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1.
Spatial and seasonal variations of fish assemblage composition were studied in three non-estuarine mangrove creeks of Zanzibar (Tanzania). Fish were collected monthly for one year at three sites (lower, intermediate and upper reaches) in each creek using a seine net (each haul covering 170 m2). Density, species number and diversity of fish were all higher at sites with dense cover of macrophytes (seagrass and macroalgae) than over unvegetated sandy sites. In general, fish assemblages mainly comprised juveniles of a few abundant taxa, e.g. Mugil cephalus, Mugilidae spp. and Leiognathus equulus at sites with mud substratum and Gerres oyena, Lethrinus harak and Sillago sihama at sites dominated by macrophytes. Multivariate analyses revealed significant separations in fish assemblage composition within the two creeks where the bottom substratum differed among sites. Overall, season seemed to have little effect on density, species number, diversity index (H′) and assemblage structure of fish. Water condition variables were also relatively stable across the season, although a short-term fluctuation primarily induced by decreased salinity, occurred during the heavy rains in April and May. Fish assemblage structure was not significantly affected by any of the abiotic factors tested. However, significant regressions were found between the other fish variables and environmental variables, but since these associations were mostly species-specific and generally inconsistent, we suggest that the overall distribution patterns of fish were mainly an effect of particular substrate preferences of fish species rather than contemporary water conditions.  相似文献   

2.
The fish assemblage structure was analyzed along an estuarine gradient of a small macrotidal estuary (the Canche, France). Fishes were collected every two months between May 2006 and July 2007 from 12 sampling stations using a 1.5-m beam trawl with a 5 mm mesh size in the cod end. To complement this information, sampling was also performed using 15-m fyke nets (8 mm mesh size in the cod end). For each sample, abiotic (temperature, salinity, pH, oxygen, turbidity, river flow, wind speed and depth) and biotic (macro crustacean species abundances) were recorded. Throughout the study, 28 fish species belonging to 20 families were collected. Fish catches were dominated by juveniles, especially Young-Of-the-Year (YOY) for the majority of the species. According to the Index of Relative Importance (IRI), common goby Pomatoschistus microps, flounder Platichtys flesus, sprat Sprattus sprattus, sea-bass Dicentrarchus labrax and plaice Pleuronectes platessa were the most abundant species, together accounting for 99.2% of the total IRI. Estuarine residents (ER = 66.2%) and marine juvenile migrants species (MJ = 31.4%) were the most important ecological guilds. The structure of the fish assemblage and its relationship to environmental variables was examined using multivariate techniques. Cluster and non-metric multidimensional scaling (nMDS) analysis defined six distinct groups in the Canche estuary, which are discriminated by specific species (SIMPER). Spatio-temporal variations in fish assemblage structure reflect the density peaks of the most abundant species. Spearman rank correlations and canonical correspondence analysis (CCA) showed that among the ten environmental variables examined, temperature, salinity and Crangon crangon (a potential predator for YOY fish or prey for older ones) are the three most important factors influencing fish species richness and abundances. Our observations reinforce the idea that certain fish species may have different life history styles in different geographic areas. The present study highlights the necessity of a better knowledge of the connectivity between estuaries and adjacent marine areas. The Canche constitutes an important ecosystem for fishes and as it is subject to little anthropogenic disturbance; its ichthyofauna can be viewed as a reference or normal assemblage for small temperate macrotidal estuaries.  相似文献   

3.
Despite substantial survey effort and a large body of literature on abiotic and biotic factors in temperate reef ecosystems, knowledge of the complex and interactive effects of environmental variables on those communities is limited. Various survey methods have been developed to study environmental predictors of biodiversity, but there remains a gap in our understanding of how survey results are influenced by environmental factors. Here, we surveyed the fish assemblage associated with southeastern U.S. temperate marine reefs with simultaneous, paired trap, and camera gears throughout a ~50,000 km2 area during 2011–2013 and assessed the influence of environmental variables on the trap‐ and video‐surveyed assemblages. Predictor variables in the multivariate general linear models included depth, temperature, month, year, location, substrate relief, percent sessile biota, biota type, and turbidity. Depth and latitude had the greatest influence on the fish assemblage for both gears. The influence of habitat variables differed between methods and percent biota explained more variation in the fish assemblage when assessed by traps, while substrate relief and biota type explained more variation in the fish assemblage when assessed by video. In general, habitat complexity was positively related to the abundance of fishes in the video survey, but there was a negative relationship in the trap survey. Differences between gears were species‐specific and the influences of environmental variables were similar for some species such as Haemulon plumierii and Hyporthodus niveatus. The methods presented here can be used to assess method‐dependent differences in fish assemblages, which is a necessary precursor to assess the effect of environmental variables on the accuracy of surveys.  相似文献   

4.
徐勇  马林  李新正  孙悦  龚琳 《海洋与湖沼》2017,48(6):1383-1391
为了研究春季长江口外海底层鱼类群聚特征及其与环境因子的关系,我们根据2015年5月长江口外海底层鱼类的调查资料,使用聚类分析(Cluster)、非参数多维标度排序(NMDS)、相似性分析(ANOSIM)、相似性百分比分析(SIMPER)以及典型对应分析(CCA)等方法对资料进行分析。本研究共记录底层鱼类58种,其中鲈形目种类数最多(21种),鲽形目次之。六丝钝尾虾虎鱼(Amblychaeturichthys hexanema)是优势种。底层鱼类可以划分为3个群组—近海组、南部中间组和南部远海组,不同群组的物种组成差异显著。南部中间组和南部远海组的物种多为东海外海种类,这可能是近岸黑潮底层分支影响的结果。物种数、Margalef丰富度、Shannon-Wiener多样性和Pielou均匀度从近海到与远海呈逐渐增加的趋势。CCA分析表明深度是显著影响底层鱼类的环境因子。短鳄齿鱼(Champsodon snyderi)、丝鳍(Repomucenus virgis)、多棘腔吻鳕(Coelorinchus multispinulosus)等东海外海种类与深度呈正相关,而优势种六丝钝尾虾虎鱼受环境因子影响较小。  相似文献   

5.
This study has analysed for the first time fish composition and assemblage structures of three small macrotidal estuaries of the Eastern English Channel (EEC) and has explored the influences of 19 biotic and abiotic variables on the fish assemblages. Fish from Canche, Authie and Somme estuaries were collected during spring (June 2006 and May 2007) and autumn (September 2006) along the estuarine gradients using a 1.5 m beam trawl. Using identical sampling protocols, the study also analysed and compared for the first time taxonomic and functional aspects of the fish assemblages in 15 estuaries located along the Atlantic and English Channel coasts. SIMPER analysis showed high similarities in fish assemblages in the three EEC estuaries and during either spring or autumn periods. However, intra-estuary similarities were relatively low, indicating that fish assemblage structures (species richnesses or abundances) were more variable within the estuary (salinity gradient) than between estuaries and/or seasons (spring vs autumn). Although numerous environmental variables were included in the study, only 47% of the variability observed in the fish distribution was explained. Fish spatial variations in the EEC estuaries are mostly driven by abiotic variables as opposed to biological interactions. As indicated by CCA, salinity and muddy sediments were the two most important factors structuring the fish assemblages. The macrobenthos being very abundant in the EEC estuaries (580–1121 ind. m−2), the availability of potential prey is probably not a limiting factor in the utilization of estuaries by fish. Contrary to the majority of French estuaries dominated by estuarine species (ES), the fish assemblages of the EEC estuaries are clearly dominated by marine migrant (MM) species (65% on average) with high abundance of juveniles (mostly young-of-the-year). Cluster and SIMPROF's analyses distinguished the functional structure of the 15 estuarine fish assemblages into different clusters. The three EEC estuaries form a similar group with the largest French estuaries (Seine, Loire and Gironde). The latters were characterized by higher MM/ES ratio compared to the other French estuaries. Freshwater flow and tidal range were the main factors associated with this group of estuaries. The results of the present study do not indicate that anthropogenic impacts (chemical contamination) on estuaries affect their ecological functioning as described by the fish ecological guilds. It is suggested that the guild approach may not be useful to provide valuable information on the ecological status of estuaries.  相似文献   

6.
东海中南部鱼类群聚结构的空间特征   总被引：7，自引：0，他引：7  
依据1997～2000年东海中南部(25°30'～29°00'N,120°30'～127°00'E)四个季节的底拖网调查资料,运用聚类分析和非度量多维标度(NMDS)的方法分析了东海中南部鱼类群聚结构的空间特征,结果显示,东海中南部鱼类群聚大致可分为近海群聚和外海群聚.在不同季节两个群聚的种类组成保持了相对的稳定,并且具有一定的持续性.近海群聚的特征种类为带鱼、发光鲷等.外海群聚的特征种类主要为黄鳍马面鲀、短尾大眼鲷、黄鲷等.鱼类群聚与环境因子水深有较好的相关关系.  相似文献   

7.
Environmental variables have been measured and sampling for ichthyoplankton has been conducted monthly, since April 2001, at three stations, located at the inner (1), middle (2) and outer (3) shelf of the central Cantabrian Sea. This paper presents the results of the study of the ichthyoplankton collected from July 2001 to June 2004. Fish larvae from 99 species, belonging to 37 families, were identified. Families with higher number of species were Gadidae, Sparidae and Labridae. The larval fish assemblage was dominated by pelagic fish species, with Sardina pilchardus, as the most abundant. There was a pronounced spring peak in larval abundance, dominated by S. pilchardus. A smaller peak, dominated by S. pilchardus and Micromesistius poutassou, was recorded in late winter at Stns 2 and 3. This pattern was evident for the three-year study. Results also indicate that this study was limited to the coastal larval fish assemblage inhabiting the central Cantabrian Sea shelf. This assemblage was temporally structured into other three assemblages: winter, late winter–spring and summer–autumn. Temperature was apparently a key factor in larval fish assemblage succession. In a scenario of global warming, this study constitutes a basis to evaluating the ongoing changes in the pelagic coastal ecosystem of the central Cantabrian Sea.  相似文献   

8.
The microhabitat preferences and depth distribution of blenniid species (Blenniidae) in the Gulf of Trieste (North Adriatic Sea) were surveyed using the all‐occurrence sampling method, a non‐destructive visual census method, aided by SCUBA diving. Fourteen species were identified during the entire survey. Four species showed to be indiscriminate in their microhabitat choice while 10 species were classified as infrequent in the surveyed coastal area. A statistically significant correlation was found between the blenniid assemblage and nine microhabitat variables. The blenniid assemblage was divided in two main groups, by the use of canonical correspondence analysis, electivity index, and the depth distribution analysis. The first group comprises species that dwell in surface waters and show a high positive correlation with boulders, the presence of Mytilus galloprovincialis, cirripeds and empty holes bored by Lithophaga lithophaga. The second group includes species that mostly inhabit deeper waters and show a high positive correlation with rocks covered by precoralligenous bioformations.  相似文献   

9.
To assess differences in fish assemblage structures among microhabitats within the surf zone of an exposed sandy beach, three microhabitats (onshore current, rip current, and river adjacent sites) were sampled at Sanrimatsubara Beach, western Japan, in May, August, and November 2002 and 2003. Although several physical variables (e.g. wave height, current speed, turbidity, salinity, and sand particle size) and major prey abundances (calanoid copepods and mysids) differed among the sites, no significant differences were apparent in the numbers of fish species and individuals among the sites in any month in either year. In addition, species and individual numbers of two dominant feeding groups (zooplankton feeders, and epiphytic and benthic crustacean feeders) did not show any significant responses to among-site differences in prey abundance. Cluster and ordination analysis confirmed the similarity in fish species composition among the three sites. These results indicated that the surf zone fish assemblage structures were similar among the microhabitats in spite of there being microhabitat-related differences in abiotic and biotic environments.  相似文献   

10.
Coastal defence structures constitute the most extensive hard substrates of the Northwestern Adriatic Sea and are known to sustain rich benthic and nektonic communities. To appreciate the pattern of colonization, we studied the fish assemblage of a recently deployed breakwater. We compared observations from two years, the different sides (landward and seaward) of the barrier, and the two fringes, characterized by timing of work completion. The results indicate that colonization, still in process, follows different patterns among species. Benthic and necto-benthic species presented a striking increase in abundance and richness in the second year of colonization, while more mobile species did not evince any variation between years. Differences in mobility among species suggest that the latter group may have reached the breakwater from nearby artificial substrates, whereas the former colonized the new structure as recruits. In addition, fish assemblages differed between the two sides, likely due to variation in the environmental characteristics, and according to depth, reflecting species preferences.  相似文献   

11.
Multivariate principal components analysis and cluster analysis were performed on data representing presence or absence of 498 species of juvenile and adult fish species in twenty-four coastal marine systems (bays, coastal lagoons, estuaries and coastal zones) distributed from southeast to southern Brazil. Five groups of coastal systems were identified based on fish assemblage similarity: estuaries and bays of the southeast area; an estuary of the southern area; coastal lagoons; rocky coastal zones; and the continental platform. Species assemblages for each zone were identified and used as surrogate habitat indicators to compare and contrast the groups. Stepwise multiple regression of environmental and physical variables as predictors of the number of species indicated that only ‘ area ’ was included in the model as the most important variable explaining the variation of the number of species in these data sets. The total number of fish species increased as surface area increased.  相似文献   

12.
The concept of essential fish habitats (EFHs) is widely accepted for conservational and management purposes. EFHs are often considered as high quality habitats for fisheries species and subsequently of high values for society. In this study, fish and Substrate-Associated Prey (SAP) were sampled during the productive summer season 1998 (fish) and 2003 and 2004 (SAP) in shallow coastal rocky- and soft-bottom habitats on the Swedish west coast. The aim was to study the spatial and monthly variation of SAP as well as abundance and biomass of fish, and to examine if food items found in the diet of the fish assemblage were derived from SAP. We also examined if the diet of Ctenolabrus rupestris, a resident and abundant fish species in the shallow coastal habitats, and the diet of four seasonally abundant and commercially important fish species (Gadus morhua, Pleuronectes platessa, Salmo trutta and Scomber scombrus) were derived from SAP. There were significantly higher mean species number and abundance of the SAP assemblage on rocky compared to soft bottoms and the highest values were found on the rocky bottoms in August and in the shallowest (0–3 m) depth strata. There were no significant differences in number of fish species caught in the two habitats, although mean number of fish and mean biomass were significantly higher on rocky bottoms. Both habitats showed the same seasonal variation and the highest values of number of fish species, abundance and biomass were observed in June. On rocky bottoms, gastropods and amphipods were the most frequent food items in the diet of the entire fish assemblage and these items were also the most abundant SAP in this habitat. The dominant food items of the soft-bottom fish assemblage were decapods and fish, which were not common SAP. However, except for S. scombrus, the diet of the selected fish species showed a strong association to the SAP availability. Gadus morhua displayed the strongest association to SAP on rocky bottoms and P. platessa and C. rupestris to SAP on soft bottoms. Further, for C. rupestris, multivariate statistical analysis showed a significant association to the SAP assemblage on both rocky and soft bottoms. These results provide vital new information for the management and conservation of Essential Fish Habitats on the Swedish west coast.  相似文献   

13.
Variability of fish assemblages across habitat structures can depend on spatial scales. A hierarchical sampling design was used to assess the spatial variability of temperate fish assemblages in different habitats and at multiple scales. Underwater visual censuses were carried out along the coasts of Elba Island (NW Mediterranean) on Posidonia oceanica beds, rocky algal reefs and sandy habitat at three spatial scales, namely tens of metres (individual replicates), hundreds of metres (sites) and tens of kilometres (locations). At the assemblage level, there was a clear relationship between fish and habitat type and the observed habitat‐related differences were largely dependent on species identity. Fish assemblages on P. oceanica beds and rocky reefs shared a high number of species, whereas overlap with sandy assemblages was negligible. Multivariate analyses revealed significant differences in fish assemblages among habitats, although there was also a significant habitat × site interaction. These differences relied mainly upon assemblage composition and species richness. Assemblages on rocky reefs and P. oceanica meadows usually harboured a higher number of species and individuals compared with sandy assemblages. Nevertheless, the patterns of habitat‐related differences in species richness and, especially, in the total number of fish, changed significantly from site to site. Eight species showed significant differences over habitats, but they were not consistent due to the interaction of habitat with site. Predictability of fish at both assemblage and population levels decreased with the scale of observation, and the spatial pattern of fish observed at the smallest scale was likely dependent on factors other than habitat type.  相似文献   

14.
Surgeonfish and parrotfish play an important role in structuring the benthic communities of coral reefs. However, despite their importance, little is known about their distribution patterns in the north sector of the Mesoamerican Reef System. This study evaluated the distribution of these fish in 34 sites in four habitats (lagoon, front, slopes and terrace) along a depth gradient (c 0.5–20 m). These herbivorous fish were assessed by visual censuses. Species dominance was evaluated for each habitat using SIMPER analysis. Habitat characteristics data were collected to determine the relationship between habitat conditions and spatial variations in herbivorous fish (using abundance and biomass as a proxy) via redundancy analysis. The herbivorous fish assemblage had a low density (fish per 100 m2) and biomass (g·100 m?2) in comparison with assemblages in similar studies. In contrast, species richness was high compared with other studies in the Caribbean. Spatial variation of the abundance, biomass and size of herbivorous fish was strongly related to coral and seagrass cover, as well as to depth and rugosity. These four variables were critical in controlling the distribution patterns of the herbivorous fish assemblages. No associations were found between fish and macroalgae or any other benthic group. The present study indicates that the species richness of surgeonfish and parrotfish was not regionally affected by the dominance of macroalgae in the habitats studied. Seagrass beds and the coral reef matrix need to be preserved for the herbivorous fish assemblages to remain healthy and capable of controlling excess macroalgae growth.  相似文献   

15.
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16.
Many of the world's fish populations are overexploited, including Ghana's fish resources. This study examines spatio-temporal trends in fish species composition in relation to biotic, abiotic and anthropogenic factors, towards achieving better-informed management of the beach-seine fisheries. Fishery-dependent data were collected between November 1999 and October 2001 from 94 beach-seine hauls fished at two stations along the Ghanaian coast. The catch consisted of fish, crustaceans, other invertebrates, and macroalgae. Generally, juveniles of species that are exploited by offshore fisheries were found in the catches. Species abundance and their occurrence generally peaked between November and January. Duration of solar radiation and tide level appeared to be important predictors of fish biomass. It is important to exploit the nearshore fish assemblage sustainably because of its nursery role. A co-managed (fishers and government) three-month ban on beach-seining (between May and July) is recommended as the most appropriate control measure towards the sustainability of Ghanaian fish stocks.  相似文献   

17.
The Wadden Sea is a shallow coastal region, with a large area of sedimentary tidal flats that extends from The Netherlands to Denmark and has been declared a site of international importance in the Dutch and German parts (Ramsar status and UNESCO World Heritage Site). Benthic macrofauna are central to the ecosystem functioning of this area, as they recycle nutrients, decompose organic matter and are an important food source for many secondary consumers, like fish and waterbirds. Due to the environmental gradients characteristic of estuarine systems, it is expected that changes in assemblage composition will be observed across the physical and environmental gradients of the Wadden Sea. First, we explored the spatial variation in assemblage composition of benthic macrofauna across the intertidal part of the Dutch Wadden Sea using 3 years of biomass data. Then, we identified the relative importance of six environmental variables for explaining and predicting changes in assemblage composition across the intertidal areas of the Wadden Sea using generalised dissimilarity modelling (GDM). In accordance with the environmental gradients across this system, the biomass contributed by a few common species differed from west to east and were distinct in the Dollard. In the west, bivalves Mya arenaria, Cerastoderma edule and Ensis directus contributed a relatively large and equal share of the total biomass, whereas C. edule contributed the sole largest share of the total biomass towards the east. The polychaete Alitta succinea became a large share of the total biomass in the upper Ems and in the Dollard estuary, but contributed little elsewhere. Similar to the observed differences in species composition, the spatial patterns in assemblage composition, as predicted by the GDM models, identified the Dollard as distinct and that the prevalence of assemblage types in the west differed to the east. Median grain size, followed by microphytobenthic biomass, and exposure time were the most important variables describing differences in assemblage composition. That the Wadden Sea forms a heterogeneous landscape where assemblage composition varies across multiple gradients has repercussions for management and monitoring.  相似文献   

18.
The spatial and temporal characteristics of trophic structure of fish communities in the southern Huanghai Sea were examined based on the data sampled from bottom trawl surveys conducted during the autumn of 2000 and the spring of 2001. Hierarchical agglomerative cluster method and bootstrap randomization were used to identify significant trophic groups for each fish assemblage in the southern Huanghai Sea. A total of six major trophic groups were identified within this system, which classified predators based upon location in the water column or prey size （ i. e. , benthic to pelagic predators or fish to small invertebrate prey predators）. The similarity level used to identify significant trophic groups in each assemblage ranged from 24% to 34%. Although planktivores were the dominant trophic group in each assemblage （60% - 79% ）, there were spatial and temporal variations in the trophic structure, which reflected the differences in the abundance and availability of dominant preys. Simplified food webs were constructed to evaluate the most important trophic relationships between the dominant prey taxa and the fishes in each assemblage within this system. Although there were some differences in the key prey species among different food webs, pelagic prey items （mainly euphausiids and copepods） represent the most important energetic link between primary producers and higher trophic level predators. The trophic level for most fishes was between 3 and d, and the weighted mean trophic level for each assemblage ranged from 3.3 to 3.4. Compared with previous study in the mid-1980s, there was an obvious downward trend in the trophic level for most fish species, which resulted mainly from the fluctuation in key prey species in the Huanghai Sea. The decrease in the importance of Japanese anchovy seems to be offset by other abundant prey species such as Euphausia pacifica and copepods （ mainly Calanus sinicus ） .  相似文献   

19.
Coral reef fish are an important source of food security and income for human coastal populations. They also underpin ecosystem processes vital for the future ability of coral reefs to generate ecological goods and services. Identifying socio-economic drivers behind the exploitation of fish that uphold these key ecosystem processes and the scales at which they operate is therefore critical for successful management. This study addresses this issue by examining the reef-associated fish value chain in Zanzibar, and how it links to functional groups of fish and maturity stage of fish within these groups. Semi-structured interviews with 188 respondents (fishers, traders and hotel staff) involved in the fisheries and trade with reef-associated fish in Zanzibar and participatory observations were used. The trade with reef fish in Zanzibar is a complex structure involving many different agents and this study shows that these different agents exhibit differential “preferences” regarding fish functional groups and/or maturity stages within these groups. Consequently, both high and low trophic species, as well as small and large fishes are fished and sold, which leaves no refuge for the fish assemblage to escape fishing. When other market agents than fishers have so much influence and there are few alternative income generating activities, it is not possible to put all burden on fishers. Management measures that extend down the value chain to include all market agents as well as their links to ecosystem processes are thus likely to be needed to reach the target of sustainable fisheries.  相似文献   

20.
The composition of the demersal nekton assemblage inhabiting the Mediterranean shelf-break of west-central Italy was examined to investigate the diel variation in species composition and abundance. Fish were obtained during four 24-h sampling periods that represented the four annual seasons. Thirty-two hauls (eight in each season) were conducted in a shelf-break area at 140–160 m depth. The density of fish, crustaceans, and decapods exhibited differences on short- and long-term time scales: multidimensional scaling suggested that season and light intensity may be important factors influencing fish assemblage composition.At the diel scale two main assemblages separating nocturnal and diurnal hauls were identified. The typical assemblage structure of shelf-break bottoms was represented during daylight by species such as Merluccius merluccius, Capros aper, Serranus hepatus, Macroramphosus scolopax and Trisopterus minutus capelanus. At night, the assemblage showed a drastic change due to the increase of small nektobenthic species, Argentina sphyraena and Glossanodon leioglossus, together with cave-dwelling fish (Gnathophis mystax, Chlopsis bicolor, Nettastoma melanurum) and nektobenthic crustaceans (i.e., Parapenaeus longirostris and Solenocera membranacea).The above-mentioned differences in species composition may be related to changes in vertical distribution for trophic needs (M. merluccius) and/or in the shoal structure (A. sphyraena and G. leioglossus). The lack of data on the pattern of trawl catchability of Mediterranean species during the day may lead to an inaccurate estimation of species abundance when only diurnal trawl survey data are used.  相似文献   

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