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1.
A new genus and species of the extinct beetle-like dictyopteran family 2Umenocoleidae, 2Laticephalana liuyani gen. et sp. nov., is described based on a well-preserved adult female from mid-Cretaceous Kachin (Myanmar/Burmese) amber. The new genus is characterized by two conspicuous autapomorphies, a triangular posterior extension of the head capsule and four large protuberances of the pronotum. Both structures can be interpreted as mechanic protective devices, but alternative/additional interpretations are possible. The large female subgenital plate is an autapomorphy of Dictyoptera. The transverse pronotal furrow suggests a placement in a clade with 2Alienopteridae and Mantodea. A close relationship between 2Laticephalana and 2Enervipraela is tentatively supported by a very narrow pronotum and relatively short antennae. The very broad head of 2Laticephalana with widely separated large compound eyes indicates predaceous habits, and the slender body and well-developed hindwings good flying abilities. The specializations of the distal legs, very similar to the conditions found in 2Alienopteridae, suggest a preference for the foliage of trees and shrubs.  相似文献   

2.
A new species of the foraminiferal genus Pseudorhapydionina is reported from the Santonian shallow-water carbonate and mixed deposits of La Cova Unit, in the Montsec and Pedraforca thrust sheets (Southern Pyrenees, NE Spain). Pseudorhapydionina bilottei sp. nov. differs from the Cenomanian species of the genus in its larger test size and the number of chambers in its early planispiral-involute stage. The discovery of P. bilottei sp. nov in the Santonian indicates that the genus Pseudorhapydionina escaped extinction during the Cenomanian-Turonian boundary (CTB) event in the Pyrenean bioprovince. A new subfamily Pseudorhapydionininae is described.  相似文献   

3.
A new large theropod, Orkoraptor burkei nov. gen. et nov. sp., from the Upper Cretaceous (Maastrichtian) Pari Aike Formation of southern Patagonia is based on a postorbital, quadratojugal, coronoid?, several teeth, an atlantal intercentrum and neurapophysis, two caudal vertebrae, and the proximal half of tibia. This new theropod exhibits characteristics of maniraptorans (i.e., coelurosaurians more derived than tyrannosaurids) including an upturned rostral process of the postorbital, the caudoventral corner of which is widely concave. Furthermore, teeth without denticles and carina in the mesial margin resemble the condition of the maniraptoran clades Compsognathidae and Deinonychosauria. The new Patagonian theropod differs from other coelurosaurians (Ornithomimosauria, Compsognathidae, Alvarezsauridae, Dromaeosauridae, Aves) in having caudal vertebrae with a single pair of small pleurocoels on each side, and in having a median depression flanked by two longitudinal and narrow furrows on each tooth. Orkoraptor represents one of the southernmost carnivorous dinosaurs yet found in South America and adds valuable information about the diversification of tetanuran theropods in Gondwana.  相似文献   

4.
5.
A new subfamily, Burmadysagrioninae Zheng, Wang and Nel, subfam. nov., for the genus and species Burmadysagrion zhangi Zheng, Wang and Nel, gen. et sp. nov. is described from mid-Cretaceous Burmese amber. This is the third dysagrionid damselfly from the Mesozoic and the second one from Burmese amber. The new specimen has a unique discoidal cell with the anterior and posterior sides not parallel, and the basal side longer than the distal side, unlike the typical ‘sieblosiid-dysagrionine’ type. It differs from other dysagrionid damselflies by the presence of a simple wing venation, the vein IR1 originating below the pterostigma and a special discoidal cell.  相似文献   

6.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight.  相似文献   

7.
Oxyporinae is a distinct staphylinid subfamily characterized by a large body, remarkably long and projecting mandibles, and large, crescent-shaped terminal labial palpomeres. Previously, only three compression fossils from the Lower Cretaceous Yixian Formation in China were known for the subfamily from the Mesozoic Era. This study describes a remarkable new species, Oxyporus cretaceous sp. nov., based on a single specimen in Upper Cretaceous Burmese amber. This finding is the earliest record of Oxyporinae preserved as amber inclusion. The well preserved mouthparts suggest that mycophagous feeding habits had already been acquired by the mid-Cretaceous.  相似文献   

8.
A new genus and species of the ant-like stone beetle supertribe Mastigitae, Cretoleptochromus archaicus gen. et sp. nov., is described and figured based on an exceptionally well-preserved individual in the Upper Cretaceous (lowermost Cenomanian, ca. 99 Ma) amber from Myanmar. Cretoleptochromus is undoubtedly placed in Mastigitae based on its distinctly geniculate antennae with strongly elongate antennal scape, and pointed labial palpomere III. Cretoleptochromus is most likely the sister taxon to the extinct Palaeoleptochromus O'Keefe from the Late Cretaceous Canadian amber. Cretoleptochromus shares with Clidicini the distinct posterior collar of pronotum; with Leptomastacini the presence of mesocoxal lateroventral bristles. The new discovery may help to elucidate the relationships among the Recent three tribes, particularly for Leptomastacini and Clidicini.  相似文献   

9.
The Cretaceous new species and genus Albocryptophagus cantabricus gen. et sp. n. is described based on a fossil specimen from the El Soplao amber deposit (Spain). The new genus is similar to the extant genera Cryptophagus and Micrambe, but differs from them in the transverse pedicel, anterior angles of pronotum, pronotal margin unmodified, pronotal pits absent. Albocryptophagus gen. n. is undoubtedly the most ancient representative for the subfamily Cryptophaginae described up today. Because the saproxylic habits in recent species of the family, a similar behavior is inferred for this new fossil. It is a new example that agrees with the idea that fossil beetles from Mesozoic ambers are characterized by a saproxylic lifestyle.  相似文献   

10.
11.
Cretoboganium gei gen. et sp. nov., a new amber inclusion of the cucujoid family Boganiidae is described and figured based on a well-preserved adult from the mid-Cretaceous Burmese amber (Hukawng Valley, northern Myanmar), some 99 million years ago. Based on the presence of a pair of pronotal callosities, Cretoboganium can be firmly placed in the extant subfamily Boganiinae, a small group currently comprising two small austral genera. Our discovery represents the first fossil record for Boganiinae. It also demonstrates another example that an apparently austral group may have its sister group occurred in today’s northern hemisphere. Together with the other fossil boganiid known from the Middle Jurassic of China, the finding suggests that Boganiidae is an ancient and relict group. Moreover, the present biogeographic distribution of Boganiinae is indicative of an earlier origin of this subfamily, which likely originated before the breakup of the Gondwanan supercontinent.  相似文献   

12.
A new genus and species, Gracilipygia canaliculata gen. et sp. nov., in the family Pygidicranidae of Dermaptera is described from the Upper Cretaceous Myanmar amber. The new species is assigned to the subfamily Pyragrinae mainly based on the following characters: broadly separated cercal forceps without segmentation and the posterior abdominal tergum with mediolongitudinal furrow and lateral ridges. The G. canaliculata gen. et sp. nov. is the earliest record hitherto for the subfamily Pyragrinae, and probably represents a stem taxa of Pyragrinae.  相似文献   

13.
The Upper Cretaceous shallow-water carbonates of the Pyrenean Basin (NE Spain) host rich and diverse larger foraminiferal associations which witness the recovery of this group of protozoans after the dramatic extinction of the Cenomanian–Turonian boundary interval. In this paper a new, large discoidal porcelaneous foraminifer, Broeckina gassoensis sp. nov., is described from the middle Coniacian shallow-water deposits of the Collada Gassó Formation, in the Bóixols Thrust Sheet. This is the first complex porcelaneous larger foraminifer of the Late Cretaceous global community maturation cycle recorded in the Pyrenean bioprovince. It differs from the late Santonian–early Campanian B. dufrenoyi for its smaller size in A and B generations and the less developed endoskeleton, which shows short septula. Broeckina gassoensis sp. nov. has been widely employed as a stratigraphic marker in the regional geological literature, under the name of “Broeckina”, but its age was so far controversial. Its middle Coniacian age (lowermost part of the Peroniceras tridorsatum ammonite zone), established in this paper by strontium isotope stratigraphy, indicates that it took about 5 My after the Cenomanian–Turonian boundary crisis to re-evolve the complex test architecture of larger foraminifera, which is functional to their relation with photosymbiotic algae and K-strategy.  相似文献   

14.
《Gondwana Research》2016,29(4):1579-1584
The subfamily Aleocharinae is a hyper-diverse group of Staphylinidae, accounting for nearly one-third of the diversity within the largest animal family on Earth. Here we describe Cretodeinopsis aenigmatica gen. et sp. nov., the first definitive Mesozoic aleocharine based on a well-preserved individual in the earliest Late Cretaceous (Cenomanian) amber from Myanmar (Burma). The new genus is clearly assigned to the ‘basal’ group of Aleocharinae based on its overall body shape, antennal insertions on the vertex of the head, and emarginate posterolateral margins of elytra. It is placed in the Recent tribe Deinopsini based on the strongly deflexed head, pectinate edge of abdominal segments III–V, tarsal structures, and non-pentamerous tarsi. The new discovery of the oldest known Aleocharinae from about 99 Mya represents a new aleocharine belonging to the ‘basal’ group of Aleocharinae, reinforcing the hypothesis that Gymnusa, Stylogymnusa and the tribe Deinopsini probably form a monophyletic lineage and suggest that the monophyly of Gymnusini is not supported based on the presence of 6 setigerous pores on the head. It indicates that the origin of the subfamily probably predates the Cenomanian. This new discovery is about 55 Ma older than the previously known Eocene-aged ones from Baltic amber, greatly broadening our knowledge on the early evolution of Aleocharinae.  相似文献   

15.
The holotype of ‘Procheneosaurusconvincens, a juvenile lambeosaurine, is the most complete hadrosaurid known from Kazakhstan. North American species of Procheneosaurus are considered juveniles of Corythosaurus, Hypacrosaurus, and Lambeosaurus, rendering the generic name unusable. A replacement name, Kazaklambia convincens comb. nov., is herein proposed as this specimen is morphologically distinct from other Eurasian taxa and known juvenile lambeosaurines at a similar ontogenetic stage in having a prefrontal process of the postorbital with a dorsal thickening forming a dome lateral to the frontal dome, doming of the nasal anterodorsal to the orbit, and a frontal length/width ratio <1. The juvenile status of Kazaklambia makes phylogenetic placement difficult; however, morphometric and morphological information (particularly in relation to the hollow cranial crest and the length of the frontal) suggest a close affiliation with the basal lambeosaurines Amurosaurus and Tsintaosaurus, and support the hypothesis for an Asian origin for Lambeosaurinae.  相似文献   

16.
The dysagrionid damselflies are relatively diverse in Burmese amber, with two genera already recorded. A new dysagrionid damselfly, Electrodysagrion lini Zheng, Nel and Wang, gen. et sp. nov., corresponding to the oldest record of the tribe Dysagrionini, is described herein. It has the unique ‘sieblosiid-dysagrionine’ type of discoidal cell. It differs from other genera of Dysagrionini in having no antenodal crossveins distal of Ax2, Arc aligned with Ax2, and only one row of cells in the cubito-anal area below the nodus. This find puts the origin of Dysagrionini back to at least the mid-Cretaceous.  相似文献   

17.
An exotic wingless female insect in mid-Cretaceous Myanmar amber is described as Aethiocarenus burmanicus sp. et gen. nov. in the order Aethiocarenodea ord. nov. and family Aethiocarenidae fam. nov. The distinguishing feature of Aethiocarenus burmanicus sp. et gen. nov. is its unique head, the dorsum of which is shaped like an isosceles right triangle with the hypotenuse at the top and vertex positioned at the base of the neck. While insects with triangular-shaped heads are common today, the hypotenuse of the triangle is always located at the base of the head and attached to the neck, with the vertex at the apex of the head. Other features of the fossil are the long narrow, flat body, long slender legs, especially the hind pair that are twice the length of the abdomen, lack of wings, protruding eyes, paired ocelli, secretory glands located on the dorsum of the neck and swollen abdomen bearing paired segmented cerci.  相似文献   

18.
A new cerambycid beetle (Qitianniu zhihaoi gen. et. sp. nov.) is described on the basis of a single specimen embedded in Cretaceous Burmese amber (ca. 99 Ma). Unusual characteristics are hairy antennae and large lateral eyes, a pronotum with lateral margin, and sinuate protibiae. Based on a phylogenetic analysis, the systematic position of Qitianniu is still uncertain and we provisionally place it as Cerambycidae incertae sedis.  相似文献   

19.
A new genus and species of Scoliidae (Hymenoptera), Sinoproscolia yangshuwanziensis gen. et sp. nov., is described from the Lower Cretaceous Yixian Formation in Inner Mongolia, China and attributed to the subfamily Proscoliinae. Some characteristics, including complete crossveins 1r-rs and 2r-m in the forewing, and a free section present in almost all longitudinal veins indicate a basal position of the new genus in Proscoliinae. The presence of vein 2A suggests its atavistic origin in Sinoproscolia gen. nov. rather than a true plesiomorphy.  相似文献   

20.
Cretadilar olei gen. et sp. nov. and Burmopsychops groehni sp. nov. (Neuroptera: Dilaridae) are described from the Late Cretaceous (earliest Cenomanian/late Albian) Burmese amber. Their dilarid affinities are supported by two autapomorphies of the family: the reduction of the ligula and the presence of pectinate antennae in males. They are considered to belong to two new subfamilies, Cretanallachiinae subfam. nov. and Cretadilarinae subfam. nov., respectively. These subfamilies are easily distinguished from other dilarids (Dilarinae, Nallachiinae, and Berothellinae) by possessing siphonate mouthparts; however they strongly differ from each other. In particular, the broad gonocoxites 9 of Cretanallachiinae females are not fused to form a distinct ovipositor, and bear long gonostyli, whereas a distinct saber-like ovipositor (short, lacking gonostyli) is present in Cretadilarinae. It is assumed that Cretanallachiinae primarily fed on pollen and Cretadilarinae on the nectar of flowers.  相似文献   

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