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1.
The well-preserved trackway of a lacertiform, lizard-like trackmaker from the Haman Formation (Cretaceous) of Korea is described as Neosauroides koreaensis ichnogen. et ichnosp. nov. This is the only example of a Cretaceous lacertiform or lizard-like trackway currently known in the global track record. Although lacertiform trackways, mostly assigned to the ichnogenus Rhynchosauroides, are common in the global Triassic, they are almost entirely absent in the Jurassic and Cretaceous. Moreover, ichnological classification criteria allow that Neosauroides is morphologically distinct from Rhynchosauroides at the genus level, and more like the tracks of the extant lizard Sceloporus. The reasons for the conspicuous lack of post-Triassic occurrences are not certain, but not due to a post-Triassic lack of potential lizard trackmakers. Thus, the preservation biases are likely due to paleobiological factors such as trackmaker ecology and paleoenvironmental preference.  相似文献   

2.
Two ichnogenus of “large-sized” ornithopods are found from the Lower Cretaceous (Aptian) Kitadani Formation in central Japan. Caririchnium isp. is characterized by the longer pes print than its width with strong mesaxony. Amblydactylus isp. is characterized by the wider pes print than its length with weak mesaxony. In the northern hemisphere, Caririchnium-type footprints are known from the Berriasian–Cenomanian strata, whereas Amblydactylus-type footprints are known from the Barremian–Maastrichtian strata. It is consistent with the temporal and geographic distribution of non-hadrosauroid iguanodontians and basal hadrosauroids. It suggests that footprint length-width ratio and mesaxony are important factors to indicate trackmakers (basal iguanodontian or hadrosauroid). Two “large-sized” ornithopod ichnogenus from the Kitadani Formation is also consistent with two iguanodontians from same site. It indicates a high diversity of the Kitadani ichnofauna and its importance to elucidate the ecosystem of the Kitadani Formation.  相似文献   

3.
Here we report a large dinosaur tracksite from an extensive fluvial sandstone surface in the Lower Cretaceous Jiaguan Formation of Sichuan Province, China. The site contains over 250 individual tracks comprising at least 18 recognizable trackways, including the longest theropod trackway (cf. Eubrontes) known from China. This exceptional theropod trackway consists of 81 successive footprints covering a distance of 69 m. The tracks are well-preserved and are expressed both as true tracks on the main “upper” surface and as transmitted undertracks on a locally exposed “lower” bed. Also recorded are six other theropod trackways, including small Grallator-like ichnites, eight sauropod trackways (cf. Brontopodus), and three small ornithopod (cf. Ornithopodichnus) trackways with a parallel orientation, which may indicate gregarious behavior. Several trackways of a larger theropod trackmaker show pes imprints with elongated traces of the metatarsals, suggesting extramorphological (substrate-controlled) variation and/or plantigrade posture, which is here interpreted as indicating a change in gait assumed in response to deep and soft sediment. The assemblage indicates a diverse dinosaur fauna in the Lower Cretaceous Sichuan Basin with variously sized theropods, sauropods, and ornithopods. The late occurrence of footprints of the Grallator-Eubrontes plexus in Lower Cretaceous strata is further evidence of the extended stratigraphic range of this morphotype and the distinct palaeobiogeographic distribution of these trackmakers in East Asia.  相似文献   

4.
基于恐龙足迹重建攀西地区白垩纪恐龙动物群   总被引:2,自引:2,他引:0  
攀西地区没有白垩纪恐龙骨骼化石记录,对该区白垩纪恐龙动物群组合的认识尚属空白。1991年,攀西地区首次发现了恐龙足迹,此后陆续发现了8个足迹点。这些足迹点共发现9种不同的非鸟恐龙足迹(6种非鸟兽脚类、1种蜥脚类和2种鸟脚类足迹),以及翼龙和龟类足迹。共11种足迹形态类型由185道行迹(与孤立足迹)组成,可能代表着同等数量的造迹者。其中飞天山组的组合多样性最强,小坝组和雷打树组较弱。这个相对全面的足迹数据库,为该地区白垩纪四足类的古生态学普查提供了动物群的组成信息,这在该缺乏骨骼化石的地区显得尤为重要。  相似文献   

5.
There are a growing number of Early Cretaceous avian tracks and trackways from around the world, with Asia (China and Korea) having the largest reported number and diversity of Mesozoic avian traces to date, and these new discoveries are increasing the Early Cretaceous avian ichnodivesrity of Laurasia. Here we report on a new Lower Cretaceous avian track locality in the Guanshan area, Yongjing County, Gansu Province, northwest China, and on a novel ichnospecies of Koreanaornis, Koreanaornis lii ichnosp. nov. Koreananornis lii is distinct from other Koreanaornipodidae in that it possesses a consistently wider digit divarication than previously described tridactyl tracks, and possess a short, small, posteromedially oriented hallux that displays a different orientation than that seen in Koreanaornis hamanensis. The lack of linear and angular data reported for digit I traces of many avian ichnotaxa has the potential to give misleading results in multivariate statistical analyses. Also, the wide divarication of Koreanaornis lii causes the ichnotaxon to not group with other Koreanornipodidae in multivariate analyses, but with Ignotornidae. Despite the results of the analyses, K. lii is morphologically distinct from these ichnotaxa. The results demonstrate that relying solely on multivariate statistical analyses without careful examination of footprint morphology will result in erroneous ichnospecies groupings. While new vertebrate ichnotaxa discoveries from Asia may support the hypotheses of the presence of a unique and endemic Asian vertebrate ichnofauna during the Cretaceous, the recent discovery of skeletal remains interpreted to be of a volant wading bird from the Early Cretaceous, and recent reports of tracks from volant avians, could suggest that flighted avians of the shore- and wading bird ecotypes could have had a Laurasian-wide distribution during the Early Cretaceous. However, strong convergence in foot morphology of shore- and wading birds suggests that avian ichnotaxa found in both present-day Asia and North America may have been made by birds endemic to eastern and western Laurasia during the Early Cretaceous.  相似文献   

6.
贵州省拥有丰富的中生代脊椎动物化石记录,多为三叠纪海生爬行类化石,侏罗纪和白垩纪的脊椎动物化石较少。目前,贵州省已发现中生代爬行类足迹点9处,分别为贞丰牛场、贞丰龙场、安顺幺铺、赫章辅处、毕节响水(2处)、仁怀茅台、赤水宝源和习水同民足迹点。文中除描述了贞丰龙场新增的一批足迹化石外,整理并总结了所有足迹点的发现史、研究现状、足迹类型与特征以及所反映的古生态信息。已发现的遗迹记录由69条行迹和420个孤立足迹组成,足迹总数达991个。所有的遗迹化石指示了至少477个造迹者,包括海生爬行类、手兽足迹类、非鸟兽脚类、鸟类、蜥脚类和鸟臀类等,其中三叠纪以海生爬行类主导,侏罗纪和白垩纪以蜥臀类主导。这个数据集为贵州中生代古生物类群的多样性提供了重要的信息,是晚中生代稀缺骨骼化石记录的良好补充,为研究贵州省中生代脊椎动物群的古生态与古环境提供了更多的证据。  相似文献   

7.
A pterosaur-bird track assemblage from a sandstone-siltstone-mudstone sequence of the Lower Cretaceous Tugulu Group of Xinjiang comprises the first pterosaur track record from this province and the largest specimen thus far known from China.The pterosaur tracks are assigned to the ichnogenus Pteraichnus based on the triangular overall-shape,the four elongate digit traces and the robust manual digit trace Ⅲ.Supposed trackmakers were dsungaripterid pterodactyloids whose skeletal remains are well known from the Tugulu Group.The bird tracks that occur on the same surface,are those of typical shorebirds,known from different other localities in southeast Asia.The congruence with Koreanaornis dodsoni described from the same stratigraphic level justifies an assignment to this ichnospecies.This is a further evidence of the co-occurrence of pterosaurs and birds in a typical iakeshore environment with possible seasonal alteration of water supply and aerial exposure indicated by wave ripples,mudcracks and repeated cycles of coarse to fine sediment.Pterosaurs and birds frequented the shoreline and may have fed also on the numerous invertebrates such as the Scoyenia tracemaker that left abundant burrows.  相似文献   

8.
The Khok Kruat Formation is the upper part of the Khorat Group, which consists of upper Lower Cretaceous non-marine sedimentary rocks in northeastern Thailand. Many dinosaur footprints have been known from the upper Lower Cretaceous (Aptian–Albian) Khok Kruat Formation at the Huai Dam Chum (Tha Uthen) site, northeastern Thailand. Approximately 600 tracks occur in thin mudstone layer of the northern part of the outcrop at the Huai Dam Chum track site. Two types of footprints, small-sized theropod and crocodylomorph are imprinted with mud cracks and ripple marks on the thin mud layer. Most of footprints are referred to cf. Asianopodus, and are imprinted by small-sized theropoda, probably ornithomimosauria. Theropod tracks are mainly separated into two groups, Group A and Group B. From ichnological viewpoints, the small-sized theropod track assemblage indicates the herd behaviour and its idiosyncratic group composition. In particular, the histogram of size-frequency measurements of Group A shows the anomalous bimodal distribution. We consider that there are two hypotheses; the first one is due to the male-female difference, and the second is a result of the different growing stage.  相似文献   

9.
A new Lower Cretaceous (lower Aptian) dinosaur tracksite, from the eastern side of Monte Cagno (Abruzzi, Italy), is described. Different styles of track formation are represented on the site surface. Most of the footprints are preserved as deep tracks, produced by trackmakers sinking into soft mud. Some tracks, better preserved than the others, are characterized by metatarsal impressions and were interpreted as the resting traces of a crouching theropod (based on their orientation and three-dimensional morphology). The 135 cm length of the track with metatarsal impressions indicates huge pedal proportions and represents the largest theropod trackmaker ever documented from the Mesozoic peri-Adriatic platforms of Italy.  相似文献   

10.
Three-dimensional tracks provide unique insights into the locomotor mechanics of their track makers. An isolated, large hadrosauriform print attributable to Caririchnium lotus from the “mid”-Cretaceous Lotus track site (Jiaguan Formation) in China permits reconstruction of the footfall, weight-bearing, and kick-off phases of the step cycle. Large-scale modifications of the pes during the step cycle indicate C. lotus trackmakers were capable of locomotory modifications in response to substrate consistency beyond the “expected” shift between bipedal and quadrupedal postures. An unusual curvature to the trace of one of the outer digits indicates substantial transverse mobility. The remaining digits demonstrate lesser degrees of transverse movement accompanied by extension of the digits during footfall. The absence of overprinted scale-scratch marks and toe drags are consistent with a vertical kick-off of the pes and concomitant flexion of the digits. This track suggests that pedal mobility in C. lotus track makers was greater than previously suspected and has implications for reconstructions of hadrosauriform locomotion.  相似文献   

11.
Ornithuromorph birds (the clade which includes modern avian radiation) first appeared in the Early Cretaceous in Asia and achieved a great diversity during the latest ages of the Late Cretaceous (Campanian and Maastrichtian). The evolutionary history of orithuromorphs during the first 17 MYAs of the Late Cretaceous (Cenomanian to Santonian ages) remains very poorly known, as the fossil record for this time interval is largely restricted to several isolated finds of the classic avian genus Ichthyornis in North America. Here we describe an isolated distal tibiotarsus of an evolutionary advanced bird, morphologically similar to Ichthyornis, from the middle Cenomanian of Saratov Province, European Russia. This is the first documentation of an Ichthyornis-like bird in the Old World. The find further constitutes only the second pre-Campanian record of the Late Cretaceous Ornithuromorpha in Eurasia, the second record of Cenomanian birds in Russia. This discovery shows that Ichthyornis-like birds enjoyed a wide geographical distribution as early as the beginning of the Late Cretaceous. Given that the earliest and the most primitive ornithuromorph birds are known from Asia, the new find supports a Eurasian origin for Ichthyornithidae.  相似文献   

12.
Cretaceous dinosaur tracks from Hunan Province are historically significant as the basis for three ichnotaxa: Xiangxipus chenxiensis, Xiangxipus youngi, and Hunanpus jiuquwanensis all representing theropodan tracks, described from a single site in 1982. Although the type locality has since been destroyed, the type specimen and replicas remained available for restudy in 2006, when paratype Hunanpus tracks and sauropod footprints were described from a second, nearby locality. Material from both localities is here re-described in detail. It is proposed that while Xiangxipus chenxiensis can be regarded as a distinct ichnospecies, probably representing an ornithomimid trackmaker, Xiangxipus youngi cannot be accommodated in the same ichnogenus. We consider it similar to the ichnogenus Wupus from the Lower Cretaceous of Sichuan Province, and therefore of possible avian affinity. We also find no justification for regarding Hunanpus jiuquwanensis as generically distinct from the widespread ichnogenus Grallator, and therefore rename it Grallator jiuquwanensis comb. nov. The Hunan track record is generally consistent with the sparse record of theropod skeletal remains in the province, but adds evidence of sauropods that was previously lacking.  相似文献   

13.
14.
Rapid radiation of the Bittacidae during the Jurassic resulted in high diversity within this family of Mecoptera. More than 40 species within 23 genera have been described from this period. However, around the end of the Jurassic the abundance and diversity of Bittacidae decreased and only seven species in five genera are known to date from the Cretaceous. A new specimen from the basal Cretaceous of England, Tytthobittacus jarzembowski sp. nov., described here, represents the eighth species and the second fossil representative of this family from the European Cretaceous. This hangingfly belongs to an extant and relictual genus previously known only from Australia. The paper also includes a review of all known Cretaceous bittacids and a re-examination of European species Antiquanabittacus nanus Petrulevičius and Jarzembowski, 2004.  相似文献   

15.
The Cretaceous witnessed the greatest diversity of Gondwanan notosuchian crocodyliforms, which displayed high levels of diversity and a notable array of specialized forms that developed in different ecological niches. Among this diversity, the advanced notosuchians are a clade of mid body sized forms which displays a remarkable abundance but is restricted to two lithological units from the Late Cretaceous of South America, the Adamantina Formation in southeastern Brazil and the Bajo de la Carpa Formation in Patagonia (Argentina). The only exceptions known so far were the Late Cretaceous Labidiosuchus from the Marilia Formation in Brazil and Yacarerani from the Cajones Formation in Bolivia. Herein we report a new Cretaceous crocodyliform, Llanosuchus tamaensis gen. nov. et sp. nov., found in the Los Llanos Formation (Campanian?) in northwestern Argentina (La Rioja Province). The small specimen includes well preserved fragments of the cranium and mandible with an estimated skull size of about 9 cm lengths. This crocodyliform shares several derived characters with Notosuchus terrestris from the Bajo de la Carpa Formation (Patagonia) and it was found in a weakly developed sandy paleosol profile formed in wet local conditions but in a region characterized by semi-arid climate and active eolian sedimentation. The presence of a new advanced notosuchian in the Cretaceous of western Gondwana, and its intermediate geographical region between the known South American species (Brazil, Argentina and Bolivia), has interesting implications, and adds another record of an advanced notosuchian in deposits with inferred warm climates and semi-arid paleoenvironments. Finally, Llanosuchus tamaensis supports a Late Cretaceous age for Los Llanos Formation with important geological implications for La Rioja Province.  相似文献   

16.
The Bajiu tracksite exposed on a fine sandstone surface at the top of the Feitianshan Formation, close the contact with the overlying Xiaoba Formation, reveals a saurischian dominated ichnofauna consisting of four sauropod and at least eight theropod trackways. The sauropod trackways display a medium-wide gauge pattern characteristic of titanosauriforms, while the theropod trackways can be subdivided into two distinct morphotypes. One is similar to the ichnogenera Eubrontes and Megalosauripus, the other (cf. Dromaeopodus) appears to represent a functionally didactyl dromaeosaur where pedal digit II is represented only by an oval basal pad. A single swim track is possibly attributable to a turtle or crocodylian. Such sauropod-theropod dominated ichnofaunas are consistently typical of the red-bed tracksites in the region and useful for characterizing a fauna that is otherwise poorly known from body fossils. The stratigraphic position and late occurrence of the (typical Jurassic) Eubrontes-Megalosauripus morphotype is well known from other Cretaceous localities in China and seems to reflect a peculiarity in the theropod faunas of East Asia.  相似文献   

17.
Although the Cretaceous is characterized by a rich fish diversity, Cretaceous continental fishes from Gondwana are poorly known and comparatively scarce. Among these fishes, the family Pleuropholidae is only known by a few species relatively poorly preserved, from the Middle Jurassic to Lower Cretaceous of Europe, Africa, North America, and South America. In this paper, two new species of the pleuropholid new genus Zurupleuropholis are described, Z. quijadensis gen. et sp. nov. and Z. decollavi gen. et sp. nov. The new fishes were recovered in the Lower Cretaceous lacustrine Lagarcito Formation of central-west Argentina. This taxon constitutes a relevant finding considering that the representation of the family Pleuropholidae is rare worldwide. Zurupleuropholis gen. nov. appears to be the youngest known member of Pleuropholidae, and it represents the second record of the family in South America and the first record in the Cretaceous of the continent.  相似文献   

18.
We describe a large collection of fossil ‘waterfowl’ bones that are referable to the extinct clade Presbyornithidae (Anseriformes). All of these fossils were collected between 1971 and 1994 from Upper Paleocene and Lower Eocene sediments at the Tsagaan Khushuu site in the Gobi Desert of southern Mongolia. The collection includes specimens referred to a new small species within the genus Presbyornis Wetmore, 1926 as well as large numbers of bones that we place in the genus Presbyornis. On this basis of the Tsagaan Khushuu collection we suggest that several species of Presbyornis likely coexisted in this region; indeed, the presence of large numbers of middle‐sized, morphologically consistent but probably ecologically disparate species at the Tsagaan Khushuu site is consistent with the range of variation seen, for example, in taxa of extant dabbling ducks (Anatini). Although the anatomy and phylogenetic position of Presbyornithidae (in particular Presbyornis) are well known, this material from Mongolia further demonstrates the prevalence of these birds in aquatic and semi‐aquatic habitats by the earliest Paleogene. Because presbyornithids are also well documented from the late Cretaceous, their palaeoecology and morphological diversity provides a clue to selective avian survivorship across the Cretaceous‐Paleogene (K‐Pg) boundary. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

19.
A Yanji Basin tracksite in the Cretaceous Tongfosi Formation in the Yanbian Korean Autonomous Prefecture, Jilin Province was reinvestigated twenty years after the original study.The re-examination confirms the presence of tracks tentatively referred to ornithopods and theropod tracks including several not previously reported. In addition to the main track-bearing level, a second footprint horizon was identified from which a probable ornithopod track was collected. Some of the formerly identified tracks were documented using photogrammetry, whereas others that had been collected, eroded, buried or obscured by dense vegetation. The age of the track-bearing beds is inferred to be late Albian to early Cenomanian. Comparison with track-bearing units of the same age in Shandong, China, and Korea indicated quite different compositions of track assemblages. In the case of the Korean sites the differences may be facies related. While the Korean ichnofauna is characterized by the co-occurrence of ornithopod, sauropod and tridactyl theropod tracks with didactyl dromaeosaurid, Minisauripus (small theropod), pterosaur and avian tracks, the latter four are absent in the Tongfosi assemblage. This could reflect a restricted environment being not favourable for activities of these trackmakers or simply a preservational bias.  相似文献   

20.
More than 125 footprints of theropods from the Cretaceous Longwangzhuang Formation have been mapped in a preliminary study at a site in the Zhucheng region of China. The tracks represent at least three morphotypes. The largest morphotype is a large theropod (footprint length ∼30 cm) represented by a single trackway and an isolated natural cast. At least 10 trackways assigned to the new ichnospecies Corpulentapus lilasia represent a medium-sized biped (footprint length ∼13 cm) with very short, wide, robust, ‘tulip-shaped’ tracks and long steps (∼5 × footprint length), and a short central digit (III) indicating weak mesaxony. Corpulentapus trackways are narrow and theropod-like even though track morphology is convergent with the footprints of some ornithopods. The third morphotype, made by a medium-sized grallatorid track maker (ichnogenus Paragrallator), is about the same size (∼13 cm) as the robust morphotype, but far more elongate and gracile, with an elongate central digit (III) indicating strong mesaxony. This ichnotaxon requires detailed comparison with Grallator sensu stricto. The contrast in morphology between the two common morphotypes is striking and demonstrates that two distinct medium-sized taxa of presumed theropod affinity frequented the same habitat in significant numbers.  相似文献   

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