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1.
Grazing in arid Mediterranean ecosystems brings about changes in species diversity, but the best way to measure such changes is unclear. In this study we compared various methods in order to identify indicators that might be useful for the management and conservation of grazed arid Mediterranean ecosystems.Changes in community structure and composition were compared along a previously studied grazing gradient in Cabo de Gata-Nijar Natural Park. Plant diversity was measured by calculating diversity (Shannon information index), evenness (Pielou index) and richness (species number). Rarefaction curves were used to measure plant species richness and the results were compared with traditional richness values. Community structure was measured as the percentage of bare ground and the coverage of overall perennial grasses, Stipa tenacissima L., perennial shrubs and annual plants. Our results showed that the proportion of bare ground, the cover of perennial shrubs and the relative abundance of S. tenacissima are good indicators of grazing effects on vegetation. Overall plant community structure was more sensitive to the grazing gradient than were the diversity indices. Finally, the adequacy of standardized methods for comparing species richness (i.e. rarefaction curves) along a grazing gradient are discussed.  相似文献   

2.
Grazing effects on patchy dryland vegetation in northern Patagonia   总被引:1,自引:0,他引:1  
In this study the spatial patterns and dynamics of vegetation patches along a grazing gradient in the steppe ofLarrea divaricataandStipaspp. in NE Patagonia (Argentina) are described. A general effect of grazing is the reduction of total plant cover resulting from the decrease in cover of perennial grasses (Stipa speciosa, Poa ligularis, Stipa tenuis) and some tall shrubs (Chuquiraga hystrix, Bougainvillea spinosa, Lycium chilense). Dwarf shrubs (Nassauvia fueguianaandJunellia seriphioides) increase their cover under medium and/or high grazing pressures. Plant species are spatially grouped into patches which alternate with areas of bare soil. Eleven types of vegetation patch differing in the dominant plant functional type or species, floristic richness and size were identified with different relative frequency along the grazing gradient. Based on these results, it is postulated that grazing forces the replacement of large patches dominated by tall shrubs with high species richness, byLarrea divaricatapatches or small dwarf shrub patches with low species richness and the extinction of grass patches. This results from: (1) disruption of local balances of species deletions and additions; (2) fragmentation of large patches; and (3) formation of new vegetation patches. These changes lead to differing plant spatial organization and heterogeneity along the grazing gradient which may be described by characteristic arrays of vegetation patches.  相似文献   

3.
This study was conducted in the arid Mediterranean Reserve located in northern Jordan, representing a continuous grazed arid rangeland ecosystem. Quantitative analyses on species diversity in addition to phytogeographical analysis were conducted. Some ecological parameters such as, life forms, growth forms, species density and frequency were investigated. A total of 93 vascular plant taxa belonging to 78 genera and 26 families were reported. This constitutes about 4.3% of the total flora of Jordan. Reported taxa include 13 species of grasses, 69 forbs and 10 shrubs and semi-shrubs. The botanical survey pointed to the presence of four rare species in Jordan. Plant species were classified into 28 functional groups. Pairwise correlation analyses among the most abundant species, showed that neutral associations dominated (75%) those communities, followed by positive associations (19%), while only (6%) showed negative association. Positive association may be attributed to direct facultative mutualism, or to physical environmental heterogeneity. Such a positive cooperation in the plant community can significantly maintain species diversity in adverse environments.The relationship between primary productivity and species richness showed a humpbacked pattern. Maximum richness corresponded to a biomass level of 300–400 g m−2. However, a sinusoidal curve-fitting model explained the relationship between graminoids biomass and species richness. These findings pointed to the adaptation of plant taxa to livestock grazing and to the small-scale spatial and environmental heterogeneity in arid Middle Eastern Mediterranean rangelands. Results suggest that optimal biodiversity conservation for the arid Middle Eastern grazing lands should consider their high level of spatial heterogeneity. Rangeland managers should adopt grazing plans that apply a range of grazing pressure across the area so as to maintain diversity at local and regional scales.  相似文献   

4.
Mongolian herders are transhumant and therefore follow a specific land use pattern. To understand their movements through a case study, we mapped and superimposed their seasonal camps on a vegetation map and a digital elevation model. We also questioned them about the reasons justifying the locations of their camps, and how they use the surrounding land. It appears that vegetation quality may play a role only in summer, whereas topography is a key driver during 3 seasons. In winter, herders seek shelter against cold winds in rugged places exposed to the south, while in spring and summer, they seek flat places. In spring, to have a clear view of their newborns and in summer, to allow wind to reduce the insect discomfort. Moreover, most of the livestock species stay within a certain distance of the camps depending on the season, but which never exceeds 5.1 km. This leads to a land use pattern where livestock is restrained to specific areas according to the season. Interestingly, during the growing season, when plants are most sensitive to defoliation, most livestock is concentrated on 30.92% of the site surface. Such information is important for range scientists working on grazing impacts at large scales.  相似文献   

5.
Heuweltjies are large circular earth mounds created by termites and are important features of the landscape in the Succulent Karoo biome in South Africa. This study demonstrates that heuweltjies and livestock grazing act together to shape vegetation patterns in the region. Using a multifactorial approach, we investigated the combined effects of heuweltjie-induced habitat conditions, grazing pressure and physical disturbance by livestock and native animals as well as the interactions among two distinct plant functional groups, on the composition and cover of vascular plant vegetation. Heuweltjie soils had significantly higher pH-values, finer texture, and higher concentrations of carbon, nitrogen, and phosphorus than those of the surroundings. Neither perennial nor ephemeral cover was different on- and off-mounds. However, species richness, cover of highly palatable plants and livestock grazing pressure were significantly higher between mounds. Grazing pressure generally resulted in a loss of perennial species and a shift to more ephemeral species, probably through the competitive release of annuals and geophytes from perennial plants. We propose that the apparent contradiction between superior nutrient conditions and inferior plant food quality on heuweltjies is caused by a combination of otherwise unfavourable soil properties and historic overgrazing of heuweltjies resulting in local degradation of the vegetation.  相似文献   

6.
Plant assemblages' dynamics have been studied to evaluate the influence of different environmental factors. The aim of our work was to assess the effect of a disturbance in the form of livestock grazing on annual plants, in a South American desert. We tested the exclusion of cattle grazing by comparing the composition of annual plants in three major vegetation types within a MaB Reserve in the Monte Desert of Argentina, with those of an adjacent grazed field. Sampling was conducted in two consecutive years that differed in precipitation. We established three sampling sites within each vegetation type at the reserve and the grazed field. Transects were set to assess plant cover, abundance, and richness. Theoretically, changes in diversity are explained by changes in one of its components: species richness. Species richness of annual plants was not different between the grazed and ungrazed sites. However, plant cover and diversity were lower in grazed sites, whereas abundance increased. Owing to a strong species-specific effect, we propose that it is evenness the main parameter involved in diversity dynamics in the heterogeneous vegetation mosaic of the Monte desert. Finally, vegetation types (spatial heterogeneity) and rainfall regime (temporal heterogeneity) greatly interacted with grazing effects. We endorse the idea that rapid responses of annual plant assemblages to changes in rainfall conditions coupled with herbivore control, might result in a restoration pathway for degraded arid landscapes.  相似文献   

7.
In view of the repeatedly reported overstocking of the high-altitude pastures on Al Jabal al Akhdar, northern Oman, plant species abundance, cover and frequency, and herbaceous mass yield were studied in ungrazed versus heavily grazed areas of this mountain range. In addition, plant species selection by goats along a gradient of 1000–2000 m and spatial extent of pasture areas were investigated after abundant rainfall and a subsequent 6-months dry spell by means of manual observation and GPS/GIS tools.The substantially higher species diversity and herbaceous mass yield in the ungrazed area illustrate the production potential of these mountain pastures or, respectively, the biodiversity and productivity loss resulting from continuous grazing. The concentration of goats' selection on only a dozen herbaceous and ligneous species favours pasture encroachment with poisonous shrubs such as Nerium mascatense in the lower and Dodonaea viscosa in the higher altitudes. Given the spatially limited extent of pasture areas, these are exposed to high stocking rates. Therefore, grazing and feeding schemes need to be developed which reduce livestock pressure on the pastures, taking into account local property rights, herding skills and the recovery potential of the vegetation, which heavily depends on unpredictable rainfall events.  相似文献   

8.
Overgrazing by livestock has caused desertification in the Monte, where ctenomyids and livestock share grasses as main food items. The diet of Ctenomys eremophilus, above-ground food availability and changes related to cattle grazing are analyzed in the arid plain of Mendoza, Argentina. The most available categories were grasses, followed by low shrubs and tall shrubs. Tuco-tucos showed dietary generalism, ate mainly above-ground plant parts, preferred grasses and avoided shrubs at both grazed and ungrazed sites. Plant cover, grass diversity and availability decreased under livestock grazing, which was reflected in the diet by a lower percentage of grasses, a shift toward low shrubs and higher number of frequently used resources. Tuco-tucos in the grazed paddock compensated for lower consumption of vegetative plant parts by increasing the use of Prosopis flexuosa pods stored inside burrows. Moreover, greater dietary variation among individuals suggests foraging restricted to the items closest to burrow holes. These feeding tactics would allow them to reduce above-ground foraging as a response to high raptor predation risk due to increased bare soil. The plant recovery detected during the rest period, favoured by moderate stocking rate and rotational grazing system, would allow coexistence of tuco-tucos and cattle.  相似文献   

9.
基于幂函数法则对放牧梯度上种群空间异质性的定量分析   总被引:3,自引:1,他引:2  
 在呼伦贝尔草原的克氏针茅群落选择轻牧、中牧、重牧3个不同的放牧梯度,沿着50 m样线设置100个50 cm×50 cm的L型样方,每个L型样方内划分4个25 cm×25 cm的S型样方,以物种出现的频度(p)作为植被空间分布的观测指标,采用幂函数法则对放牧梯度上草原植物群落物种的空间异质性进行了定量研究。结果表明,在不同放牧强度下,种类结构发生明显改变,轻度放牧区羊草和星状刺骨藜、中度放牧区藜和糙隐子草、重度放牧区猪毛菜和多根葱具有高的出现频度,群落物种数随着放牧压力的增大而减小。整体来看,放牧梯度上物种的空间分布呈现明显的聚集分布,物种空间异质性(δi)随着放牧强度的增大而增加,拟合于幂函数规律曲线;群落的空间异质性(δc)随着轻牧-中牧-重牧放牧演替的进展呈现降低趋势,种群个体分布呈现片断化特征。在群落内部各个种群具有不同的异质化分布特点,物种结构主要表现出密集型种群和疏散型种群两大类型。这些趋势与种群的繁殖策略、生活史对策、群落的再生产策略等生理与生态特性密切相关。  相似文献   

10.
Camel grazing is recognized as a primary cause of ecological degradation in the UAE. A study of perennial plant species <1 m in height was conducted along a fence separating continuously camel grazed land from land in which camels had been replaced by oryx and gazelle species for 5 years (Al Maha). Vegetation regeneration in Al Maha in the absence of camels was considerable on all substrates (gravel, stable sand, and semi-stable sand) but was greatest on the gravel substratum, indicating that ecology in this habitat is most at risk. Observed regeneration was primarily through vegetative reproduction and growth of existing plants, showing that existing species can tolerate heavy grazing. Therefore, an equilibrium grazing model of continuous and reversible vegetation dynamics is most suitable for management of this ecological zone. Species richness was greater in Al Maha due to the greater number of plants, but biodiversity was unaffected. There was some evidence of localized dune stabilization within Al Maha due to increased vegetative cover. Further recovery of vegetation within Al Maha is discussed. This study highlights the need for reduced grazing pressure throughout the Dubai inland desert, and in particular on gravel substrata.  相似文献   

11.
The Southern Kalahari Duneveld is an arid savannah, which has been used for permanent stock farming since around 1900. Farming with high numbers of animals has led to changes in the vegetation which have been associated with excessive grazing and trampling. To determine whether altered seed dispersal could be an alternative or additional explanation, we investigated how replacing antelope with sheep affects seed dispersal. We determined the dispersal potentials of plant species by various vectors using standardized experiments and studied the variation of dispersal potential along a grazing intensity gradient and among different habitats.Wind dispersal potential was low, but diaspores of many plant species were well adapted to dispersal by water. Biotic long-distance dispersal potentials were relatively high and varied between sheep and antelope: the species of plants dispersed differed, as did the number of seeds transported and the time of year that seeds were dispersed in.The importance of zoochory and the differences found between antelope and sheep dispersal suggest that dispersal is affected by a change of animal type. Thus, sheep can not replace antelope as seed dispersers. Therefore, sustainable grazing management or restoration efforts should benefit from seed input of sensitive species where antelope are rare.  相似文献   

12.
The arid rangelands of Syria cover over half of the nation's landmass. Punctuating this landscape are broad, dry basins, or wadis, and gentle landscape depressions that exhibit localized elevated vegetation productivity and unique edaphic and hydrologic properties. Historically, continuous heavy grazing and aggressive agricultural activities resulted in excessive ecological degradation within these sensitive environments. Information is needed to determine the influence of livestock grazing on plant communities in landscape depressions and the impact that this has on ecosystem resilience. The purpose of this research is to evaluate the effect of short-term sheep grazing on vegetation characteristics and plant community structure within depressions, and to provide recommendations for improved grazing management. Study plots were randomly located within paired topographic depressions located in northwestern Syria. Vegetation samples were collected along transects including plant biomass, plant density, herbaceous cover, and species diversity. In grazed plots, plant biomass was 49 g DM/m2 compared to 234.4 g DM/m2 in protected plots. Average plant density was 65 plants/m2 in grazed plots compared to 1013 plants/m2 in protected plots (P = 0.001). Herbaceous cover was 175% higher on protected sites compared to grazed plots. Average diversity (Shannon–Wiener index value) was 0.8 in grazed plots compared to 2.3 in protected plots. These results suggest that plant community structure will be impacted from short-term grazing and that a site's ability to positively respond to disturbance over time may be limited. We conclude that carefully planned grazing management should result in greater plant productivity and diversity.  相似文献   

13.
Snow cover strongly influences plant growth in Arctic and alpine ecosystems. Snow characteristics and snowmelt timing are likely to change in a warmer climate. We studied year rings and shoot growth of the dwarf shrub bilberry ( Vaccinium myrtillus ), and species abundances of the vegetation, in response to early or late snowmelt at a study site in the Central Alps, near Davos, Switzerland. Snowmelt was manipulated on experimental plots for 3 and 30 years. Additional plots were set up along a natural snowmelt gradient, and at high and low elevation. Growth ring data showed an increasing trend in annual growth increment over the last 20 years, especially in the extraordinarily hot summer of 2003. Comparing high and low elevation sites, growth rings were wider at low elevation, but only in cold years. In years with relatively cold summers, however, xylem ring width was greater in plots with late rather than early snowmelt along the natural snowmelt gradient, possibly indicating drought stress in early snowmelt plots. Snow cover had a strong influence on species abundances along the natural snowmelt gradient, and change (not yet significant) was beginning to be seen in plots with 30 years of snow manipulation. Our results indicate that beneficial effects of early snowmelt for shrub growth may be offset in cold summers. Although early snowmelt prolongs the growing season, harsh conditions and frost events early in the growing season may become more likely, and hamper plant growth, and this could affect plant growth in all Arctic and alpine snow-dominated ecosystems.  相似文献   

14.
Six sites between 0 m and 4000 m were sampled for plant and soil chemical characteristics along the Río Loa, Atacama Desert, Chile. Sites located between 0 m and 1500 m showed lower species richness, higher plant cover and higher herbaceous productivity than the upper part of the altitudinal gradient. The number of species varied non-linerly with precipitation along the altitudinal gradient. Plant cover and herbaceous productivity in the lowlands is characterized by thePluchea absinthioides Distichlis spicataassociation of anthropic origin. We propose that vegetation structure along the altitudinal gradient has been affected by past and present human activities, and climatic and edaphic factors.  相似文献   

15.
Camel grazing plays a crucial role in the desert ecosystems of the UAE. In this study, we compare areas grazed by small antelope (Al Maha Resort – the AMR) with areas grazed by both camels and small antelope (Dubai Desert Conservation Reserve – the DDCR). A total of 126 plots were selected during the growing season 2006/07 on three soil substrates: gravel plains, sand flats and sand dunes. In each plot, several vegetation parameters were assessed: density, frequency, percent cover and diversity indices. The replacement of camels with wild antelope has significantly increased the number of species on gravel plains, vegetation density on sand dunes and diversity indices on both sand flats and sand dunes, but significantly decreased plant cover on sand flats and sand dunes. The increase in species diversity in the AMR was attributed to moderate grazing by antelope. Replacement of camels by antelope in the AMR has resulted in change in plant community composition of the three substrate types. Species recovered after protection from camel grazing are palatable, especially for camels, except Heliotropium kotschyi and Aerva javanica. The absence of most of the palatable species from the DDCR was attributed to both selective foraging and overgrazing by camels.  相似文献   

16.
Our study was focused on the effect of abandonment on above-ground biomass and net primary productivity (ANPP) in a Montado in Southern Portugal. The Montado has a long history of human management and control of invasion by shrubs is achieved by clearing, ploughing and grazing. When these cease, it is invaded by Mediterranean matorral species. We hypothesized that the change in life form dominance would affect both biomass and productivity, but while the total biomass was expected to increase, the effects on ANPP were less clear. We tested our hypothesis by determining above-ground biomass and ANPP along a gradient of decreasing land use intensity, ranging from extensive grazing to 20 years of abandonment.Above-ground biomass increased with abandonment, which was related with the increase in shrub cover. In addition, we found a decrease in herbaceous ANPP that was more than compensated by an increase in shrub ANPP in plots abandoned for longer time, resulting in a significant increase in total ANPP. This increase was strongly related with the increase in the cover of Cistus ladanifer, a pioneer species that colonises degraded areas and forms one of the first stages of succession of woody communities.  相似文献   

17.
研究了科尔沁沙质草地放牧和封育下植被盖度、密度、丰富度与地上生物量关系的季节变化。结果表明:放牧和封育草地4月的植被盖度、丰富度和生物量均低于生长旺季8月,而4月植物密度要高于8月;封育草地生长季盖度高于放牧草地,6月和8月生物量高于放牧草地, 4月和6月植物密度则低于放牧草地。封育草地8月物种丰富度与生物量呈极显著的负线性关系(p<0.01);放牧草地植物密度与生物量4月表现出极显著的正线性关系(p<0.01)、8月为极显著的负线性关系(p<0.01)。放牧草地优势植物糙隐子草4月的密度和生物量呈极显著的正线性关系(p<0.01)、8月为极显著的负线性关系(p<0.01);黄蒿和狗尾草8月的密度和生物量均呈极显著的负线性关系(p<0.01);猪毛菜4月和6月的密度和生物量呈极显著的正线性关系(p<0.01)。封育草地达乌里胡枝子、猪毛菜8月的植物密度和生物量呈极显著的正线性关系(p<0.01)。持续放牧和季节变化及其二者的交互作用对植被盖度、密度和生物量均有重要的影响(p<0.05);放牧通过影响草地生长季中的优势植物密度变化和生物量积累,引起了草地植物密度和生物量关系的季节变化,也导致生长季物种丰富度和生物量无显著关系;封育草地植物的竞争导致了物种丰富度和生物量关系的季节变化。  相似文献   

18.
How species diversity–productivity relationships respond to temporal dynamics and land use is still not clear in semi-arid grassland ecosystems. We analyzed seasonal changes of the relationships between vegetation cover, plant density, species richness, and aboveground biomass in grasslands under grazing and exclosure in the Horqin Sandy Land of northern China. Our results showed that in grazed and fenced grassland, vegetation cover, richness, and biomass were lower in April than in August, whereas plant density showed a reverse trend. Vegetation cover during the growing season and biomass in June and August were higher in fenced grassland than in grazed grassland, whereas plant density in April and June was lower in fenced grassland than in grazed grassland. A negative relationship between species richness and biomass was found in August in fenced grassland, and in grazed grassland the relationship between plant density and biomass changed from positive in April to negative in August. The relationship between the density of the dominant plant species and the total biomass also varied with seasonal changes and land use (grazing and exclosure). These results suggest that long-term grazing, seasonal changes, and their interaction significantly influence vegetation cover, plant density, and biomass in grasslands. Plant species competition in fenced grassland results in seasonal changes of the relationship between species richness and biomass. Long-term grazing also affects seasonal changes of the density and biomass of dominant plant species, which further affects the seasonal relationship between plant density and biomass in grasslands. Our study demonstrates the importance of temporal dynamics and land use in understanding the relationship between species richness and ecosystem function.  相似文献   

19.
Responses of herbaceous and suffrutescent species to fire, grazing, and presence of Prosopis glandulosa were examined in a Chihuahuan desert grassland in south-central New Mexico. Treatments were assigned randomly to eight 12×8 m plots within each of two blocks. Following fires in June 1995, unfenced plots were exposed to livestock grazing over 4 years. Plots were established that either included or excluded P. glandulosa. Perennial grass cover, primarilyBouteloua eriopoda , decreased by 13% in burned plots but increased 5% in unburned areas. Conversely, perennial forb cover was 4% greater after fire. Perennial grass frequency decreased 30% more and perennial forb frequency increased 10% more following burning. Further, increases in evenness after fire resulted in a 225% increase in species diversity. Grazing also resulted in a decrease in perennial grass cover while frequency decreased 22% more in grazed than ungrazed plots. Only frequency and not cover of perennial forbs and annual grasses increased more following grazing. Presence of P. glandulosa had no differential effect on responses of non-shrub species. Fires were conducted during near drought conditions while grazing occurred during years of precipitation equivalent to the long-term average. Precipitation immediately following fire may be critical for recovery of B. eriopoda -dominated desert grasslands; relationships between fire and post-fire precipitation patterns require future investigation.  相似文献   

20.
Eleven plant communities of the central Tibetan ecotone (31°20′-33°00′N/89°00′-92°10′E) between the Kobresia pygmaea grazing pastures of the eastern highlands and the open short grass steppe with cushion plants (“Alpine Steppe”) of the north-western highlands have been described on the basis of 189 plot-based floristically complete vegetation records. In parallel, remote sensing techniques distinguish four main land-use cover types. Our hypotheses are: (1) The plant communities of the central Tibetan highlands are grazing-adapted and resilient to degradation. (2) In contrast to grazing resilient plant functional types, the turf cover of the K. pygmaea pastures and the Kobresia schoenoides wetlands is degradable through desiccation, periglacial processes, soil-dwelling small mammals and livestock. Five grazing-related plant functional traits are introduced. Grazing tolerance is the prevalent functional type. Species with no specific protection against grazing make up not more than 6% of the total cover. Unpalatable plants cover up to 8%. Only the azonal K. schoenoides swamps - the indispensable winter grazing reserve - have been widely degraded and depleted by 75%, being replaced by Carex sagaensis grazing pastures. It can be foreseen that governmental policy of sedentarisation of nomads will lead to reduced grazing mobility and degradation of winter grazing reserves.  相似文献   

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