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1.
Seven planktic foraminiferal zones are distinguished in the Maastrichtian-Paleocene succession at the north Farafra Oasis. These are the Rugoglobigerina hexacamerata (CF8b), Gansserina gansseri, and Contusotruncana contusa zones in the Maastrichtian topped by a well-known unconformity across the Cretaceous/Paleogene (K/Pg) boundary. The Danian is subdivided into two biozones: Globanomalina compressa/Praemurica inconstans-Praemurica uncinata Subzone (P1c) and Praemurica uncinata–Morozovella angulata (P2) Zone. The Late Paleocene is divided into two zones: Morozovella angulata-Globanomalina pseudomenardii (P3) and Globanomalina pseudomenardii (P4). A minor hiatus between the Danian/Selandian and Selandian/Thanetian boundaries are also recorded. These time gaps across the stage boundaries may be related to the tectonic events that affected the sedimentation regime throughout the Upper Cretaceous–Lower Paleogene interval in the Farafra Oasis.  相似文献   

2.
Planktonic foraminiferal analysis of the Erto section in the Vajont valley (Southern Alps, northern Italy) reveals a relatively complete succession across the Cretaceous–Tertiary (K–T) boundary. The turnover of planktonic foraminiferal fauna was studied for a stratigraphic interval spanning theAbathomphalus mayaroensisZonep.p., Pseudotextularia deformisZone,Guembelitria cretaceaZone,Parvularugoglobigerina eugubinaZone,Eglobigerina eobullioidesSubzone, andParasubbotina pseudobulloidesZonep. p.The extinction of most large, ornate, late Maastrichtian species occurs below a black ‘boundary clay’ (2–4 cm thick); however, part of the Late Cretaceous species, mainly heterohelicidids and hedbergellids, were found over an interval of more than 100 cm above the boundary. Although a relatively high number of species occur for the last time in the main extinction phase, the abundance of these outgoing species is less than 20% of the total population; unkeeled or weakly keeled, simple-shaped forms (heterohelicids, globotruncanellids, hedbergellids) constitute the bulk of the planktonic foraminiferal population both in uppermost Maastrichtian and lowermost Danian beds. The first Tertiary species (‘Globigerina’minutulaand ‘Globigerina’fringa) appear just above the ‘boundary clay’;Parvularuglobigerina eugubinaoccurs a few centimeters above. A marked increase in abundance and diversity in the Tertiary planktonic foraminiferal population occurs at the base of theEoglobigerina eobulloidesSubzone.  相似文献   

3.
Geochemical and isotopic analyses of the Cretaceous–Paleogene (K/Pg) boundary deposits were conducted at the Caravaca section (External Subbetic, southeast of Spain) in order to evaluate the recovery of the macrobenthic tracemaker community and the bioturbational disturbance. Samples from the infilling material of several lower Danian dark-colored trace fossils (Chondrites, Planolites, Thalassinoides and Zoophycos) located in the uppermost 8-cm of the light upper Maastrichtian strata, as well as samples from the host sedimentary rock of these trace fossils, were analyzed and compared with data from the lower Danian deposits. The values of element ratios indicative of extraterrestrial contamination (Cr/Al, Co/Al and Ni/Al) are higher in the infilling trace fossil material than in the upper Maastrichtian and lower Danian deposits, which suggests a contribution of the ejecta layer. Regarding the isotope composition, the δ13C values are lower in the infilling material than in the Maastrichtian host sedimentary rocks surrounding the traces, while the δ18O are higher in the infilling material. The geochemical and isotopic compositions of the infilling material evidence the unconsolidated character of the sediment, including the red boundary layer. Softground conditions confirm a relatively rapid recovery by the macrobenthic tracemaker community, starting a few millimeters above the K/Pg boundary layer. The mixture of the infilling material of the trace fossils moreover reveals a significant macrobenthic tracemaker activity affecting K–Pg boundary transition sediments that may have significantly altered original signatures.  相似文献   

4.
This study of the upper Maastrichtian to Danian sedimentary succession from the northern part of the Romanian Eastern Carpathians (Varniţa section) aims to establish an integrated biostratigraphy based on calcareous nannofossils, organic-walled dinoflagellate cysts (dinocysts) and foraminiferal assemblages, and to reconstruct the depositional environments of the interval. The stratigraphic record across the studied section is incomplete, considering that an approximately 16 m thick strata interval from the top of the Maastrichtian to lowermost Danian cannot be analyzed due to a landslide covering the outcrop. The upper Maastrichtian is marked by a succession of biostratigraphic events, such as the First Appearance Datum (FAD) of the nannoplankton taxon Nephrolithus frequens and FAD of the dinocyst species Deflandrea galeata and Disphaerogena carposphaeropsis, and the Last Appearance Datum (LAD) of Isabelidinium cooksoniae in the lower part of the section. These bioevents are followed by the LAD of the Dinogymnium spp. and Palynodinium grallator dinocyst markers in the top of the Maastrichtian deposits analyzed. In terms of foraminiferal biostratigraphy, the upper Maastrichtian Abathomphalus mayaroensis Zone is documented in the lower part of the studied section. Some bioevents, such as the bloom of the calcareous dinoflagellate genus Thoracosphaera and the FAD of the organic-walled dinocysts Damassadinium californicum, Senoniasphaera inornata, Xenicodinium lubricum and X. reticulatum suggest an early Danian age for the middle part of the section. From the Danian deposits in the Varniţa section, we describe a new organic-walled dinocyst species, Pentadinium darmirae sp. nov., which is until now the only species of the Pentadinium genus discovered in the Paleocene. The occurrence of the global Danian dinocyst marker Senoniasphaera inornata in the top of the section, suggests an age not younger than middle Danian (62.6 Ma) for the analyzed deposits.The palynofacies constituents, as well as the agglutinated foraminiferal morphogroups, used to reconstruct the depositional environments, show that the late Maastrichtian sediments were deposited in an outer shelf to distal (bathyal) environment, followed by a marine transgression during the Danian.  相似文献   

5.
The Cretaceous outcrop belt of the Mississippi Embayment in the Gulf Coastal Plain (GCP) spans the Cretaceous/Paleogene (K/Pg) boundary. A detailed reconstruction of this time interval is critical for understanding the nature of biotic and environmental changes preceding the end-Cretaceous Mass Extinction event and for deciphering the likely extinction mechanism (i.e., bolide impact versus volcanism). Eight sections encompassing the K/Pg succession across the Mississippi Embayment were analyzed using biostratigraphic sampling of ammonites, dinoflagellates, and nannofossils. An upper Maastrichtian ammonite zonation is proposed as follows, from oldest to youngest: Discoscaphites conradi Zone, D. minardi Zone, and D. iris Zone. Our study documents that the ammonite zonation established in the Atlantic Coastal Plain (ACP) extends to the GCP. This zonation is integrated with nannofossil and dinoflagellate biostratigraphy to provide a framework to more accurately determine the age relationships in this region. We demonstrate that ammonites and dinoflagellates are more reliable stratigraphic indicators in this area than nannofossils because age-diagnostic nannofossils are not consistently present within the upper Maastrichtian in the GCP. This biostratigraphic framework has the potential to become a useful tool for correlation of strata both within the GCP and between the GCP, Western Interior, and ACP. The presence of the uppermost Maastrichtian ammonite D. iris, calcareous nannofossil Micula prinsii, and dinoflagellates Palynodinium grallator and Disphaerogena carposphaeropsis suggests that the K/Pg succession in the GCP is nearly complete. Consequently, the GCP is an excellent setting for investigating fine scale temporal changes across the K/Pg boundary and ultimately elucidating the mechanisms causing extinction.  相似文献   

6.
The upper Campanian to upper Maastrichtian sedimentary sequence of the Kiseiba Formation in south Western Desert is sampled and described in two surface sections (Sinn El Kaddab and Wadi Abu Siyal). Forty-four agglutinated foraminiferal species are identified from 42 samples in the studied succession. The benthic foraminiferal assemblages are dominated by agglutinated foraminifera which comprise more than 90% of the assemblage. The agglutinated foraminifera are subdivided into five morphogroups (A, B, C, D, E) according to shell architecture, integrated with the supposed microhabitat and feeding strategy. The foraminiferal assemblage is assigned to mixohaline shallow water environments. These assemblages with Ammoastuta megacribrostomoides and Ammotium bartheli suggest lagoonal environments with considerably reduced salinity in warm climates and high runoff for the late Campanian-Maastrichtian interval.  相似文献   

7.
The Maastrichtian regressive sequence of the Hekimhan-Malatya area in Eastern Turkey consists of dolomitic limestones, limestones and calcareous mudstones which are dominated by rudists and Loftusia (foraminifera) assemblages. Several Loftusia species indicating middle to late Maastrichtian age such as Loftusia anatolica Meriç, L. baykali Meriç, L. coxi Henson, L. harrisoni Cox, L. minor Cox and L. morgani Douvillé have been recorded. Other benthic foraminifers present include Orbitoides medius d’ Archiac, Omphalocyclus macroporus (Lamarck), Laffitteina conica Drooger, Laffitteina mengaudi (Astre) and Laffitteina oeztuerki Inan. The rudists are abundant in the dolomitic limestones in the study area. The genus Miseia Patrulius is dominant and represented by Miseia bilacunosa Özer and Miseia hekimhanensis Karacabey-Öztemür. The Loftusia species and the Rudists assemblage indicates middle to late Maastrichtian age for the formation. The paleobiogeographic distribution of the assemblage has been discussed taking this find into account.  相似文献   

8.
The Late Cretaceous (Maastrichtian) to early Paleocene (Thanetian) shallow water (<100 m) agglutinated foraminifera from a section at Dakhla Oasis (Western Desert, Egypt) were analyzed for their assemblage, species and genera distribution, diversity, depositional environment, community structure and palaeobathymetry with respect to regional tectonics, climate and global eustasy. Data suggest an equitable benthic environment with low species dominance deposited in a brackish littoral and/or marsh setting. Sea level curves using characteristic benthic foraminiferal species, genera and assemblages corroborate quantitatively generated estimate and statistical analysis. Data suggests that in the absence of or of an impoverished benthic foraminiferal fauna, a high resolution agglutinated foraminiferal dataset can be as good a predictor of the benthic community structure and environment, as its calcareous counterpart, at least for shallow settings (<100 m). Present data also provides a good window in better understanding the distribution and interrelationship between the three dominant genera, Haplophragmoides, Trochammina and Ammobaculites. Faunal changes at boundaries (Cretaceous/Paleogene, Danian/Selandian and Selandian/Thanetian) are also evaluated.  相似文献   

9.
Cretaceous climate, volcanism, impacts, and biotic effects   总被引：5，自引：0，他引：5  
Cretaceous volcanic activities (LIPs and CFBPs) appear to have had relatively minor biotic effects, at least at the generic level. Major biotic stress during the Cretaceous was associated with OAEs and related to nutrient availability largely from weathering, greenhouse warming, drowning of platform areas, and volcanism. The biotic effects of OAEs were often dramatic at the species level, causing the extinction of larger specialized and heavily calcified planktonic foraminifera (rotaliporid extinction) and nannoconids (nannoconid crises), the temporary disappearances of other larger species, and the rapid increase in r-selected small and thin-walled species, such as the low oxygen tolerant heterohelicids and radially elongated taxa among planktic foraminifera and thin walled nannofossils. Biotic diversity increased during cool climates, particularly during the late Campanian and Maastrichtian, reaching maximum diversity during the middle Maastrichtian. High biotic stress conditions began during greenhouse warming and Deccan volcanism about 400 ky before the K-T boundary; it reduced abundances of large specialized tropical planktic foraminiferal species and endangered their survival. By K-T time, renewed Deccan volcanism combined with a large impact probably triggered the demise of this already extinction prone species group.Evidence from NE Mexico, Texas, and the Chicxulub crater itself indicates that this 170 km-diameter crater predates the K-T boundary by 300,000 years and caused no species extinctions. The Chicxulub impact, therefore, can no longer be considered a direct cause for the K-T mass extinction. However, the K-T mass extinction is closely associated with a global Ir anomaly, which is considered too large, too widespread, and too concentrated in a thin layer to have originated from volcanic activity, leaving another large impact as the most likely source. This suggests that a second still unknown larger impact may have triggered the K-T mass extinction.  相似文献   

10.
The uppermost Cretaceous (upper Campanian–Maastrichtian) marine deposits of the central south Pyrenees host a rich larger benthic foraminiferal fauna and several rudist-rich levels. These marine deposits are directly overlain by the continental facies of the Arén and Tremp Formations, which are famous for their fossil dinosaur remains. Larger benthic foraminiferal distribution documents an important faunal turnover in all the carbonate platform environments within the photic zone, from open marine to littoral areas. Biostratigraphy indicates that this turnover occurred close to the Campanian-Maastrichtian boundary. This is also confirmed by strontium isotope stratigraphy which indicates an earliest Maastrichtian age for the appearance of the larger benthic foraminiferal assemblage constituted by Lepidorbitoides socialis, Clypeorbis mammillata, Wannierina cataluniensis, Orbitoides gruenbachensis, Siderolites aff. calcitrapoides, Fascispira colomi, Omphalocyclus macroporus and Laffiteina mengaudi. In particular, a numerical age of 71 Ma is obtained for the Hippurites radiosus level, just a few meters below the first continental deposits of the Arén sensu stricto Formation. The youngest marine sediments of the central south Pyrenees are early Maastrichtian in age. This is also an important constraint for the age of the end-Cretaceous dinosaur fossil localities of the Tremp basin.  相似文献   

11.
Located to the north of the Stevns Klint Peninsula (Denmark), Kulstirenden shows the transition from Cretaceous chalks to Danian carbonates across the Cretaceous - Paleogene (K/Pg) boundary. The K/Pg boundary at Kulstirenden is represented by the Fiskeler Member (Fish Clay), a streaked marl interval with a smectite rich, red oxidised layer at its base marking the boundary. The Fiskeler Member is important as it includes the iridium concentration linked to the bolide impact which may have caused the end-Cretaceous mass extinction and is at its maximum thickness at Kulstirenden (c.45 cm). Calcareous dinoflagellate cysts (Calciodinelloideae) have been investigated from within the fine fraction (45-125 μm) at Kulstirenden in order to understand the ecological implications of these enigmatic microfossils. Several species were found within the Fiskeler Member, including Orthopithonella collarisWendler et al. (2001, Rev. Palaeobot. Palynol. 115, 69-77). This species is of particular interest as it demonstrates reduced paratabulation and may indicate a transgressive period immediately after the K/Pg boundary event. O. collaris has been described as a morphotype formed under post-K/Pg environmental conditions and is, therefore, described as a “disaster” taxon that marks the K/Pg boundary at Stevns Klint. The distribution of this taxon is more extensive than previously documented, disappearing as carbonate sedimentation returned in the Cerithium Limestone Member of the Danian. Several calcareous microfossil specimens that were found cannot be associated with the dinoflagellates and display Bolboforma-like features. While the origins of Bolboforma are enigmatic, the current finding expands their stratigraphic range and suggests an appearance of this group of organisms found at Stevns Klint may give an insight into the inception of this group into the earliest Danian.  相似文献   

12.
This study is based on calcerous nannofossil assemblages changes and fluctutions of stable carbon and oxygen isotopes was collected clayey limestones, limestones, and marls in the Maastrichtian to Selandian from Akveren Formation (Western Black Sea). As the relative abundances of species of Micula spp, Watznauera barnesiae, and Arkhangelskiella cymbiformis, which tolerated changes of temperature and nutrition, carbon and oxygen isotopes compositions, and low species richness imply strong diagenesis effect at the Maastrichtian, there is no important diagenesis effect at Paleocene. Just after the Cretaceous/Tertiary (K-Pg) boundary, Thoracosphaera spp. and Braarudosphaera bigelowi were dominant species; Danian is characterized by Thoracosphaera, Ericsonia ovalis, Cruciplacolithus spp., Coccolithus pelagicus, and Ericsonia subpertusa. Generally, the nutrition productivity is good–moderate in Lower Maastrichtian, and decreasing carbon isotope values during the Uppermost Maastrichtian shows the presence of oligotrophic environmental conditions suitable with global nutrition crises before the K-Pg boundary and diagenesis in study area. Throughout the Danian, mesotrophic–oligotrophic environmental conditions dominate; however, the decrease in nutrition before Selandian represents oligotrophic environmental conditions. The increasing nutrition at Selandian is related to the change in the environmental conditions.  相似文献   

13.
The Upper Cretaceous succession outcropping in the Anamas–Akseki Autochton, consists of approximately 500 m thick purely platform carbonate sediments. It begins with Cenomanian limestones intercalated with limestone breccias (Unit-1) containing mainly Pseudorhapydionina dubia, Pseudonummoloculina heimi, Spiroloculina cretacea (Assemblage I) and unconformably overlies the Lower Cretaceous (Barremian–Aptian) limestones with Vercorsella laurentii, Praechrysalidina infracretacea and Salpingoporella hasi. The Cenomanian limestones include foraminiferal packstone–wackestone, peloidal packstone–wackestone and mudstone microfacies deposited in restricted platform conditions. The Cenomanian succession is truncated by an unconformity characterised by locale bauxite deposits. Immediately above the unconformable surface, dolomitic limestones and rudistid limestones (Unit-2) are assigned to the upper Campanian based on the benthic foraminiferal assemblage (Assemblage II) comprising mainly Murciella gr. cuvillieri, Pseudocyclammina sphaeroidea, Accordiella conica, Scandonea samnitica and Fleuryana adriatica (smaller-sized populations). The upper Campanian limestones composed of dominantly foraminiferal-microbial packstone–wackestone microfacies deposited in shallow water environment with low energy, restricted circulation. The following limestones of the Unit-2 is characterised by sporadic intercalation of “open shelf” Orbitoides, Omphalocyclus, Siderolites assemblage (Assemblage III), assigned to the Maastrichtian, in addition to pre-existing “restricted platform” species. In the upper part of this biozone, the Rhapydionina liburnica/Fleuryana adriatica concurrent range subzone (Assemblage IIIb) is distinguished by the presence of Valvulina aff. triangularis, Loftusia minor as well as the nominal species. The Maastrichtian limestones with sporadically open marine influence consist of bioclastic (rudist-bearing) packstone–floatstone, foraminiferal packstone–wackestone with rudist fragments and peloidal/intraclastic packstone–wackestone microfacies deposited in shallow subtidal–subtidal (lagoonal) environments. The Upper Cretaceous succession passes upwardly into 70 m thick limestones and clayey limestones (Unit-3) which do not contain rudists and pre-existing foraminiferal assemblage with one exception Valvulina aff. triangularis. Variable amounts of ostracoda, discorbids, miliolids, dasycladacean algae and Stomatorbina sp. (Assemblage IV) occur into mud-rich microfacies suggesting restricted conditions with low water energy. A probable Danian age is proposed for the Unit-3 based on the occurrence of Valvulina aff. triangularis and Stomatorbina sp. which were previously recorded from Danian of peri-Tethyan platforms.  相似文献   

14.
Though the Maastrichtian Phosphates of Morocco have yielded very rich marine vertebrate assemblages, plesiosaurs remain very scarce in these strata. The only hitherto recognized taxon was Plesiosaurus mauritanicus Arambourg, 1952, regarded here as a nomem dubium. Here we describe a new genus and species of elasmosaurid plesiosaur, Zarafasaura oceanis, which represents the first valid elasmosaurid plesiosaur described from the latest Cretaceous of Africa, and the second one from this continent. A phylogenetic analysis of plesiosauroids indicates that Zarafasaura oceanis has close affinities with elasmosaurids from the Late Cretaceous of North America and Japan. Among its distinctive suite of characters, the general shape and organisation of its squamosal and palate are unique among elasmosaurids. This new taxon completes our understanding of Late Cretaceous plesiosaur palaeobiodiversity and palaeobiogeography, and shows that Maastrichtian plesiosaurs were characterized by a quite high degree of endemism. They were also highly diversified and distributed worldwide, which supports the hypothesis of a catastrophic extinction of plesiosaurs at the K/T boundary.  相似文献   

15.
The Bir Dakhl section which is located in the southern Galala plateau, north eastern desert was sampled for microfacies analysis of the upper Cretaceous–lower Paleogene succession. Microfacies analysis led to the recognition of eight mixed clastic-carbonate facies types (black shale lithofacies MF-1, pure shale lithofacies MF-2, sandy shale lithofacies MF-3, marly shale lithofacies MF-4, mudstone microfacies MF-5, foraminiferal wackestone microfacies MF-6, bioclastic wackestone microfacies MF-7, and bioclastic packstone microfacies MF-8) of the studied interval. These microfacies can be grouped into three depositional environments: inner, middle and outer ramp. The interpreted depositional environments have been suggested to range from neritic to middle bathyal (~ 700 m). Based on cyclostratigraphy, five deepening upward cycles and three shallowing upward cycles have been differentiated in Bir Dakhl section.  相似文献   

16.
A preliminary integrated study of a Cretaceous-Tertiary (K/T) boundary section in Pernambuco, northeastern Brazil, provides evidence for an extraterrestrial bolide impact in the earliest Danian. A non-graded, nodular carbonate mudstone/wackestone bed is interpreted as a slump or a mud Flow deposit, induced by a tsunami event or by a gravity mass movement during a sea-level lowstand, the former possibly generated by the K/T boundary Yucatán impact. This bed overlies marlstones deposited in an upper bathyal environment and marks the top of the Cretaceous. Nearly all known latest Maastrichtian planktonic foraminifera are recovered from the Cretaceous strata. Iridium shows a marked peak in a thin hemipelagic claystone layer, about 75 cm above the K/T boundary, deposited in a middle to deep neritic environment. The claystone overlies alternating beds of finegrained limestones and marlstones and a 50 cm-thick graded bioclastic packstone, which rests upon a marly limestone breccia of the lowermost Danian. Abundant reworked Cretaceous and rare lowermost Danian microfossils (e.g. E. edita, E. eobulloides, E. fringa, G. irregularis, P. eugubina, P. cf. pseudobulloides, W. claytonensis and W. hornerstownensis ) have been recovered from these strata. These lowermost Danian beds record the sudden appearance of abundant shock-metamorphosed quartz grains, with several sets of intersecting deformation lamellae and microtektite-like microspherules. This suggests that these deposits were formed by an additional tsunami caused by a second impact event in the earliest Danian, near or at the boundary between the Palaeocene P α and P 1a foraminiferal zones. Impact-derived material has not been found in the uppermost Cretaceous beds.  相似文献   

17.
The Kopet-Dagh sedimentary basin is located in northeast of Iran and southeast of Turkmenistan in the Middle East. The Khangiran formation represents the last marine deposition in the Kopet-Dagh sedimentary basin. The early Eocene planktonic and benthic foraminifera from the Khangiran formation which belongs to the lower 376 m thickness of this formation are identified and biostratigraphically evaluated. Due to rarity of Morozovella species, planktonic foraminiferal zonation was difficult to determine in this formation. The determination of upper part of the late Paleocene for the lower 124 m of this formation is according to the lowest occurrence of Acarinina sibaiyaensis species. From E5 to near middle of E7 biozone, increasing trend of Acarinina frequency and a peak in Morozovella species and decreasing trend of frequency of Subbotina and Pseudohastigerina species indicate the warm and oligotrophic condition of the seawater during sedimentation of the studied interval. Toward the Ypresian-Lutetian boundary, the increasing trend of Pseudohastigerina and Subbotina species and decreasing in frequency of Acarinina species suggest the low-oxygen level, eutrophic and intermediate condition of the seawater. In this formation, the high abundance of the epifaunal taxa such as Anomalinoides spp., Cibicidoides spp., Gyroidinoides spp., and Lenticulina spp. from the base (late Paleocene sediments) up to E6 biozone reflects oligotrophic and oxic shallow water conditions. The occurrence of several peaks in abundance of Bulimina and Uvigerina species at the Ypresian/Lutetian boundary suggests eutrophic condition. These paleoecological conditions could be correlated with fluctuations in the numbers of the planktonic foraminifera.  相似文献   

18.
Benthic foraminiferal fauna are analyzed quantitatively and qualitatively at three stratigraphic sections in Eastern Desert of Egypt (Serai, Duwi, and Um El Huetat). These sections embrace the Paleocene–Eocene Thermal Maximum (PETM) interval which is represented by the occurrence of five distinctive beds. These beds constitute the Dababiya Quarry Member at the lower part of Esna Formation. The occurrence of them indicates an expanded and relatively continuous record across the P/E boundary. The organic-rich clay layer (bed no. 1 of the Dababiya Quarry Member) marks the start of the PETM event. This bed is characterized by the extinction of all benthic foraminiferal fauna except for the occurrence of rare agglutinated foraminiferal species. The presence of these species indicates an oceanic anoxic event at the sea floor. High concentration of phosphatic contents including fish remains occurred in the middle part of the PETM (bed nos. 2 and 3 of the Dababiya Quarry Member) with the continuous absence of benthic foraminiferal fauna except for few specimens at the top of bed 3. Bed nos. 4 and 5 of the Dababiya Quarry Member represent the upper part of the PETM and the initial stage of sea floor recovery. Low diversity and abundance of benthic foraminiferal taxa occurred within these beds, represented by Valvulineria scrobiculata, Lenticulina midwayensis, Loxostomoides appliane, and Siphogenerinoides eleganta. This phenomenon continues upward during the post-PETM event. The Paleocene velasco-type benthic foraminifera Angulogavelinella avnimelechi and Coryphostoma midwayensis species are extinct within the advent of the PETM event. The benthic foraminiferal assemblages at the studied sections are dominated by midway-type fauna with little representative of velasco-type fauna. The velasco-type species are represented with high abundance at Serai section and with low densities at Um El Huetat section, while at Duwi section, they rarely occurred. This suggests outer neritic-upper bathyal (150–400 m) setting at Serai section and mostly middle-outer neritic (50–150 m) setting at Um El Huetat and Duwi sections.  相似文献   

19.
Cold-water coral ecosystems represent unique and exceptionally diverse environments in the deep-sea. They are well developed along the Irish margin, varying broadly in shape and size. The Moira Mounds, numerous small-sized mounds, are nestled in the Belgica Mound Province (Porcupine Seabight, North-East Atlantic). The investigation of living (Rose Bengal stained) and dead benthic foraminiferal assemblages from these mounds allowed to describe their distribution patterns and to evaluate their response to environmental variability. Quantitative data was statistically treated to define groups of species/genera associated to specific habitats. The Moira Mounds differ from their larger neighbours by the reduced spatial variability of benthic foraminiferal assemblages, living assemblages only distinguishing coral-rich and coral-barren areas. The ecological needs of corals are highlighted by the abundance of Alabaminella weddellensis and Nonionella iridea, phytodetritus-feeding species in coral supporting sediments. Living foraminifera in sediments from the Moira Mounds concentrate in the upper first centimetre. Infaunal species may be affected by bioturbation and/or reworking by the strong currents in the area. Dead foraminiferal assemblages from the Moira Mounds resemble those described for the sandwave facies in adjacent giant mounds, suggesting similar processes in facies deposition.  相似文献   

20.
Compared to earlier representatives of the family, pachycephalosaurids are less well known from upper Maastrichtian deposits around the world. Here, we report on a nearly complete left postorbital attributable to the pachycephalosaurid Sphaerotholus cf. Sphaerotholus buchholtzae from the upper Maastrichtian Frenchman Formation of Saskatchewan. This marks a probable northern range extension for the species into Canada, and the first occurrence of a pachycephalosaurid from this formation. We further demonstrate the taxonomic distinction between Stegoceras edmontonense and S. buchholtzae, which has been debated, based on postorbital sutural proportions. The northerly occurrence of Sphaerotholus cf. S. buchholtzae is consistent with the hypothesis of low beta diversity during the late Maastrichtian of North America, and its high stratigraphic incidence documents the persistence of non-pachycephalosaurin pachycephalosaurines in a critical interval for understanding the end-Cretaceous mass extinction event.  相似文献   

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