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1.
Coastal and estuarine systems provide nursery grounds for many marine fish species. Their productivity has been correlated with terrigeneous inputs entering the coastal–estuarine benthic food web, thereby favouring the establishment of fish juveniles. Studies in these ecosystems often describe the nursery as a single large habitat without verifying nor considering the presence of contiguous habitats. Our study aimed at identifying different habitats based on macrozoobenthic communities and morpho-sedimentary characteristics and assessing the trophic interactions between fish juveniles and their benthic preys within these habitats. It included 43 sampling sites covering 5 habitats in which we described taxonomically and quantitatively the invertebrates and fish communities with stable isotopes and gut contents. It suggested that the benthic common sole Solea solea displayed feeding plasticity at the population level, separating the juveniles (G0) from the older fish (G1) into different “feeding sub-populations”. Size-based feeding plasticity was also observable in the spatial occupancy of that species in the studied bay. The demersal pouting, Trisopterus luscus, equally used the different habitats but displayed low feeding plasticity across and inside each habitat. Stable isotopes proved to be powerful tools to study the spatial distribution of trophic interactions in complex ecosystems like the bay of Vilaine and to define optimal habitats for fish that use the coastal–estuarine ecosystem as nursery grounds.  相似文献   

2.
It is known that the structure of benthic macrophyte and invertebrate habitats indicate the quality of coastal water. Thus, a large-scale analysis of the spatial patterns of coastal marine habitats makes it possible to adequately estimate the status of valuable coastal marine habitats, provide better evidence for environmental changes, and describe the processes behind the changes. Knowing the spatial distribution of benthic habitats is also important from the coastal management point of view. Our previous results clearly demonstrated that remote sensing methods can be used to map water depth and distribution of taxonomic groups of benthic algae (e.g., red, green, and brown algae) in the optically complex coastal waters of the Baltic Sea. We have as well shown that benthic habitat mapping should be done at high spatial resolution owing to the small-scale heterogeneity of such habitats in Estonian coastal waters. Here we tested the capability of high spatial resolution hyperspectral airborne image in its application for mapping benthic habitats. A big challenge is to define appropriate mapping classes that are also meaningful from the ecological point of view. In this study two benthic habitat classification schemes??broader level and finer level??were defined for the study area. The broader level classes were relatively well classified, but discrimination among the units of the finer classification scheme posed a considerable challenge and required a careful approach. Benthic habitat classification provided the highest accuracy in the case of the Spectral Angle Mapper classification method applied to a radiometrically corrected image. Further processing levels, such as spatial filtering and glint correction, decreased the classification accuracy.  相似文献   

3.
Post-settlement processes are a major focus in the study of the dynamics of marine populations and communities. Post-settlement movement of juveniles is an important, but often ignored, process which affects local predator–prey and competitive interactions. We used benthic suction sampling and pitfall traps to examine density and locomotory activity of Carcinus maenas juveniles in different intertidal habitat types in the Rio Mira Estuary, Portugal, to better understand intra-specific interactions in a system where density-dependent processes are known to regulate population dynamics. As expected, significantly higher densities of juvenile shore crabs were found from bare mud compared to densely vegetated habitats. At the time of sampling, small and intermediate stages together outnumbered by far the larger juveniles. Conversely, larger crabs were much more frequent than smaller ones in traps. A locomotory index (LI), i.e. the ratio between crab catch in pitfall traps and their density within their moving range, is proposed as a measure of movement. LI analyses indicated that: (1) movement is an order of magnitude higher in large than small juveniles and much higher in sparse than dense vegetation cover; (2) activity of small juveniles is mostly crepuscular, regardless of vegetation cover; and (3) movement of large juveniles is very limited in dense Zostera patches, but very high in sparsely vegetated areas, during the day and night. These results suggest that small juveniles are relatively protected under dense vegetation cover due to lower mobility of larger crabs, and provide evidence of temporal segregation of activity windows between juvenile crabs of different sizes, which may be a key mechanism to reduce cannibalism and therefore increase the carrying capacity of nursery habitats.  相似文献   

4.
The distribution of nepheloid layers across the outer shelf and upper continental slope off Namibia was studied during a cruise with R.V. Meteor in late austral summer 2003. Optical measurements, carried out with a transmissometer and a backscattering fluorometer, are correlated with suspended particulate matter (SPM) and particulate organic carbon (POC) values from water sample filtration. Conductivity-temperature-depth and oxygen data are used to relate the nepheloid layers to hydrographic structures. The particle content of surface water at the continental slope is controlled primarily by the offshore extension of highly productive upwelling filaments. A pronounced bottom nepheloid layer (BNL) covers the entire area of study with maximum intensity above the outer shelf and at the shelf break—an area where erosional forces dominate. The detachment of this BNL at the shelf break feeds a major intermediate nepheloid layer (INL) at 25.5°S. This INL is positioned at 250–400 m depth, at the lower boundary of an oxygen minimum zone, and is likely connected to the poleward flow of South Atlantic Central Water (SACW) across the shelf break. Together, these strong subsurface nepheloid layers are indicators of intensive lateral particle transport from the outer shelf towards a depocenter of organic matter on the upper continental slope.  相似文献   

5.
Tropical shallow-water habitats such as mangroves and seagrass beds are widely acknowledged as important juvenile habitats for various coral reef fish species, most of which are commercially important to fisheries. Spatio-temporal variability in ontogenetic habitat use by fish among these tropical coastal ecosystems has rarely been investigated, yet there are sufficient reasons to believe that this plays an important role. In the present study, we test the spatio-temporal variability in patterns of ontogenetic habitat use by some mangrove/seagrass-associated coral reef fishes (Lethrinus harak, Lethrinus lentjan, Lutjanus fulviflamma and Siganus sutor). Abundances of these four species were investigated during two years in Tanzanian coastal waters, using underwater visual census in mangrove, seagrass, shallow and deep mudflat, and shallow and deep coral reef habitats. The study covered four distinct seasons of the year and was done at two spatially separated (>40 km) locations. Averaged across locations, seasons and years, juveniles (≤10 cm length) of the four study species had significantly higher relative densities in shallow-water (mangroves and seagrass beds) than in deep-water habitats (deep mudflats or coral reefs), whereas the opposite pattern was found for the adults (>15 cm). These findings suggest a strong and general pattern of ontogenetic habitat shifts from shallow- to deep-water habitats. However, specific habitat-use patterns of juveniles as well as adults differed significantly in time and space. Various species showed subtle to considerable flexibility in juvenile as well as adult habitat use across seasons, years, or at different locations. Furthermore, for some species the data suggest presence of ontogenetic habitat shifts at one location but lack thereof at the other location. In summary, ontogenetic habitat use needs to be considered at various spatial and temporal scales for the interpretation of habitat utilization by fish during different life stages. This is important for conservation and management of these habitats, as essential habitats or seasons may be ignored or over-emphasized with respect to their importance for fish during different parts of their life cycle.  相似文献   

6.
7.
Optical transmissometer measurements were coupled with particulate organic matter (POM) observations to understand suspended sediment composition and distribution in the eastern Cariaco Basin during the rainy seasons of September 2003 and 2006. Our results suggest that nepheloid layers originating at the mouth of small mountainous rivers discharging into the eastern Basin are a major delivery mechanism of terrigenous sediments to the Basin interior. Intermediate nepheloid layers (INL) were observed near the shelf break (~100 m) and appear to effectively transport terrigenous material laterally from the shelf to deep waters, thereby providing a plausible supply mechanism of the terrestrial material observed in sediment traps. These findings highlight the importance of small, local rivers in the Cariaco Basin as sources of terrestrial material. In contrast, these nepheloid layers contained only limited POM. When this information is combined with published sediment trap POM data, it suggests that nepheloid layers may not be a primary mechanism for delivering terrigenous POM to the deeper waters of the basin during the rainy season. Rather, BNL may redistribute marine-derived POM from shallow waters to the Basin's interior by providing ballast materials, particularly during episodic events driven by wind and precipitation. Though we have determined that nepheloid layers play an important role in the seaward transport of particulate material in the Cariaco Basin, their composition and temporal variability have not been fully characterized. This is critical to understand lateral particle transport, since nepheloid layers constitute a significant source of sediment to the deep Cariaco Basin.  相似文献   

8.
太平洋悬浮体特征及近底雾状层(雾浊层)探讨   总被引:2,自引:0,他引:2       下载免费PDF全文
2005年在太平洋12个测站分层采集了悬浮体水样及连续的水体温度、盐度、深度等数据.利用水体悬浮体含量在垂向上的分布特征初步分析了大洋雾状层的特点,并讨论了水体中悬浮体颗粒物质的来源,对不同海洋环境中悬浮体分布进行了对比,发现大洋水体中悬浮体含量总体很小,一般不超过0.30mg/dm3,但在一定水层内悬浮体含量相对较高,形成雾状层.雾状层在大洋内广泛分布,一般以表层雾状层和近底雾状层形式存在.大洋水体中悬浮体颗粒物来源广泛.表层雾状层的形成主要受透光层内生物作用的影响,而底质沉积物的再悬浮是影响底部雾状层形成的重要因素.因受水体物理海洋环境和物质来源的影响,不同区域水体中悬浮体含量不同,雾状层的分布不同.海山顶部受水流及岩石基底影响雾状层分布不明显,海山底部与开辟区洋盆海域雾状层更普遍.开辟区东西区由于物源差异水体中悬浮体含量差别明显,靠近火山喷口的西区水体中悬浮体含量明显高于东区的.  相似文献   

9.
A challenge for marine ecologists is to explain distinct and recurrent patterns in the distribution of marine faunas by developing new methods that identify and link environmental processes responsible for these patterns. Methods that describe and predict the distribution of benthic faunas using single factors such as sediment type or water depth are generally inadequate, particularly when applied on a broad scale. When a combination of factors such as near-bed tidal velocity, surface seawater temperature and salinity are evaluated in conjunction with sediment type and depth, however, they more clearly characterise benthic habitats. Using principal component analysis (PCA) patterns in the distribution and abundance of different echinoderm and crustacean species were shown to be predictable and characterised by a suite of physical factors. Characterising benthic habitats using factors from the environment provided a potential mechanism for predicting patterns in their spatial distribution. A new analytical method for characterising a species habitat was constructed using a combination of PCA and a generalised additive model. The method is able to predict the habitat preferences of individual species based on their association with physical factors characterising their habitat. These preferences were then used to describe the probability of a species occurring across a range of different habitats, which is referred to as the habitat-envelope. This method enables one species habitat range to be compared directly to another. The strong correlation between species patchiness and its habitat-envelope was used to develop an index to identify species that are potentially more sensitive to habitat change. Distinct patterns in the habitat preferences of echinoderms were generally stronger than those identified for crustaceans. Thus, crustaceans were found more likely to exploit a wider range of habitats than echinoderms, suggesting that they may be less sensitive to habitat change.  相似文献   

10.
The gastropod Terebralia palustris often dominates the surface of muddy to sandy substrates of intertidal mudflats and mangrove forests, where they clearly destabilize the sediment. In the present study, it was investigated whether and to what extent the behaviour of juvenile and adult snails differs among habitats (mudflat vs. mangrove stand) in a Sonneratia alba mangal at Gazi Bay, Kenya. For this purpose we: (1) examined their distribution along three land–sea transects; and (2) applied stable isotope analysis to determine the feeding patterns of different-sized snails from the mangrove and mudflat habitats. Additionally, we investigated if these gastropods exert an impact on microphytobenthic (diatom) biomass, and whether this is size-dependent. The latter objective was met by either enclosing or excluding different-sized snails from experimental cages on the intertidal mudflat and the subsequent assessment of a change in pigment concentration of the sediment surface. In agreement with several previous studies conducted in other mangroves and geographical locations, a spatial segregation was demonstrated between juveniles (more common on the mudflat) and adults (more common in the mangrove forest). On the intertidal mudflat juveniles avoided sediment patches characterized by highly saline water in intertidal pools and a high mud content, while adults tended to dwell on substrates covered by a high amount of leaf litter. Stable carbon isotope analysis of the foot tissue of snails sampled from the S. alba stand and the mudflat indicated a transition in food source when a shell length of 51 mm is reached. Considering the δ13C value of juveniles, it seems they might be selecting for microphytobenthos, which might explain their preference for the mudflat. The diet of size classes found in both habitats did not differ significantly, although juveniles inhabiting the mangrove forest were slightly more depleted in 13C compared to those residing on the mudflat. Assuming juveniles feed on benthic microalgae and considering the lower microalgal biomass inside the mangrove forest, this may be a consequence of a higher contribution of other, more 13C depleted organic carbon sources, like phytoplankton, to their diet. Experimental results indicate a negative, but insignificant, impact on benthic diatom biomass by juveniles (due to grazing) and adults (due to physical disturbance). This finding seems to be in agreement with the results of the stable carbon isotope analysis, strongly suggesting the selective feeding of juvenile T. palustris on benthic diatoms.  相似文献   

11.
Spatial and temporal characteristics of the water masses and the dispersion of the suspended particulate matter were investigated using current meter, hydrographic and nephelometric observations, gathered during the ECOFER experiment (1989–1991) in the Cap-Ferret Canyon on the Aquitanian margin of the Bay of Biscay. While characteristics of the deep water masses were stable from one year to another, large hydrographic change in the upper 500 m related to winter renewal induced by poleward advection of warm and saline water along the continental slope. The slope circulation and seasonal eddy activity appear as important dynamical mechanisms that control the entrainment and the dispersion of the suspended particulate matter from the neritic domain to the deep ocean. A predominantly northward along-slope current with occasional reversal characterizes this circulation. The nephelometric structures also showed seasonal changes in the overall suspended particulate matter content, but recurrent features, such as the presence of intermediate nepheloid layers at the shelf-break depth and various depths along the slope (∼500, 1000 and 2000 m), were observed. These nepheloid layers extended off the slope to about 10–30 km, but especially laterally along the slope. Their presence indicated that suspended particulate matter exchanges between the shelf and the slope occurred mainly in the head of the canyon and along the southern open slope. The intermediate nepheloid layers around 500 m depth detached from the slope particularly in regions where the bottom slope is close to critical for the M2 internal tide.  相似文献   

12.
南海北部海洋雾状层特征及影响因素初步研究   总被引:1,自引:0,他引:1  
在粤东上升流区和南海北部陆架陆坡区共选取7条断面,通过分析水体的温度、盐度、密度、浊度、叶绿素浓度特征和悬浮体现场粒度平均粒径、体积浓度特征等,初步研究了南海北部海洋雾状层的特征及其受控机制。研究发现粤东上升流区底部雾状层普遍发育,在个别站位存在表层雾状层,高能的近岸浪流作用和来自港湾、河流的泥沙为表层雾状层和底部雾状层提供物源;南海北部陆架陆坡区底部雾状层也普遍发育,在珠江口外的内陆架和外陆架陆坡区表层雾状层比较常见,陆坡区中层雾状层比较发育,本区雾状层的物源主要来自珠江和台湾岛西南部河流的入海泥沙、海底沉积物以及生源颗粒物。南海北部底部雾状层的强度和扩散范围主要受水动力条件、河流入海物质的扩散和底质再悬浮等控制。  相似文献   

13.
Seawater along the southern margin of the Cretan Sea (May 1994–September 1995) has been found to have light transmission values ranging from 79% to 94%, corresponding to SPM values ranging from 1.5 mg l−1 to 0.2 mg l−1. The highest SPM concentrations (mostly of terrigenous origin) were found close to the sea-bed over the shelf-break and upper slope. The origins of SPM in the surface waters (<150 m) is principally biogenic. The occurrence of nepheloid layers at intermediate depths within the upper water column is mostly a result of density stratification. The dynamics of SPM distributions are governed by the 2-gyre system which induces a general onslope flow; and so inhibits the seaward dispersion of the relatively more turbid coastal/shelf waters. This is in agreement with the virtual absence of suspensates of terrigenous origin offshore of the shelf-break. Near bottom nepheloid layers (BNL) and detached intermediate nepheloid layers occur in the vicinity of the shelf-break and over upper slope region; these may be explained by resuspension induced by near-bed current activity and breaking of internal waves. High concentrations of SPM near the seabed may be caused by anthropogenic (trawling) activity. Occasionally, the formation of BNL may result from local seismic activity resulting in gravity-driven mass movements.  相似文献   

14.
Caribbean spiny lobster (Panulirus argus) settle preferentially in macroalgal‐covered hard‐bottom habitat, but seagrass is more prevalent in Florida (United States) and the Caribbean, so even low settlement of lobsters within seagrass could contribute substantially to recruitment if post‐settlement survival and growth were high. We tested the role of seagrass and hard‐bottom habitats for P. argus recruitment in three ways. We first explored possible density‐dependent regulation of early benthic juvenile lobster survival within cages deployed in seagrass and hard‐bottom habitats. Second, we compared settlement and survival of P. argus in both habitats, by comparing the recovery of microwire‐tagged early benthic juveniles from patches of seagrass and hard‐bottom. Finally, we assessed the relative abundance of juvenile lobsters in each habitat by deploying artificial structures in seagrass sites and compared these data with data from similar deployments of artificial structures in hard‐bottom habitat in other years. More early benthic juvenile lobsters were recovered from cages placed in hard‐bottom than in seagrass, but mortality of the early benthic life stage was high in both habitats. In regional surveys, the mean number of lobsters recovered from artificial shelters deployed within seagrass was lower than in any year that we sampled hard‐bottom, indicating that fewer lobsters reside naturally in seagrass, particularly large juveniles >40 mm carapace length. The greater abundance (and likely survival) of juvenile P. argus that we observed in hard‐bottom habitat as opposed to seagrass, combined with previous studies demonstrating that postlarval P. argus are attracted to, settle in, and metamorphose more quickly in red macroalgae, confirm that macroalgae‐dominated hard‐bottom habitat appears to be the preferred and more optimal nursery for Caribbean spiny lobster.  相似文献   

15.
The significance of a sheltering effect of seagrass against predators influencing the distribution patterns of benthic juveniles of the streaked goby Acentrogobius sp. was investigated by field experimentation in and around seagrass habitats at Moroiso and Aburatsubo Bays, Miura Peninsula, Japan. In the former bay, juveniles were always restricted to the seagrass bed, whereas at the latter, they also occurred over the surrounding bare sand substrate. Juveniles never occurred inside predator-exclusion cages over unvegetated sand in Moroiso Bay. Additionally, a tethering experiment resulted in similarly small numbers of juveniles being preyed upon in both the seagrass bed and over bare sand, suggesting that the predation risk for juveniles may not differ between the habitat types and was thus not responsible for their distribution patterns. The availability of symbiotic shrimp burrows, which were scarce in bare sand in Moroiso Bay but abundant in a similar area in Aburatsubo Bay, was also hypothesized as a determinant of distribution pattern. In a manipulative experiment at Aburatsubo Bay, however, juveniles showed no response to alterations in the availability of symbiotic shrimp burrows, indicating that neither the sheltering effect of seagrass against predators nor the availability of symbiotic shrimp burrows was a significant factor in streaked goby juvenile distribution. Food availability may be a determinant, because food abundance patterns were concordant with the juvenile distribution pattern.  相似文献   

16.
Boundary currents and internal waves determine cross-slope zonation of erosion and deposition in the Faeroe-Shetland Channel. Currents were measured at 8 and 34–50 m above the bottom at three mooring sites (502, 595 and 708 m depth) for 14 days. The structure of the water column was evaluated from CTD sections, and included nepheloid layers and particulate matter concentrations. Indicators for recent deposition in the sediment (organic carbon, phytopigments, 210Pb) were measured at eight stations across the slope. Strong near-bottom currents at the upper slope sustain down-slope particle transport in a benthic nepheloid layer, which is eroded under the influence of critically reflecting M2 internal tidal waves at 350–550 m, where the major pycnocline meets the sloping bottom. Beam attenuation profiles confirmed the presence of intermediate nepheloid layers intruding into the Channel along the major pycnocline, and elevated concentrations of particulate matter and chlorophyll-a were measured at this depth. Near-bottom currents decreased with depth, thus allowing particle deposition down the slope. Inventories of excess 210Pb activity in the sediment deeper than 600 m were higher than what was expected on the basis of atmospheric input of 210Pb and production in the water column, thus indicating additional lateral inputs. Simple calculations showed that off-slope input of particles from areas shallower than 600 m may be responsible for the enhanced deposition at greater depths.  相似文献   

17.
海洋雾状层的成因及其对海洋碳循环过程的影响   总被引:2,自引:0,他引:2  
海洋雾状层既是陆源物质进入海底的输送通道,又是海洋水体中沉降颗粒及底部再悬浮颗粒物的停留场所。雾状层物质来源主要有陆源、生源以及海底表层沉积物的再悬浮,不同海区、不同层位的雾状层的物质来源有所差异;雾状层的成因具有复杂性,既有物理作用,又有生物及化学作用,大量研究表明,海底洋流、内波(潮)等物理作用是雾状层形成的主要控制因素。雾状层中碳的存在形态主要有颗粒有机碳(POC)、溶解有机碳(DOC)、胶体有机碳(COC)以及无机碳,雾状层与其上下海水之间、雾状层与海底表层沉积物之间不同形态碳在生物-化学-物理动力系统作用下不断发生物质交换与迁移,对海洋碳循环生物地球化学过程起重要的控制作用,是整个海洋碳循环的一个不可忽视的环节。  相似文献   

18.
Abstract.  Few studies have described the spatial patterns of rocky sublittoral benthic communities in Brazil, where these habitats are almost exclusively located on the southeast and south coastline. The fauna and flora were analyzed using a quadrat method, and the species abundance and distribution were determined along transects down to the bottom sand plain on 10 rocky shores at Armação dos Búzios, Brazil. The 16 most common species accounted for around 97% of the benthos coverage and included cnidarians, algae, poriferans, and echinoderms. Cnidarians and algae were the most important groups, with abundance ranging from 13–66% and 27–68%, respectively. The species distribution indicated a clear depth zonation pattern. The algal community dominated the shallowest depths in sites more exposed to waves and steeper substrate slope, whereas the cnidarian community, divided into the zoanthidean Palythoa caribaeorum and the coral Siderastrea stellata , was more abundant from intermediate to greater (4 m) depths. Different community patterns were found between sites, demonstrating that different spatially discrete factors may be acting on each rocky shore.  相似文献   

19.
The productivity and health of seagrass depend on the combined inputs of nutrients from the water and sediments in which they grow and the microbiota with which they live intimately. However, little is known about the composition and diversity pattern of single-celled benthic eukaryotes in seagrass meadows. Here, we investigated how the structure and diversity of the benthic microeukaryotic community vary with respect to season, location, and seagrass colonization, by applying 18S rRNA gene amplicon sequencing for 96 surface sediment samples that were collected from three different seagrass habitats through four seasons. We found that benthic microeukaryotic communities associated with seagrass Zostera japonica exhibited remarkable spatial and seasonal variations, as well as differences between vegetated and unvegetated sediments. Diatoms and dinoflagellates predominated in the benthic microeukaryotic communities, but they were inversely correlated and displaced each other as the dominant microbial group in different seasons or habitats. Mucoromycota was more prevalent in vegetated sediments, whereas Lobulomycetales and Chytridiales had higher proportions in unvegetated sites. Total organic carbon and total organic nitrogen were the most important environmental factors in driving the microeukaryotic assemblages and diversity. Our study expands the available knowledge on the biogeographic distribution patterns and niche preferences for benthic microeukaryotes in seagrass systems.  相似文献   

20.
Nepheloid layer dynamics in the northern Portuguese shelf   总被引:1,自引:0,他引:1  
A general hydrographic and nephelometric survey of the NW Portuguese continental shelf and upper slope was undertaken, under winter and spring conditions. The nepheloid layer dynamics along the shelf during three cruises were controlled, principally by the following factors: (a) the hydrography of the shelf-slope waters, i.e. the nepheloid layers followed isopycnals and water masses; (b) prevalence of upwelling or downwelling circulation over the shelf; (c) dispersion of material by river discharge (mainly the Douro river); (d) resuspension of mid-shelf fine deposits induced by swell; (e) peculiar morphology with the presence of Porto Canyon and outer shelf rock outcrops.The general circulation controls the seaward extension of the nepheloid layers. In winter, under dominant downwelling conditions, an intense bottom nepheloid layer (BNL) was observed on the shelf, due to river borne particle supply and remobilization of mid-shelf muddy sediments (depth ≈100 m). In this period the BNL increased in thickness to the top of the slope, following the isopycnals. Near the shelf-break the BNL detached to form intermediate nepheloid layers (INL). Small INLs appear deeper in some areas of the slope. A surface nepheloid layer (SNL) appears in the surface water over the shelf and slope. In spring, biological particles mainly contribute to the SNL, which is separated from the BNL by a zone of clear water. In winter the SNL is restricted to the inner shelf where there are high inputs of fluvial particles.Downwelling circulation probably induces transport of shelf particles to deeper waters in the BNL. In spring, the predominant circulation was southward (upwelling), the water column was highly stratified, and dispersion of particles in the SNL was offshore.  相似文献   

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