to the Nephrolithus frequens Zone) are discussed. Additionally, the zonal subdivision of this section based on calcareous nannofossils, is correlated with planktonic foraminiferal zones (Dicarinella asymetrica Zone to Abathomphalus mayaroensis Zone). We can also learn about the predominant conditions of the studied sedimentary basin that was in fact a part of the Neotethys basin with the existence of index species of calcareous nannofossils indicating a warm climate and high depths of the basin in low latitudes.  相似文献   

2.
The first data on the distribution of calcareous nannofossils in the Behbehan section, the Kuh-e-Rish, are considered. According to the distribution of nannofossils, the Upper Cretaceous deposits of the section are subdivided into nine biostratigraphic zones. CC17 (Calculites obscurus zone) indicate the Late Santonian. Biozones CC18 (Aspidolithus parcus zone), CC19 (Calculites ovalis zone), CC20 (Ceratolithoides aculeus zone), CC21 (Quadrum sissinghii zone), and CC22 (Quadrum trifidum zone) represent the Campanian. Biozone CC23 (Tranolithus phacelosus zone) indicate the Late Campanian–Early Maastrichtian. Biozones CC24 (Reinhardtites levis zone) and CC25 (Arkhangelskiella cymbiformis zone) suggest the Middle and Late Maastrichtian, respectively. In the late Late Maastrichtian, due to decreasing in water depth at the study area, Nephrolithus frequens zone (CC26) defined in Tethysian domain was not recognized. The boundary between Gurpi–Pabdeh Formations represented a non-depositional period from the late Late Maastrichtian to the end of Early Paleocene. Also, it seems that predominant conditions of the sedimentary environment of Neotethys basin with the presence of index species calcareous nannofossils specified, which itself indicates that the warm climate and high depth of the basin in Late Santonian to Late Maastrichtian, in low latitudes has been prevalent.  相似文献   

3.
Calcareous nannoplankton biostratigraphy has been performed on five sedimentary sections through the marine Akveren Formation from the Bartin region of northern Turkey, on the southern Black Sea coast. This new biostratigraphy provides an age for the formation of the Early Campanian (nannofossil zone UC15aTP) to Early Selandian (nannofossil zone NP5), and highlights the presence of the Campanian/Maastrichtian, Cretaceous/Tertiary (K/T), and Danian/Selandian boundaries in this intermediate palaeolatitude location. Micula murus was identified below the K/T boundary, but Micula prinsii and Nephrolithus frequens were not, which implies that the K/T boundary interval is not complete in the study area. These dates are in agreement with previous micropaleontological studies.  相似文献   

4.
A biostratigraphical and palaeoecological survey employing calcareous nannofossils and planktonic and benthonic foraminifera has been carried out in four sections of hemipelagic marls and chalks of the Late Maastrichtian Abathomphalus mayaroensis Zone of eastern Sinai, in order to evaluate the mechanisms controlling the composition of the well preserved microfauna and nannoflora.The Abathomphalus mayaroensis Zone in eastern Sinai can be easily identified by the wide occurrence of the index fossil A. mayaroensis and can be further subdivided by the first occurrences of Plummerita reicheli (ex. P. hantkeninoides) and Micula prinsii. Microfossil abundances and lithologies are characterised by pronounced repetitive distribution patterns. These include low and high frequency fluctuations of the planktonic/benthonic (P/B) foraminiferal ratio, repetitive changes in the abundance of calcareous nannofossils and benthonic foraminifera, as well as the development of chalk-marl couplets and thinning upward chalk packets. both microfossil distribution patterns and the occurrence of rhythmites are attributed to changes in primary palaeoproductivity. Semiquantitative investigations of calcareous nannofossils and a few selected benthonic foraminifera yield evidence of the presence of high (HP) and low (LP) productivity assemblages.The interpreted HP assemblage is dominated by Glaukolithus diplogrammus, Manvitella pemmatoidea, Microrhabdulus decoratus and Micula murus and the benthonic foraminifera Neoflabellina jarvisi; the LP assemblage is characterised by Lithraphidites quadratus and Bolivinoides draco. However, further quantitative studies are necessary to reconstruct the exact composition of these assemblages and to explain deviatory developments. The chalk-marl couplets, thinning-upward chalk packets and the high frequency P/B patterns are interpreted to reflect productivity changes related to orbital forcing. These hemipelagites were deposited during the latest phase of the southern Tethyan upwelling system, which was active from the Santonian to the Late Maastrichtian with a peak in the Campanian. Termination of upwelling just before the K/T boundary also provides a good explanation for the change towards a palaeobathymetric control on foraminiferal distribution, as observed for the Palaeocene of central east Sinai.  相似文献   

5.
A moderately various calcareous nannofossil assemblage of 44 species assigned to 24 genera is identified in samples collected from the Shahdar section and 18 genera and 43 species in Namazgah section. Testing of calcareous nannofossil has permitted recognition of few important coccolith events in stratigraphic interval occupied by the uppermost layers of Cretaceous in the Shahdar and Namazgah sections. These events are correlated to the CC24–CC26 of Sissingh (Geol Minjbouw 56: 37–65, 1977) in two sections. According to these biozones, the age of the studied sections in Shahdar and Namazgah is Early Maastrichtian–Late Maastrichtian. On the basis of paleoecological interpretation, the last layers related to the Cretaceous deposits in two sections were deposited in shallow marine environment in relatively low latitude, and the depth of the basin from the Cretaceous deposits toward Fajan Formation is minimized.  相似文献   

6.
One hundred and thirty nine samples have been studied from the Late Campanian–Early Maastrichtian of three deep wells drilled in Jiza’-Qamar Basin, Eastern Yemen to determine the calcareous nannofossil zones and the age of the sediments. Forty-seven calcareous nannofossil species were identified and four biozones were determined in the present study (CC21–CC24). These biozones are assigned to the Late Campanian–Early Maastrichtian ages. Most of the studied species in this work refer to tropical–subtropical environment. The Campanian–Maastrichtian Boundary was determined in Al-Fatk well based on the last occurrence of Eiffelithus eximus and the last occurrences of Uniplanarius sissinghii and Uniplanarius trifidus.  相似文献   

7.
The Cretaceous outcrop belt of the Mississippi Embayment in the Gulf Coastal Plain (GCP) spans the Cretaceous/Paleogene (K/Pg) boundary. A detailed reconstruction of this time interval is critical for understanding the nature of biotic and environmental changes preceding the end-Cretaceous Mass Extinction event and for deciphering the likely extinction mechanism (i.e., bolide impact versus volcanism). Eight sections encompassing the K/Pg succession across the Mississippi Embayment were analyzed using biostratigraphic sampling of ammonites, dinoflagellates, and nannofossils. An upper Maastrichtian ammonite zonation is proposed as follows, from oldest to youngest: Discoscaphites conradi Zone, D. minardi Zone, and D. iris Zone. Our study documents that the ammonite zonation established in the Atlantic Coastal Plain (ACP) extends to the GCP. This zonation is integrated with nannofossil and dinoflagellate biostratigraphy to provide a framework to more accurately determine the age relationships in this region. We demonstrate that ammonites and dinoflagellates are more reliable stratigraphic indicators in this area than nannofossils because age-diagnostic nannofossils are not consistently present within the upper Maastrichtian in the GCP. This biostratigraphic framework has the potential to become a useful tool for correlation of strata both within the GCP and between the GCP, Western Interior, and ACP. The presence of the uppermost Maastrichtian ammonite D. iris, calcareous nannofossil Micula prinsii, and dinoflagellates Palynodinium grallator and Disphaerogena carposphaeropsis suggests that the K/Pg succession in the GCP is nearly complete. Consequently, the GCP is an excellent setting for investigating fine scale temporal changes across the K/Pg boundary and ultimately elucidating the mechanisms causing extinction.  相似文献   

8.
Forty-seven samples from Upper Cretaceous sections penetrated by the Kachi-1 and Inga-1 wells in the South Yellow Sea Basin have been analysed for their spore and pollen content. Thirty-five species of 18 spore genera and 54 species of 28 pollen genera are documented. One new monotypic genus, Diporocolpopollenites, and its type species, D. kachiensis sp. nov., are erected, and Dilwynites Harris, 1965, and its type species, D. granulatus Harris, 1965, are emended. There are also three new combinations: Ephedripites eocaenicus (Selling, 1944), E. praeclarus (Chlonova, 1961), and Retitricolpites anguloluminosus (Anderson, 1960). Two palynological zones are erected: anAquilapollenites attenuatus Assemblage Zone, which encompasses deposits that are considered to be latest Campanian–Early Maastrichtian in age, and an Aquilapollenites eurypteronus Assemblage Zone for sections that have been dated as Late Maastrichtian. The assemblages are typical of the Yenisey-Amur Subprovince of the Aquilapollenites (floral) Province. Lowland floodplain to shallow, commonly mesotrophic, lacustrine environments of deposition are indicated. The climate was probably wet subtropical, with rainfall being somewhat higher during the Late Maastrichtian than through the latest Campanian–Early Maastrichtian.  相似文献   

9.
《Cretaceous Research》2008,29(1):40-64
The proposed definition of the Turonian/Coniacian boundary, at the first occurrence of the inoceramid bivalve Cremnoceramus deformis erectus (Meek) (= Cremnoceramus rotundatus (sensu Tröger non Fiege)), prompted a rigorous study of the calcareous nannofossil events through this interval, both for calibration of the calcareous nannofossil biostratigraphy, and to provide an assessment of the suitability, in calcareous nannofossil terms, of the proposed stratotype section. New calcareous nannofossil data are presented here, detailing the biostratigraphy of the boundary interval from four locations. These include the candidate boundary-stratotype, the Salzgitter-Salder Quarry section (northern Germany), as well as the Slupia Nadbrzezna outcrop (central Poland), a potential secondary reference section. Also included is the Brezno Pd-1 Borehole and outcrops in the Brezno Formation (= Priesener Schichten) type-area (north-western Czech Republic), which represents an original boundary candidate (Copenhagen Stage Boundaries Meeting, 1983), and the Langdon Stairs coastal section (south-eastern England), part of the British Chalk succession. The calcareous nannofossil events derived from each section provide a sequence across the boundary of (in stratigraphical order): below the boundary, the first occurrence of Lithastrinus septenarius followed by that of Broinsonia parca expansa; above the boundary, the last occurrence of Helicolithus turonicus followed by the first occurrence of Micula staurophora (= Micula decussata of some authors). This places the boundary within Nannofossil Subzone UC9c. A similar sequence of events has previously been determined from sections in north-eastern England and in the south-eastern Indian Ocean. The presented data and correlations suggest that either the Salzgitter-Salder Quarry section or the Slupia Nadbrzezna outcrop section would make a suitable Global Stratotype Section for the Turonian/Coniacian boundary, as far as calcareous nannofossils are concerned. The use of the calcareous nannofossil Marthasterites furcatus, widely quoted as an indicator of this boundary, is discussed and proved to be untenable.  相似文献   

10.
New carbon (δ13C) isotope records calibrated by planktonic bioevents provide general support for a late Campanian age assignment of the Shiranish Formation (Fm.) and its boundaries in the Dokan section (NE Iraq). The Shiranish Fm. is characterised at the base by a mid-Campanian unconformity as can be interpreted by absences of nannofossil zones CC20-21. The Shiranish Fm. then spans nannofossil biozones CC22-CC23a (UC15d-eTP to UC16aTP). Results obtained on carbon isotopes suggest that diagenesis affected and compromised a few carbonate samples in the uppermost 50 m of the section. However, once these samples are discarded, pristine trends suggest that the top of the section records a negative carbon isotope excursion that is interpreted as CMBa-c events that straddle the Campanian–Maastrichtian boundary. This interpretation is supported by the lowermost occurrence of planktic foraminifers Rugoglobigerina scotti and Contusotruncana contusa some 30 m above the base of the negative excursion and 10 m below a positive excursion identified as the Maastrichtian M1+ event. Discrepancies in the stratigraphic range of several planktic foraminifer bioevents are highlighted and advocate for the need of many more integrated records of planktic foraminifer and nannofossil biostratigraphy alongside carbon isotope stratigraphy in the eastern Tethys in order to improve regional and global schemes.  相似文献   

11.
Benthic foraminifers from borehole sections recovered by drilling in the Yamal Peninsula, West Siberia, characterize the Ceratobulimina cretacea Beds (the upper Campanian-lower Maastrichtian) and the Spiroplectammina variabilis-Gaudryina rugosa spinulosa and Spiroplectammina kasanzevi-Bulimina rosenkrantzi regional zones of the lower and upper Maastrichtian, respectively. The Danian Stage is missing from the sections, which include marine deposits of the Selandian Stage attributed to the Ceratolamarckina tuberculata Beds. Foraminiferal assemblages of the beds include the Siberian endemic species associated with Paleocene foraminifers of the Midway-type fauna of subglobal distribution range. Occurrence of the latter suggests that warm-water surface currents from the North Atlantic reached southern areas of the Kara Sea.  相似文献   

12.
This study is based on calcerous nannofossil assemblages changes and fluctutions of stable carbon and oxygen isotopes was collected clayey limestones, limestones, and marls in the Maastrichtian to Selandian from Akveren Formation (Western Black Sea). As the relative abundances of species of Micula spp, Watznauera barnesiae, and Arkhangelskiella cymbiformis, which tolerated changes of temperature and nutrition, carbon and oxygen isotopes compositions, and low species richness imply strong diagenesis effect at the Maastrichtian, there is no important diagenesis effect at Paleocene. Just after the Cretaceous/Tertiary (K-Pg) boundary, Thoracosphaera spp. and Braarudosphaera bigelowi were dominant species; Danian is characterized by Thoracosphaera, Ericsonia ovalis, Cruciplacolithus spp., Coccolithus pelagicus, and Ericsonia subpertusa. Generally, the nutrition productivity is good–moderate in Lower Maastrichtian, and decreasing carbon isotope values during the Uppermost Maastrichtian shows the presence of oligotrophic environmental conditions suitable with global nutrition crises before the K-Pg boundary and diagenesis in study area. Throughout the Danian, mesotrophic–oligotrophic environmental conditions dominate; however, the decrease in nutrition before Selandian represents oligotrophic environmental conditions. The increasing nutrition at Selandian is related to the change in the environmental conditions.  相似文献   

13.
The Elk Butte Member of the Pierre Shale of southeast South Dakota and northeast Nebraska yields a late Maastrichtian cephalopod fauna of nautiloids, belemnites and ammonites of theFeletzkytes nebrascensisZone, best known from the near-shore facies of the Fox Hills Formation. ThenebrascensisZone is the highest distinct marine assemblage that can be recognised in the Western Interior, although ammonites occur as rarities high in the Lance Formation in Wyoming. Elements of the fauna occur in the Gulf Coast and Atlantic Seaboard, and extend into the highest Maastrichtian nannofossil Subzone CC26b, ofMicula prinsii, in Texas. These occurrences point to the existence of a southerly marine route for migration into and out of the northern Interior during the late late Maastrichtian. An analysis of Maastrichtian ammonite occurrences in West Greenland reveals no evidence for a marine link to the western Interior at this time, but rather indicates an open marine link to the North Atlantic region.The presence of upper upper Maastrichtian Pierre Shale in southeast South Dakota and northeast Nebraska, deposited in water depths that are conservatively estimated at 100-200 m, suggests that marine conditions (evidence for which has been removed by post-Cretceous erosion) may have extended well to the north of the shoreline position indicated in recent palaeogeographic reconstructions.  相似文献   

14.
Field investigations in the Upper Benue rift basin of Nigeria highlight the necessity for a review of the Cenomanian-Turonian biostratigraphy of the area. Three ammonite zones of Vascoceras bulbosum (Reyment), Paravascoceras costatum (Reyment) and Pseudotissotia (Bauchioceras) nigeriensis (Woods) have been recognized at sections exposed at the quarry of the cement factory near the village of Ashaka. The sediments within the Vascoceras bulbosum zone also contain specimens of Exogyra olisiponensis (Sharpe), Kanabiceras septemseriatum (Cragin) and Epengonoceras dumbli (Cragin). These are all indicative of an uppermost Cenomanian age and reveal that at least the lower portion of the Gongila Formation as exposed at Ashaka is faunistically equivalent to the Pindiga Formation as exposed in the type section at Pindiga.The shales in the Gombe region in which Libycoceras ismaeli (Zittel) was found are probably part of the Maastrichtian Fika shales outcropping extensively in the northern part of the area. The discovery of Cyclolithes nov. sp. within Gombe sandstones at Dakiti near Kumo confirm that they are at least partially marine in origin and Maastrichtian in age. This could be considered as an additional evidence in favour of the presence of marine waters in the Upper Benue during the last part of the Cretaceous.  相似文献   

15.
This study of the upper Maastrichtian to Danian sedimentary succession from the northern part of the Romanian Eastern Carpathians (Varniţa section) aims to establish an integrated biostratigraphy based on calcareous nannofossils, organic-walled dinoflagellate cysts (dinocysts) and foraminiferal assemblages, and to reconstruct the depositional environments of the interval. The stratigraphic record across the studied section is incomplete, considering that an approximately 16 m thick strata interval from the top of the Maastrichtian to lowermost Danian cannot be analyzed due to a landslide covering the outcrop. The upper Maastrichtian is marked by a succession of biostratigraphic events, such as the First Appearance Datum (FAD) of the nannoplankton taxon Nephrolithus frequens and FAD of the dinocyst species Deflandrea galeata and Disphaerogena carposphaeropsis, and the Last Appearance Datum (LAD) of Isabelidinium cooksoniae in the lower part of the section. These bioevents are followed by the LAD of the Dinogymnium spp. and Palynodinium grallator dinocyst markers in the top of the Maastrichtian deposits analyzed. In terms of foraminiferal biostratigraphy, the upper Maastrichtian Abathomphalus mayaroensis Zone is documented in the lower part of the studied section. Some bioevents, such as the bloom of the calcareous dinoflagellate genus Thoracosphaera and the FAD of the organic-walled dinocysts Damassadinium californicum, Senoniasphaera inornata, Xenicodinium lubricum and X. reticulatum suggest an early Danian age for the middle part of the section. From the Danian deposits in the Varniţa section, we describe a new organic-walled dinocyst species, Pentadinium darmirae sp. nov., which is until now the only species of the Pentadinium genus discovered in the Paleocene. The occurrence of the global Danian dinocyst marker Senoniasphaera inornata in the top of the section, suggests an age not younger than middle Danian (62.6 Ma) for the analyzed deposits.The palynofacies constituents, as well as the agglutinated foraminiferal morphogroups, used to reconstruct the depositional environments, show that the late Maastrichtian sediments were deposited in an outer shelf to distal (bathyal) environment, followed by a marine transgression during the Danian.  相似文献   

16.
The late Turonian to early Campanian calcareous nannofossil biostratigraphy of the Austrian Gosau Group is correlated with ammonite and planktonic foraminiferal zones. The standard Tethyan zonations for nannofossils and planktonic foraminifers are applied with only minor modifications. The basal marine sediments of the Gosau Group, bearing late Turonian-early Coniacian macrofossils, belong to the Marthasterites furcatus nannofossil Zone (CC13). The Micula decussata Zone (middle Coniacian to early Santonian) is combined with the Reinhardtites anthophorus Zone because of the rare occurrence of Renhardtites cf. R. anthophorus already in the Coniacian and taxonomic problems concerning the correct identification of this species. The Santonian-Campanian boundary lies within the Calculites obscures Zone (CCl7).  相似文献   

17.
The palaeogeographic setting of the studied Ain Medheker section represents an Early Campanian to Early Maastrichtian moderately deep carbonate shelf to distal ramp position with high rates of hemipelagic carbonate production, periodically triggered by mass-flow processes. Syndepositional extensional tectonic processes are confirmed to the Early Campanian. Planktonic foraminifera identified in thin sections and calcareous nannofossils allow the identification of the following biozones: Globotruncanita elevata, Contusotruncana plummerae (replacing former Globotruncana ventricosa Zone), Radotruncana calcarata, Globotruncana falsostuarti, and Gansserina gansseri. The following stable C-isotope events were identified: the Santonian/Campanian boundary Event, the Mid-Campanian Event, and the Late Campanian Event. Together with further four minor isotopic events, they allow for correlation between the western and eastern realms of Tunisia. Frequently occurring turbidites were studied in detail and discussed in comparison with contourites.  相似文献   

18.
Conodonts from the Cambrian–Ordovician transition at the Liangcun section in Xishui County, Guizhou and at the Huangcao section in Wulong County, Chongqing are examined for the first time. Both sections are located at the southeast margin of the Sichuan Basin. A total of 1367 specimens were recovered, representing 30 species and 15 genera. Based on the ranges of conodonts generalized from these two sections and another six sections previously studied in the same region, three conodont zones, Cordylodus proavus, Monocostodus sevierensis and Cordylodus angulatus zones are recognized. The index species of the Cambrian–Ordovician boundary at the global stratotype section and point (GSSP), Iapetognathus fluctivagus and its substitute in China Iapetognathus jilinensis are not observed in the study sections, therefore it is impossible to determine the Cambrian–Ordovician boundary exactly. However, it probably lies within the lower part of M. sevierensis zone (the upper part of the Loushanguan Group), correlating with the GSSP in Canada and the Dayangcha section in China. Chronological sequences of the FAD (First Appearance Datum) of C. angulatus, Chosonodina herfurthi and Rossodus manitouensis are not obvious in the study, so the C. angulatus zone here is correlated with zones defined by C. angulatus, Ch. herfurthi and R. manitouensis in the lower Yangtze Platform.  相似文献   

19.
The uppermost Cretaceous (upper Campanian-Maastrichtian) pelagic successions from the Malatya Basin (NW Malatya, eastern Anatolia) were studied by 688 samples, which were collected from five stratigraphic sections in the Hekimhan area. The pelagic deposits conformably overlie rudist bearing shallow-water limestones and are overlain conformably by Maastrichtian dolomites and unconformably by Paleocene-Eocene deposits, respectively.The pelagic successions in the Hekimhan area comprise the Kösehasan Formation at the base and the Zorbehan Formation at the top and reach up to 1100 m in thickness. The Kösehasan Formation rests over the neritic rudist-bearing limestones of the Güzelyurt Formation along a sharp contact and consists mainly of flysch-type sandstone-mudstone alternation with complete and partial Bouma sequences. The carbonate content of abundant planktonic foraminifera and nannoplankton-bearing 980-m-thick succession increases upwards and the formation passes gradually to the clayey limestones and marlstones of the Zorbehan Formation to the top. Occurrences of nannoplankton Lithraphidites quadratus Bramlette and Martini and Micula praemurus (Bukry) in the first beds of the Kösehasan Formation indicate that the age of the Kösehasan Formation and overlying Zorbehan Formation is of late Maasthrichtian. Another late Maastrichtian taxa Cribrosphaerella daniae Perch-Nielsen and Arkhangelskiella maastrichtiana Burnett are observed from the lowermost part of the succession. Maastrichtian planktonic foraminifera such as Contusotruncana walfischensis (Todd) and Globotruncanita pettersi (Gandolfi) were recorded through the successions. Although planktonic foraminifera are diverse and abundant particularly in the Kösehesan Formation, index late Maasthrichtian species were not encountered. Campanian and Santonian-Campanian planktonic foraminifera, e.g. Radotruncana calcarata (Cushman) and Globotruncanita elevata (Brotzen), obtained particularly from the lower part of the succession and calcareous nannofossils such as Broinsonia parca parca Bukry, Reinhardtites anthophorus (Deflanre) and Eiffellithus eximius (Stover) are interpreted as reworked from older strata. Trace fossils are common throughout the succession.Rareness of planktonic foraminifera and nannoplankton in the uppermost part of the succession (Zorbehan Formation) indicates maximum shallowing of the latest Maastrichtian sea in this part of the basin. Rare echinoids, bivalves and ammonites are observed in that part of the sequence.The obtained data indicate that sediment accumulation rate of the pelagic deposits is rather high and about 27.5 cm/ky for this part of the basin. Changes in thickness of the formations along short distances in the five stratigraphic sections analysed in this study should be related to the diachroneity of the depositional and erosional events.  相似文献   

20.
In the Izeh section (Zagros Basin, SW Iran), about two third of the planktonic foraminiferal species, representing less than 20% of the individuals, present in the Maastrichtian polytaxic assemblages, became suddenly extinct at the K/Pg boundary. This mass species extinction testifies end-Cretaceous catastrophic bioevent. The Cretaceous species remain became apparently extinct gradually during the earliest Danian. In fact the unkeeled globotruncanids (Globotruncanella and Rugoglobigerina) as well as the small heterohelicids (Heterohelix and Pseudoguembelina) were also affected by extinction and they were reworked in the basal Danian. Except guembelitriids which persisted longer time. The sudden species extinctions were selective, eliminating geographically restricted large, complex, and deeper dwelling forms (i.e., globotruncanids and large heterohelicids). Contrary, few cosmopolitan small, simple surface-subsurface dwellers (i.e., guembelitriids among small heterohelicids and hedbergellids) crossed the K/Pg boundary and survived the catastrophic event. This selective bioevent is related to the end-Cretaceous bolide and produced ejecta impact inducing major environmental changes as decrease in temperature, collapse of nutriment, and flow breakdown in the water mass stratification. Our geochemical results (δ13C negative shift, δ 18O positive shift, and trace elements anomalies) are consistent with the cooling due to the collision winter triggering collapse of nutriment. Especially the δ 18O values across the upper Maastrichtian and the K-Pg transition in Izeh section although they are lower; they report similar fluctuations to those in El Kef section. Thus these lower values may be indicative of somewhat warmer water at the sea floor. Besides, like as elsewhere, in Izeh, this Cretaceous end catastrophic bioevent is followed by a delayed gradual recovery. Thus, immediately after the K/Pg boundary was created a nearly empty niche in which persisted few Cretaceous survivors with dwarf individuals. These survivors despite they suffered very stressful environmental conditions, they constituted the root stock of globigerinids from which the Cenozoic planktonic foraminiferal species evolved and developed carnivorous food diet.  相似文献   

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