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1.
The responses of desert rams to diet composition (concentrate vs. lucerne hay) in unshaded and shaded environments were investigated. The rectal temperature was higher (p < 0·01) with the concentrate diet but only in the afternoon, whereas the respiration rate was higher (p < 0·01) with the concentrate diet in unshaded and shaded environments during both morning and afternoon. Dry matter intake (DMI) was higher (p < 0·05) with the concentrate diet, while the ratio of water intake to DMI was higher (p < 0·01) with lucerne hay. Water gain through preformed and metabolic avenues and water loss through faeces and evaporative avenues were higher (p < 0·01) with the concentrate diet. With both diets, water turnover rate was higher (p < 0·05) in the unshaded environment. Feeding lucerne hay increased plasma total protein (p < 0·01) and albumin (p < 0·05) in unshaded and shaded environments. Plasma urea was higher (p < 0·01) with the concentrate diet. Plasma glucose level and plasma osmalality were higher (p < 0.01) and level of free fatty acids was lower (p < 0·01) with the concentrate diet in both thermal environments. Plasma cholesterol level was higher (p < 0·05) with the concentrate diet in the shaded environment, but lower (p < 0·05) in the unshaded environment. With lucerne hay, plasma cholesterol level remained the same in both environments.  相似文献   

2.
Three-week old soybean (Glycine max) plants were subjected to a factorial combination of four regimes of soil matric water potential (ψm=−0·03, −0·5, −1·0 and −1·5 MPa), two levels of supplementary Zn (O and 20 mgl−1) and two levels of foliar IAA application (O and 10 mgl−1). Under control conditions (no Zn, no IAA), increasing soil drying progressively retarded shoot and root growth (length and dry mass production), reduced leaf relative water content (RWC) and decreased the contents of chlorophyll (Chl) and shoot soluble sugars (SS), but increased soluble sugar content of roots and lowered osmotic water potential of shoots and roots (osmotic adjustment). Total free amino acid (TAA) content increased in shoots but decreased in roots whereas contents of soluble proteins (SP) decreased in shoots and roots. The effect of water stress was statistically significant (p<0·05) and had a major effect (as indicated by η2values) on leaf RWC, shoot and root dry masses and osmotic potential. Supplementary Zn improved root growth at all levels of stress and shoot growth under severe stress. Improvement of growth was positively correlated with the internal tissue Zn concentrations (r=0·91 and 0·86 for shoot and 0·94 and 0·82 for root length and dry mass respectively). Exogenous IAA raised (p<0·05) RWC, Chl, DM (slightly), root SS, and SP, whereas shoot TAA was lowered. Effects on root TAA and shoot SS were more complex: they were lowered at zero stress and raised under severe stress. IAA and Zn in combination had additive effects on Chl, growth and osmotic potential, but their combined effects on SP and TAA were more complex. It is concluded that the treatment of soybean plants grown under conditions of low soil water potentials and Zn deficiency with Zn and IAA solutions counteracted the deleterious effects of stress, especially at high stress levels, and helped stressed plants to grow successfully under these adverse unfavourable conditions.  相似文献   

3.
We studied the relationship between primary productivity and species richness of small mammals at both large and small spatial scales in the arid and semi-arid grasslands of north China. The productivity (x)–species richness (y) pattern at a large spatial scale can be described by a unimodal quadratic regression curve (y=7·41+0·1*x−0·0003*x2,p =0·008, r2=0·788). At a small spatial scale, however, neither linear nor quadratic regression fit the data for 1980 and 1994 (p>0·25). Primary productivity may not be an appropriate predictor of the species richness of small mammals at a small spatial scale. We conclude, therefore, that the primary productivity–species richness pattern of small mammals may be scale-dependent in the arid and semi-arid grasslands of north China. Landscape complexity should be considered in future studies of productivity–richness relationships.  相似文献   

4.
A greenhouse experiment was conducted to investigate the effects of moderate (14 cm) vs. severe (7 cm) defoliation on tiller recruitment and mortality within the interior and on the perimeter of weeping lovegrass (Eragrostis curvula) tussocks. Tiller number per marked shoot within the interior was significantly greater in plants defoliated at 14-cm stubble height than at 7-cm stubble height (2·37 vs. 0·43, p<0·05) 28 weeks after defoliation; likewise, tiller number per marked shoot was also greater on the perimeter of tussocks in the higher stubble than the lower stubble defoliation treatment (5·10 vs. 2·03, p<0·05). Tiller natality increased significantly 4 weeks after defoliation except for tillers cut to 7-cm stubble height within the interior of tussocks where tiller natality reached a peak at 4·5 weeks after defoliation. During the later stages of canopy development, tiller numbers per marked shoot gradually declined as tiller mortality increased. Tillers began senescing 13 weeks after defoliation, and the senescence rate was 52% greater in the 7-cm stubble height treatment than in the 14-cm stubble height treatment (2·33 vs. 1·53, p<0·0001). Within the interior of tussocks, number of the marked tillers more than doubled under the 14-cm stubble height treatment, whereas the plants lost 57% of the monitored tillers under the 7-cm stubble height treatment. We concluded that the central dieback process in weeping lovegrass is accelerated by severe defoliation.  相似文献   

5.
The profile distribution of total, diethylene triamine penta-acetic acid (DTPA)- and 0·1 HCl-extractable Fe and Mn were determined in 12 pedons formed on three contiguous sand dunes in the semi-arid savanna of Nigeria. The total Fe and Mn contents varied from 100 to 3750 and 40 to 11,375 μg g−1, respectively. Values of 0·1 HCl-extractable Fe and Mn varied from 8·0 to 123 and 1·5 to 43·5 μg g−1, respectively. The corresponding values of DTPA-extractable Fe and Mn were 4·5 to 16·0 and 1·0 to 38·8 μg g−1. Total Fe and Mn correlated significantly with clay in nearly all the dunes (p≤ 0·01) but were not significantly correlated with organic matter. The 0·1 HCl-extractable Mn had a positive significant correlation with soil pH (r= 0·58*) in Illela dunes but a negative significant correlation with pH in the Sangiwa dunes (r= −0·75***). The values of extractable Fe and Mn in the sand dune soils are above the critical limits reported in other literature, thus deficiencies of these micronutrients do not pose a problem to crop production in the study area.  相似文献   

6.
Nitrogen amendments (0, 25, 50 and 100 kg NH4NO3/ha) were used to study the responses of primary production, microbial biomass and nematode population in desert soil. The study was conducted in the Israeli Negev Desert, a region characterized by low and randomly distributed rainfall. Over a 1-year study period, nitrogen amendments resulted in a significant (p<0·01) increase in soil microbial biomass. Soil microbial biomass also increased concomitantly with the increase in soil organic matter. The number of free-living nematodes in the soil increased with the increase in soil moisture, ranging from 43,000 individuals per square metre at the end of the summer to 351,000 individuals per square metre during the rainy season. No significant correlation was found between the nitrogen treatments and the nematode population, whereas a significant positive correlation was found between the nitrogen amendments and the above-ground biomass (r2=0·94, p<0·03). The nitrate proportion of the total soluble nitrogen in the soil also increased with the increase in soil moisture. This study provides baseline data for nitrogen amendments on soil microbial status, as well as insights into the importance of nitrogen in fertility in arid environments.  相似文献   

7.
The Southern Alps of New Zealand are the topographic expression of active oblique continental convergence of the Australian and Pacific plates. Despite inferred high rates of tectonic and climatic forcing, the pattern of differential uplift and erosion remains uncertain. We use a 25-m DEM to conduct a regional-scale relief analysis of a 250-km long strip of the western Southern Alps (WSA). We present a preliminary map of regional erosion and denudation by overlaying mean basin relief, a modelled stream-power erosion index, river incision rates, historic landslide denudation rates, and landslide density. The interplay between strong tectonic and climatic forcing has led to relief production that locally attains 2 km in major catchments, with mean values of 0.65–0.68 km. Interpolation between elevations of major catchment divides indicates potential removal of l01–103 km3, or a mean basin relief of 0.51–0.85 km in the larger catchments. Local relief and inferred river incision rates into bedrock are highest about 50–67% of the distance between the Alpine fault and the main divide. The mean regional relief variability is ± 0.5 km.Local relief, valley cross-sectional area, and catchment width correlate moderately with catchment area, and also reach maximum values between the range front and the divide. Hypsometric integrals show scale dependence, and together with hypsometric curves, are insufficient to clearly differentiate between glacial and fluvial dominated basins. Mean slope angle in the WSA (ψ = 30°) is lower where major longitudinal valleys and extensive ice cover occur, and may be an insensitive measure of regional relief. Modal slope angle is strikingly uniform throughout the WSA (φ = 38–40°), and may record adjustment to runoff and landsliding. Both ψ and φ show non-linear relationships with elevation, which we attribute to dominant geomorphic process domains, such as fluvial processes in low-altitude valley trains, surface runoff and frequent landsliding on montane hillslopes, “relief dampening” by glaciers, and rock fall/avalanching on steep main-divide slopes.  相似文献   

8.
Sedimentary deposits from the Smith Canyon dune field, south-central Columbia Basin, Washington, U.S.A. document climatically-influenced Late Pleistocene and Holocene aeolian and fluvial deposition in a region impacted by glacial outburst floods and tephra falls. The depositional history is summarized by five environmentally distinctive and climatically sensitive sedimentary units (temporal limits estimated): Unit 1 (c. 15·5–8 ka), pedogenically altered glacial outburst flood and minor aeolian silt and clay; Unit 2 (c. 8–6·9 ka), fluvial and minor aeolian sand; Unit 3 (c. 6·9–6·8 ka), flood-induced fluvial sand with gravel-sized tephra clasts; Unit 4 (c. 6·8–3·9 ka), aeolian dune sand; Unit 5 (c. 3·9 ka to present), pedogenically altered, stabilized dune sand. Estimated age ranges are based on stratigraphic position, tephrochronology, and correlation with temporally constrained strata from elsewhere in the region.  相似文献   

9.
The selective use by cattle of range resources in the Sahel was assessed in terms of quantity and nutritional quality of the herbage grazed and diet selected. Peak standing crop was 1951 kg ha−1dry matter (DM) with 47·9 kg nitrogen (N) ha−1and 7·4 kg phosphorus (P) ha−1. Esophageally fistulated cattle diets were consistently (p<0·05) higher in N, P and organic matter digestibility (OMD) than the herbage grazed. Cattle consumed 48% of the mean annual herbage production. The results indicate grazing ruminants select a higher quality diet from Sahelian rangelands than can be predicted on the basis of pasture evaluation alone. Therefore, the foraging behaviour of the animals needs to be considered in the evaluation of Sahelian rangelands for animal production.  相似文献   

10.
Field and laboratory studies were conducted during the period May 1993–April 1994 on the cocoons and hatchlings ofAporrectodea calignosa(Savigny 1826), a common earthworm found in Benghazi, Libya. Cocoon production started in August 1993 and continued until April 1994. A positive correlation between soil moisture (r=0.74;p< 0.05) and cocoon production, and a negative one between soil temperature (r=−0.85;p< 0.05) on cocoon production were found. Mean incubation time was 47 days. Some cocoons contained a single worm (single hatchling) whereas the majority of them (68%) contained two, the first hatchling and second hatchling. The rate of change of body mass (F=2.18;p>0.05) and the average body mass (F=1.25;p>0.05) of the single, first and second hatchlings were similar.  相似文献   

11.
The natural abundance15N/14N method was used to estimate the influence of silvicultural and P fertilization treatments on N accretion, N2fixation and N partitioning among tissues in a mature mesquiteProsopis glandulosavar.glandulosastand in Texas. The silvicultural treatments consisted of understory removal, herbicide treatment of brushy resprouts, thinning trees to single stems and 100 kg ha−1P fertilization. The trees had a mean basal diameter of 17·8 cm with 8 to 35 cm range. The stand was slow growing with the increase in dry matter ranging from 0·465 Mg ha−1year−1to 0·701 Mg ha−1year−1for the 8 years after the treatments were applied. N accretion after 8 years ranged from 3·1 kg ha−1year−1to 4·4 kg ha−1year−1.Due to the range in δ15N of the leaves, twigs, branches and trunk, we used the weighted (by biomass) average δ15N per tree in calculations of the percent N derived from N2fixation (%Ndfa). There was considerable variability in δ15N of the reference plants, i.e. from 3·3 to 5·9. In contrast there was low variability in the background δ15N of nearby soils (7·0±1·0). As the total above-ground biomass δ15N of a grass grown outside the influence of mesquite (7·8±0·58) had the same δ15N as the soil (7·5±1·0), we used the grass outside the influence of mesquite and the weighted tree mean δ15N to calculate % of N derived from N2fixation.The decrease in intraspecific competition by thinning multistemed trees to single stemmed trees was the only treatment that significantly (p= 0·0001) increased growth. Interspecific competition, i.e. understory removal, did not increase growth. There were no significant differences in total N production or N fixation among treatment means. The most striking result was the highly positive correlation between tree δ15N and total N per tree and biomass per tree (R2= 0·90,F= 164·4, df. = 18, mean square error (MSE) = 0·155,p= 0·0001). This implies that the younger trees colonizing infertile soils relied more heavily on N2fixation than larger trees which accumulated 1200 kg ha−1more N under their canopies. The percentage N derived from N2fixation ranged from 63 to 73% in the various treatments. Despite the high percentage of N derived from N2fixation, the N2fixation of the stand was very low, i.e. 1·98 to 2·80 kg N ha−1year−1, due to the low growth of the stand. We believe that comparisons of the whole tree weighted δ15N to background soil δ15N provides a more reasonable approach to estimate % N2fixation than comparisons of leaves of fixers and reference plants.  相似文献   

12.
The impact of range plant toxins on the reproductive performance of female cattle is economically important to the livestock industry. Nevertheless, there is little information on the significance of toxicosis from range plants on semen quality in domestic ruminants. A total of 18 adult mixed-breed bucks under range conditions were used to evaluate the effect of diet composition (microhistological analysis of fecal samples) on some semen characteristics, and blood and fecal components in April and August. Forages in the buck diets were grouped into 2 classes: low or high levels of each plant in the buck diets. Bucks with higher proportion of Acacia greggii, Flourensia cernua and Lindleya mespiloides in their diets yielded 23–50% less (p<0.05) semen than bucks with low proportion of these shrubs in their diet. Sperm motility was reduced by 3–8 percent units (p<0.05) by high levels of five rangeland shrubs in the buck diets. Bucks consuming high levels of Acacia greggii showed significantly lower percentages of normal sperm (92±10 vs. 96±3; p<0.05) than bucks with low intake of this forage. High proportions of Rhus virens and Solanum elaeagnifolium in the buck diet increased the percentage of normal sperm (5 units, p<0.05) when compared to bucks with low utilization of these forages. Increased intake of Acacia greggii, Dalea bicolor, Opuntia rastrera, Larrea tridentata and Cowania plicata reduced percentage of live sperm by 5 units (p<0.05). Blood metabolites analysis showed that increased levels of Acacia greggii and Parthenium incanum were related to better nutritional status of bucks. Zn (−), Cu (−) and glucose (+) were significant predictors of semen volume (r2=0.30). Cu (−) and cholesterol (−) accounted for 37% of the variation in percentage of normal sperms. Forty-four percent of the variation in secondary abnormalities was accounted by Zn (+) and cholesterol (+), whereas fecal N, serum total proteins and creatinine positively affected percentage of live sperms (r2=0.37). These results indicate that semen quality and metabolic profiles of grazing bucks were sensitive to the ingestion of some Chihuahuan desert forages.  相似文献   

13.
We have monitored initiation conditions for six debris flows between May 2004 and July 2006 in a 0.3 km2 drainage basin at Chalk Cliffs; a band of hydrothermally-altered quartz monzonite in central Colorado. Debris flows were initiated by water runoff from colluvium and bedrock that entrained sediment from rills and channels with slopes ranging from about 14° to 45°. The availability of channel material is essentially unlimited because of thick channel fill and refilling following debris flows by rock fall and dry ravel processes. Rainfall exceeding I = 6.61(D)− 0.77, where I is rainfall intensity (mm/h), and D is duration (h), was required for the initiation of debris flows in the drainage basin. The approximate minimum runoff discharge from the surface of bedrock required to initiate debris flows in the channels was 0.15 m3/s. Colluvium in the basin was unsaturated immediately prior to (antecedent) and during debris flows. Antecedent, volumetric moisture levels in colluvium at depths of 1 cm and 29 cm ranged from 4–9%, and 4–7%, respectively. During debris flows, peak moisture levels in colluvium at depths of 1 cm and 29 cm ranged from 10–20%, and 4–12%, respectively. Channel sediment at a depth of 45 cm was unsaturated before and during debris flows; antecedent moisture ranged from 20–22%, and peak moisture ranged from 24–38%. Although we have no measurements from shallow rill or channel sediment, we infer that it was unsaturated before debris flows, and saturated by surface-water runoff during debris flows.Our results allow us to make the following general statements with regard to debris flows generated by runoff in semi-arid to arid mountainous regions: 1) high antecedent moisture levels in hillslope and channel sediment are not required for the initiation of debris flows by runoff, 2) locations of entrainment of sediment by successive runoff events can vary within a basin as a function of variations in the thickness of existing channel fill and the rate of replenishment of channel fill by rock fall and dry ravel processes following debris flows, and 3) rainfall and simulated surface-water discharge thresholds can be useful in understanding and predicting debris flows generated by runoff and sediment entrainment.  相似文献   

14.
We present a remotely-sensed degradation assessment technique based on the measurement of landscape heterogeneity (a key determinant of degradation status). The moving standard deviation index (MSDI) is derived by passing a moving standard deviation filter across the Landsat TM band 3 (red band). Degraded/unstable landscapes exhibit higher MSDI values than their undisturbed/stable counterparts. Significant differences in MSDI were detected (p<0·001) across four fence-lines which separated rangeland of contrasting condition. The relationship of the index to normalized difference vegetation index (NDVI) was tested in five different ecosystems and significant correlations were obtained in all cases (p<0·001). We propose the MSDI as a powerful adjunct to vegetation indices.  相似文献   

15.
Impacts of six initial treatments and rainfall on maximum above-ground herbaceous standing crop were assessed in 1978 and annually from 1984 to 1992 in rangelands of east-central semi-arid Argentina. Treatments applied in 1977–1978 were: (1) untreated control; (2) burning; (3) herbicide application (shrub control); (4) Old Field 1 or (5) Old Field 2 (areas previously exposed to different degrees of mechanical soil disturbance for 25 years); and (6) overgrazing. Domestic herbivores were excluded thereafter from all treatments until 1993.During 1984–1992, total above-ground herbaceous standing crop often remained greater in both Old Fields than in the other treatments; highest mean values were 2336 and 1640 kg ha−1for Old Field 1 and Old Field 2 treatments, respectively. Lowest total above-ground standing crops in all treatments (M = 296–475 kg ha−1) occurred in 1989, the year with the lowest annual rainfall (257·5 mm). Desirable perennial grasses contributed most (44–100%) to total herbaceous standing crop in all treatments during the study period. Most of this standing crop was made up of the cool-season grassesPiptochaetium napostaense, Poa ligularis, Stipa clarazii, S. papposaandS. tenuis. Annual rainfall was closely related (p< 0·05) to total herbaceous and desirable perennial grass standing crops in most treatments, and accounted for most of the variation in herbage production between years.  相似文献   

16.
The objectives of this study were to determine the composition, density and spatial distribution of the soil seed bank of woody species, as well as their regeneration pattern in two different land use systems, controlled (ranch) and open grazing, in an Acacia woodland of the Rift Valley in Ethiopia. We also compared the species composition of the soil seed bank and the above-ground vegetation to find out if differences exist in the soil seed bank and advance regeneration between the two land use systems. The germination requirements of seeds of the woody species were also investigated under laboratory conditions. Acacia senegal, Acacia seyal, Acacia tortilis, Dichrostacys cinerea and Balanites aegyptiaca were encountered in the above-ground vegetation in both systems. Seeds of only A. tortilis (90±32 seeds m−2) from the ranch, and A. senegal (5±3 seeds m−2) and A. tortilis (72±34 seeds m−2) from the open system were found in the soil seed bank along transects with patchy horizontal pattern. The two systems were not significantly different in density of soil seed banks of A. senegal andA. tortilis . Jaccard's similarity index showed that only a few woody species were common in the soil seed flora and above ground vegetation. However, all of the species accumulated seeds (58±43–331±130 seeds m−2) in/on the soil under the canopy. Very large numbers of seeds of A. tortilis (19382±9722 seeds m−2) and D. cinerea (1278±494 seeds m−2) were also found in barns. Most of the seeds recovered from the soil samples (60–80%) were found in the litter layer. Acid and mechanical scarification improved legume germination (36–99% and 60–99%, respectively) over boiling water (0–48%). Treatment means differed significantly for all the legumes (p<0·001) but not for Balanites. Height and diameter class distribution of regeneration of A. tortilis and D. cinerea in both systems and A. senegal in the open system had a negative exponential correlation (rs=−0·5, −0·25 and −0·86, respectively). A. seyal and B. aegyptiaca showed poor regeneration. Horizontal distribution of advance regeneration of all the species was patchy. Advance regeneration of A. seyal, A. tortilis and B. aegyptiaca were not significantly different, while that of A. senegal and D. cinerea were significantly different between the two systems. Poor representation of species in the soil seed bank along transects and in the different height and diameter classes may be attributed to the low density of mature trees as well as the mode and strategy of seed dispersal. Ungulate and wind dispersed species (e.g. A. senegal, A. tortilis and D. cinerea) were highly favoured. Patchiness in the distribution of seeds and advance regeneration was also a result of endozoochory. Dispersal of non-ungulate dispersed seeds (A. senegal, A. seyal and B. aegyptiaca) was restricted to the canopy zone. Piles of seeds ofA. tortilis and D. cinerea that were found in barns were a result of consumption of their pods by cattle. High concentration of seeds in the litter layer may be due to low soil disturbance and larger size of seeds. The height and diameter class distribution of A. senegal (in the open system), A. tortilis and D. cinerea also indicated that the species have good regeneration. Results from the germination tests indicated that seeds of the legumes require pre-sowing treatments to give a rapid, uniform and improved germination. Intervention through artificial regeneration should be employed to improve the density and regeneration capacity of those species with hampered regeneration at both systems.  相似文献   

17.
As an approach to understand how diurnal and seasonal plant water potentials (Ψ) are related to soil water-content and evaporative demand components, the responses of six thornscrub shrubs species (Havardia pallens, Acacia rigidula, Eysenhardtia texana, Diospyros texana, Randia rhagocarpa, and Bernardia myricaefolia) of the north-eastern region of Mexico to drought stress were investigated during the course of 1 year. All study species showed the typical diurnal pattern of variation in Ψ. That is, Ψ decreased gradually from predawn (Ψpd) maximal values to reach minima at midday (Ψmd) and began to recover in the late afternoon. On a diurnal basis and with adequate soil water-content (>0.20 kg kg−1), diurnal Ψ values differed among shrub species and were negatively and significantly (p<0.001) correlated with air temperature (r=−0.741 to −0.883) and vapor pressure deficit (r=−0.750 to −0.817); in contrast, a positive and significant (p<0.001) relationship was found to exist with relative humidity (r=0.758–0.842). On a seasonal basis, during the wettest period (soil water-content>0.20 kg kg−1), higher Ψpd (−0.10 MPa) and Ψmd (−1.16 MPa) values were observed in R. rhagocarpa, whereas lower figures (−0.26 and −2.73 MPa, respectively) were detected in A. rigidula. On the other hand, during the driest period (soil water-content<0.15 kg kg−1), Ψpd and Ψmd values were below −7.3 MPa; i.e. when shrubs species faced severe water deficit. Soil water-content at different soil layers, monthly mean relative humidity and monthly precipitation were significantly correlated with both Ψpd (r=0.538–0.953; p<0.01) and Ψmd (r=0.431–0.906; p<0.05). Average soil water-content in the 0–50 cm soil depth profile explained between 70% and 87% of the variation in Ψpd. Results have shown that when gravimetric soil water-content values were above 0.15 kg kg−1, Ψpd values were high and constant; below this threshold value, Ψ declined gradually. Among all shrub species, A. rigidula appeared to be the most drought tolerant of the six species since during dry periods it tends to sustain significantly higher Ψpd in relation to B. myricaefolia. The remaining species showed an intermediate pattern. It is concluded that the ability of shrub species to cope with drought stress depends on the pattern of water uptake and the extent to control water loss through the transpirational flux.  相似文献   

18.
Distinct rock fragment displacements occur on the ambas, or structurally determined stepped mountains of the Northern Ethiopian Highlands. This paper describes the rock fragment detachment from cliffs by rockfall, quantifies its annual rate, and identifies factors controlling rock fragment movement on the scree slopes. It further presents a conceptual model explaining rock fragment cover at the soil surface in these landscapes. In the May Zegzeg catchment (Dogu'a Tembien district, Tigray), rockfall from cliffs and rock fragment movement on debris slopes by runoff and livestock trampling were monitored over a 4-year period (1998–2001). Rockfall and rock fragment transport mainly induced by livestock trampling appear to be important geomorphic processes. Along a 1500-m long section of the Amba Aradam sandstone cliff, at least 80 t of rocks are detached yearly and fall over a mean vertical distance of 24 m resulting in a mean annual cliff retreat rate of 0.37 mm y− 1. Yearly unit rock fragment transport rates on scree slopes ranged between 23.1 and 37.9 kg m− 1 y− 1. This process is virtually stopped when exclosures are established. Corresponding mean rock fragment transport coefficients K are 32–69 kg m− 1 y− 1 on rangeland but only 3.9 kg m− 1 y− 1 in densely vegetated exclosures. A conceptual model indicates that besides rockfall from cliffs and argillipedoturbation, all factors and processes of rock fragment redistribution in the study area are of anthropogenic origin.  相似文献   

19.
The standard metabolic rate ofUromastyx philbyiwas measured at 20–40 °C, using constant pressure manometric respirometers. Standard metabolic rate was mass-dependent. Values for the mass exponent ‘b’ ranged from 0·75 at 20 °C to 0·82 at 40 °C. Standard metabolic rate increased as temperature increased with high Q10values at low temperatures (20–25 °C). Low standard metabolic rates in this species are probably an adaptation to herbivory in hot desert environments.  相似文献   

20.
《自然地理学》2013,34(6):561-573
The semi-arid forest-steppe ecotone in China is characterized by a patchy pattern of forest and steppe, with forest patches restricted to shady slopes. To address the effect of topography on forest distribution through regulation of available water, we calculated evaporation as a function of slope aspect and inclination. Field vegetation records from randomly selected sites with minimum slope inclination were used to test the simulated forest distribution. Seasonal and diurnal changes of surface soil temperature and moisture of shady and sunny slopes were recorded. Soil water content was measured during two growing seasons on both sunny and shady slopes with the same forest type at three sites located along the mean annual precipitation (MAP) gradient. Evaporation decreases with slope inclination on shady slopes, but increases with inclination on sunny slopes. The shady slope received 35% of the annual direct solar radiation received by the sunny slope when the slope inclination was 25°, and the contrast in annual direct solar radiation between the shady and sunny slopes further widens as slope inclination increases. Steeper shady slopes can support forests in dryer climates, with log-linear regression revealing a minimum slope inclination for forest distribution along the MAP gradient. The simulated minimum slope inclination for forest growth was larger than the observed minimum inclination, and the difference was greater in wetter conditions. A larger forest area fraction was considered to lead to a reduction in soil temperature and evaporation, as verified by soil temperature and moisture records and soil water content measurements. The slope-specific forest distribution in the semi-arid region of China can be explained by a topography-controlled soil water supply. Lower evaporation, resulting from lower direct solar radiation on shady slopes, allows shady slopes to retain a water supply sufficient for sustaining forests, and the existence of forests on shady slopes further reduces evaporation. Different tree species coexist at the xeric timberline due to regulation by slope inclination and aspect.  相似文献   

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