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1.
One of the present concerns of fish biologists involves defining and identifying nursery habitats in the context of conservation and resource management strategies. Fish nursery studies usually report upon nursery occupation during the latter juvenile stages, despite the fact that recruitment to nurseries can start early in life, during the larval phase. Here we investigated the use of a temperate estuarine nursery area, the Lima estuary (NW Portugal), by initial development stages of flatfish species before and after metamorphosis, integrating the larval and juvenile phases. The Lima estuarine flatfish community comprised twelve taxa, seven of which were present as pelagic larvae, six as juveniles and three as adults. There was a general trend of increasing spring–summer abundance of both larvae and juveniles, followed by a sharp winter decrease, mainly of larval flatfishes. The Lima estuary was used by Solea senegalensis, Platichthys flesus and Solea solea as a nursery area, with direct settlement for the two first species. In contrast, indirect settlement was suggested for S. solea, with metamorphosis occurring outside the estuarine area. Estuarine recruitment of S. senegalensis varied between years, with young larvae occurring in the estuary throughout a prolonged period that lasted 6–9 months, corroborating the protracted spawning season. P. flesus, the second most abundant species, exhibited a typical spring estuarine recruitment, without inter-annual variations. Developed larvae arrived in the estuary during spring, whereas the 0-group juveniles emerged in the following summer period. The present study contributes new insight to our understanding of the economically important S. senegalensis, and highlights the importance of integrating the planktonic larval phase into traditional flatfish nursery studies.  相似文献   

2.
The recruitment variability of the marine fish species Dicentrarchus labrax, Platichthys flesus and Solea solea was evaluated in the Mondego estuary (Portugal) from 2003 to 2007. The relationships between sea surface temperature, NAO index, coastal wind speed and direction, precipitation and river runoff prior to the estuarine colonization and the abundance of 0-group fish were evaluated using gamma-based Generalized Linear Models. Dicentrarchus labrax and P. flesus 0-group decreased in abundance towards the end of the study period, while S. solea, despite low abundance in 2004, increased in abundance in 2007. For D. labrax, river runoff, precipitation and east–west wind were significant; for P. flesus, precipitation, river runoff and both north–south and east–west wind components were significant parameters, while for S. solea only river runoff was important. Results were compared with recent projections for climate change scenarios, to evaluate their effects on future recruitment levels.  相似文献   

3.
We tracked the duration and intensity of the euphausiid spawning season through biweekly sampling along a transect off Newport, OR (latitude 44°40′N) over a six year period from 1996 to 2001. Our sampling consisted of vertical plankton tows, CTD casts, and collection of water for determination of chlorophyll a. Here, we report on data collected from two stations, 5 and 15 nautical miles (9.3 and 27.8 km) offshore. The density of euphausiid eggs in our samples was highly variable spatially and temporally; we saw the most striking differences in egg densities and length of the spawning season, when we compared spawning before and after 1999. This year corresponded to the time when the Pacific Decadal Oscillation switched from warm phase (pre-1999) to cool phase (1999–present). The years 1996 and 1997 were characterized by one large, late summer peak in egg density at our inshore station. 1998, an El Niño year, followed this pattern for our offshore station, but eggs were nearly absent at our inshore station. Starting in 1999, we saw multiple peaks in egg density and found that the spawning season extended from spring through early fall. For example, in spring (March–May) at the inshore station, the abundance of eggs increased from an average of 0.4 m−3 (1996–1998) to 51.3 m−3 (1999–2001), and for summer (July–September), 27.8 m−3 to 132.6 m−3 for the same time period. At the offshore station, egg abundances doubled over the same two time periods: 7 m−3 versus 11 m−3 (spring) and 55 m−3 versus 186 m−3 (summer). Peaks in egg densities were often associated with phytoplankton blooms, but not in a predictable way. Peaks in egg densities often followed cold-water upwelling events, especially at the inshore station. It is not yet clear whether this connection is due to changes in advection or changes in upwelling-induced productivity.  相似文献   

4.
Micronuclei (MN), nuclear buds (NB) and fragmented-apoptotic cells (FA) were analyzed in mature peripheral blood and immature cephalic kidney erythrocytes of flounder (Platichthys flesus), dab (Limanda limanda) and cod (Gadus morhua) from 12 offshore sites in the Baltic Sea (479 specimens) and 11 sites in the North Sea (291 specimens), which were collected during three research vessel cruises in December 2002, 2003 and in September 2004. The highest levels of environmental genotoxicity (frequencies of MN up to 0.5‰, NB – up to 0.75‰) and cytotoxicity (FA – up to 0.53‰) were observed in flatfishes from areas close to oil and gas platforms in the North Sea and in zones related to the extensive shipping and potentially influenced by contamination from large European Rivers (Elbe, Vistula, Oder). In dab from the offshore zones of the North Sea, the levels of nuclear abnormalities were higher as compared to those in dab from the Baltic Sea. Responses in immature kidney erythrocytes were higher than in mature erythrocytes from peripheral blood. MN frequency lower than 0.05‰ (the Baltic Sea) and lower than 0.1‰ (the North Sea) could be suspected as a reference level in the peripheral blood erythrocytes of flatfish.  相似文献   

5.
The influence of distinct demographic factors on determining population size in a littoral nesting fish (Symphodus roissali) was studied. Differences in the overall abundances were studied at three sites in the NW Mediterranean Sea for three years, to examine whether: 1) the adult population determined the number of successful nests; 2) the number of successful nests determined recruitment levels; 3) recruitment levels were subsequently related to the number of young of the year (YOY) joining the population after the recruitment period; and 4) the YOY value determined the size of the year class 1 cohort in the following year's adult reproductive population. The results show that there was a significant relationship between the number of spawners and the number of successful nests. However, the number of successful nests does not always correlate well with recruitment level. Furthermore, recruitment levels are not good indicators of the Young of the Year (measured three months after settlement) in number joining the populations, though YOY in number was related to the strength of the year class joining the adult reproductive population the following year. Therefore, the magnitude of the recruitment and probably the size of the adult population did not determine the year-class strength in S. roissali.  相似文献   

6.
In early summer plaice (Pleuronectes platessa L.) in the Clyde Sea Area eat Tellina siphons and spionid palps. As they grow, they take progressively larger siphons, fragments of spionids and some crustaceans. In late summer they eat entire spionids and other sedentary polychaetes. There is no significant predation on fish. The common dab [Limanda limanda (L.)] took a wider range of prey species than the plaice, but also depended mainly on polychaetes, crustaceans and bivalves. Depth distribution may limit both intra- and interspecific competition between these fish and also influence the choice of prey. The results from the main nursery areas of Irvine and Ayr Bays were similar to those from the other bays throughout the Clyde Sea Area and support earlier work at Firemore Bay in north-west Scotland and in the Dutch Wadden Sea.The growth rate of the fish increases during the early summer and may be related to increasing food availability, feeding experience of the fish and longer day length. Growth rate between areas appeared to be correlated with benthic productivity. 0-group plaice in Irvine, Ayr and Firemore Bays grew to different mean lengths by the end of the summer growth season, being 82 mm, 73 mm and 61 mm, respectively, for the period 1972–1974. This agrees with the known differences in standing stocks of intertidal macrobenthos in these bays (17·0, 11·1 and 1·3 g m?2 dry weight, respectively). The significance of this in relation to survival is discussed. Offshore migration of the larger fish in the autumn may result in apparent slowing of growth rates prematurely in inshore areas. The year-class strength depends more on the food availability in the nursery areas than on the number of plaice that settle on the bottom.  相似文献   

7.
Cross-shelf distribution and abundance of copepod nauplii and copepodids were measured during three summer upwelling seasons (2000–2002) in a coastal upwelling zone off northern California. These 3 years varied considerably in the intensity of winds, abundance of chlorophyll, and water temperature. The cruises in 2000 were characterized by relaxation conditions, with generally high levels of chlorophyll and high water temperature. The cruises in 2001 and 2002 were dominated by strong and persistent upwelling events, leading to lower chlorophyll and water temperatures. The copepod assemblage was dominated by Oithona spp., Acartia spp. and Pseudocalanus spp., with Metridia pacifica (lucens), Microsetella rosea, Oncaea spp. and Tortanus discaudatus also common during all 3 years. The cross-shelf distribution of copepods was generally shifted offshore during upwelling and onshore during relaxation events, although some variability between species occurred. Abundance of all life stages generally exhibited a negative correlation with cross-shelf transport averaged over at least 1–4 days and lagged by 0–3 days, indicating lower abundances during and immediately after active upwelling. However, copepod nauplii seemed to respond positively to wind events lasting 1–5 days followed by a period of relaxation lasting 6 or 7 days. These rapid rates of change in abundance are probably too great to be due to in situ growth and reproduction alone; physical processes must also play a role. These results suggest a highly dynamic relationship between copepods and upwelling events off northern California, with species-specific responses to upwelling to be expected.  相似文献   

8.
The aim of the research was to investigate the diet of herring at different stages of its life cycle. For that purpose feeding of 0-group and immature herring in the Barents Sea, as well as of mature fish from the Norwegian Sea, was studied. 0-Group herring was sampled in the Barents Sea in August–September 2002–2005 during the international 0-group and trawl-acoustic survey of pelagic fish, as well as during the trawl-acoustic survey of demersal fish in November–December 2003–2004. Stomach samples of immature herring (1–3 years) were collected in late May and early of June 2001 and 2005 in the south-western part of the Barents Sea during the trawl-acoustic survey for young herring. Stomach samples of mature herring were collected in the Norwegian Sea in 1996, 1998, 1999, 2001, and 2002 in the course of the international trawl-acoustic survey of pelagic fish. Feeding intensity of herring of all age groups varied considerably between years and this was probably associated with availability and accessibility of their prey. The 0-group herring was found to have the most diverse diet, including 31 different taxa. In August–September, copepods, euphausiids, Cladocera, and larvae Bivalvia were most frequent in the diet of 0-group herring, but euphausiids and Calanus finmarchicus were the main prey taken. In November–December, euphausiids and tunicates were major prey groups. It was found that C. finmarchicus in the diet of 0-group herring was replaced by larval and adult euphausiids with increasing fish length. C. finmarchicus was the principal prey of immature herring and dominated in the diet of both small and large individuals and mainly older copepodites of C. finmarchicus were taken. Larval and adult euphausiids were found in stomachs of immature herring as well, but their share was not large. The importance of different prey for mature herring in the Norwegian Sea varied depending on the feeding area and length of the herring. On the whole C. finmarchicus and 0-group fish were the most important prey for mature herring diet, but fish prey were only important in a small sampling area. Hyperiids, euphausiids, tunicates, and pteropods were less important prey, and in 2002 herring actively consumed herring fry and redfish larvae.  相似文献   

9.
In this study, seasonal and annual variability in the use of estuarine and ocean beaches by young-of-the-year bluefish, Pomatomus saltatrix, was evaluated by indices of abundance in coastal areas of southern New Jersey (1998–2000). Biological and physical factors measured at specific sites were correlated with bluefish abundance to determine the mechanisms underlying habitat selection. In addition, integrative and discrete indicators of bluefish growth were used to examine spatio-temporal dynamics in habitat quality and its effect on habitat selection by multiple cohorts of bluefish. Intra-annual recruitment to coastal areas of southern New Jersey was episodic, and resulted from the ingress of spring-spawned bluefish (hatch-date April) to estuarine beaches in late May to early June, followed by the recruitment of summer-spawned fish (hatch-date early July) to ocean beaches from July to October. Bluefish utilized estuarine and ocean beaches in a facultative manner that was responsive to dynamics in prey composition and temperature conditions. The recruitment and residency of bluefish in the estuary (1998–1999) and ocean beaches (1998), for example, was coincidental with the presence of the Atlantic silverside Menidia menidia and bay anchovy Anchoa mitchilli, the principal prey species for bluefish occupying these respective habitat-types. Bluefish abundance in the estuary (2000) and ocean beaches (1999–2000) was also correlated with water temperature, with the greatest catches of juveniles coinciding with their optimal growth temperature (24 °C). Bluefish growth, estimated as the slope of age–length relationships and daily specific growth rates, equaled 1.27–2.63 mm fork length (FL) d−1 and 3.8–8.7% body length increase d−1, respectively. The growth of sagittal otoliths was also used as a proxy for changes in bluefish size during and shortly before their time of capture. Accordingly, otolith growth rates of summer-spawned bluefish were greater at ocean beaches relative to the estuary and were explained by the more suitable temperature conditions found at ocean beaches during the mid- to late summer. Notwithstanding the fast growth of oceanic summer-spawned bluefish, individuals spawned in the spring were still larger in absolute body size at the end of the summer growing season (240 and 50–200 mm FL for spring- and summer-spawned bluefish, respectively). The size discrepancy between spring- and summer-spawned bluefish at the onset of autumn migrations and during overwintering periods may account for the differential recruitment success of the respective cohorts.  相似文献   

10.
Estimates of abundance and size of three commercially exploited grunt species indicate ontogenetic changes in habitat utilization concentrate their juveniles within the lagoon of the Bay of La Parguera, Puerto Rico. Eleven biotopes, defined by four benthic structures (reef, mangrove, vegetation beds and unconsolidated sediments) and three geographic zones (inner lagoon, outer lagoon and bank shelf) were sampled randomly by visual surveys. French, bluestriped and white grunt (Haemulon flavolineatum, Haemulon sciurus and Haemulon plumeri) were common in the bay and appeared to exhibit similar life history patterns of cross-shelf migration and habitat selection. Recently settled grunts were dispersed over vegetated and unconsolidated soft-bottom sediments of the bay. The juvenile stage occurred in highest densities in shallow lagoon biotopes among the submerged prop-roots of mangrove stands and on inshore reefs. Length data indicates that grunts migrate offshore to adult habitat via increasingly deep reefs. Indices of biotope nursery function based on standing stock estimates of juveniles identified three biotopes, all within the inner lagoon as essential habitat for juveniles of 5–10 cm length interval. This concentration of juveniles within biotopes of the lagoon could represent a bottleneck to recruitment for grunt stocks. Evidence that quantity and quality of lagoon nurseries may limit recruitment indicates that these areas represent a key component of a marine protected area designed to restore fisheries within the bay.  相似文献   

11.
The spatial distribution of stage-specific abundance and reproduction of the copepod Paracalanus parvus were studied from October 2005 to September 2006 in the Jiaozhou Bay. This copepod occurred continuously in this bay throughout the year. The species reached the lowest abundance in April and peaked in June. From October to December, distribution center mainly occurred in offshore water and at the mouth of the bay. In winter, early copepodites and adults gradually decreased and till February, most of the population was only comprised of CIV–CV stages. Overwintering copepodites matured in March and males tended to mature before female. From May to September, each stage occurred in the population and gradually reached high abundance. Temperature and chlorophyll a (Chl-a) concentration in the three stations can't clearly explain the seasonal variation in stage-specific abundance, so we surmised the important effect of the Yellow Sea. Egg production rate (EPR) reached its lowest in winter and peaked in June at 60.8 eggs female−1 day−1 in nearshore water. In the warming period, EPR in nearshore water was statistically higher and EPR > 10 eggs female−1 day−1 lasted longer than that in offshore water, showing the importance of nearshore water for recruitment of P. parvus. Our study showed that EPR was positively related to temperature and total chlorophyll a in offshore water and mouth of the bay. In nearshore water, the relationships between EPR and temperature and Chl-a in three size fractions were not the same as those in offshore water, suggesting complicated ecosystem in such a eutrophic area in warming period.  相似文献   

12.
Estuarine marshes are known as suitable nursery areas for many marine migrant fishes, such as flounder. The potential nursery value of such habitats was investigated in the Venice lagoon, by using growth and production of 0-group flounder as indicators. Size–frequency distribution analysis was performed on fish samples collected fortnightly, from March 2004 to June 2005, in two marsh sites, Dese and Tessera, differing in their origin and environmental conditions. Samples were mostly composed of juvenile individuals, belonging to 0- and 1-group cohorts (the latter being present in Tessera only). A higher total production, either annual or monthly, and faster growth of 0-group flounder was observed in Dese, associated to a higher ecological performance of 0-group individuals in this site, as indicated by the higher P:B ratio values. Dese is a site located in a marsh complex characterized by the relevant influence of a nearby river, and the observed higher potential nursery value of this marsh area with respect to the other is discussed in the light of the higher trophic status and other environmental conditions in this site. The production results were also compared to those from other estuarine environments commonly acknowledged as important nurseries for European flounder.  相似文献   

13.
A study was performed in the Tyrrhenian Sea in front of S. Rossore Park (Pisa) to ascertain the degree of Cu, Pb, Cd and Cr pollution and to investigate whether the rivers flowing into the sea may be considered potential sources of seawater contamination. Owing to the variability of such a system, the research, covering a one-year period, involved five surveys on 13 selected sampling stations and two surveys following the floats along the river Arno and its plume. The results refer to the total concentration of the above metals in the water phase after filtration. The content of Cu, Pb and Cd was always found to be higher than in the reference station (about 8 km offshore), generally reaching maximum values around the mouths of the rivers. Concentrations exceeding the minimal risk concentration were observed in the case of Cd. As for Cr, its concentration was always below the detection limit of the analytical method used. Rivers thus appear to be the main sources of coastal metal pollution.By the samplings carried out along the river Arno and its plume (involving 3–5 km from the mouth), it was found that the dilution model represents the concentration pattern for Cu and Pb fairly well over a large range of salinity.  相似文献   

14.
Dissolved gaseous mercury (DGM) was measured continuously using two newly developed techniques and a manual technique. The continuous techniques were based on the equilibrium between the aqueous and gaseous phase (DGM = Hgextr / H', Hgextr is the measured mercury concentration in the gas phase, H' is the Henry's Law coefficient at the desired temperature). In order to calculate the annual mercury evasion from the Mediterranean Sea, diurnal and seasonal measurements of DGM, total gaseous mercury in air (TGM), water temperature and wind speed were performed. During August 2003, March–April 2004 and October–November 2004 measurements of these parameters were conducted on board the RV Urania. The continuous measurements of DGM showed a diurnal variation in concentration, at both coastal and off shore sites, with higher concentrations during daytime than nighttime. The concentration difference could be as large as 130 fM between day and night. The degree of saturation was calculated directly from the measurements, S = Hgextr / TGM and was found to vary between the different seasons. The highest average degree of saturation (850%) and the largest variation in saturation (600–1150%) was observed during the summer. The spring showed the lowest variation (260–360%) and the lowest average degree of saturation (320%). The autumn also showed a large variation in saturation (500–1070%) but a lower average (740%) compared to the summer cruise. This might be explained by the temperature difference between the different seasons, since that parameter varied the most. The flux from the sea surface was calculated using the gas exchange model developed by Nightingale et al. [Nightingale, P.D., Malin, G., Law, C.S., Watson, A.J., Liss, P.S., Liddicoat, M.I., Boutin, J., Upstill-Goddard, R. C., 2000. In situ evaluation of air–sea gas exchange parameterization using novel conservative and volatile tracers. Global Biogeochemical Cycles, 14(1):373–387]. The evasion varied between the different seasons with the highest evasion during the autumn, 24.6 pmol m− 2 h− 1. The summer value was estimated to 22.3 pmol m− 2 h− 1 and the spring to 7.6 pmol m− 2 h− 1. Using this data the yearly evasion from the Mediterranean Sea surface was estimated to 77 tons.  相似文献   

15.
Control of walleye pollock (Theragra chalcogramma) recruitment in the Eastern Bering Sea involves complex interactions between bottom-up and top-down processes, although the mechanisms are poorly understood. We used statistical models to test the leading hypotheses linking recruitment variability to biotic and abiotic factors. Consistent with a “cold-pool hypothesis”, recruitment of pollock was significantly stronger if winters preceding the larval (age-0) and juvenile stages (age-1) were mild. However, our results did not support the proposed top-down mechanism (cannibalism) underlying this hypothesis. Several empirical relationships support an “oscillating control hypothesis”. As predicted by it, the effect of ice conditions on survival during the larval and early juvenile stages was modified by the abundance of adult pollock, implying stronger bottom-up control when adult abundance (hence cannibalism) was low. The proposed bottom-up mechanism predicts that the survival of pelagic-feeding walleye pollock (benthic-feeding yellowfin sole), should be higher during years with an early (late) ice retreat, which was confirmed by our analysis. Our results also provide additional evidence for a “larval transport hypothesis”, which states that cannibalism of larval and juvenile pollock is reduced in years when strong northward advection separates juveniles from cannibalistic adults.In addition to testing existing hypotheses, we identified new relationships between spawner-to-recruit survival rates of walleye pollock and several indicators of mixed layer dynamics during the spring and summer. Survival rates and recruitment were significantly reduced when larval or early juvenile stages experienced a delay in the (non-ice-associated) spring bloom as a result of stormy spring conditions, suggesting that the timing of the spring bloom is critical to both first-feeding larvae and age-1 juveniles. Furthermore, a dome-shaped relationship between pollock survival and summer wind mixing at the early juvenile stage is consistent with modeling and laboratory studies showing an increase in survival at low to moderate levels of wind mixing, but a decrease in feeding success at high levels of wind mixing.Top-down controls also regulate recruitment of walleye pollock. At least one-third of the variability in spawner-to-recruit survival could be accounted for by predation mortality at the early juvenile stage (age-1). Predation of juvenile pollock can be attributed largely to cannibalism, which varies with the abundance of adult pollock and with the availability of juveniles to adult predators. A simple index reflecting the spatial overlap between juvenile and adult pollock explained 30–50% of the overall variability in recruitment, similar to the variability explained by the best environmental predictors. Although environmental effects are difficult to separate from the effects of predation, we conclude that bottom-up and top-down processes are equally important in controlling the survival of pollock from spawning to recruitment at age 2. However, the magnitude of top-down control is itself modified by environmental factors that control the availability of juvenile pollock to adults (through impacts on spatial distribution) and the abundance of adult predators (through effects on productivity and carrying capacity).  相似文献   

16.
On the basis of the climatic data array on temperature and salinity including the data of observations carried out at 102,000 oceanographic stations in 1910–1998, we estimate the amount of available potential energy of large-scale processes in the 0–300-m layer of the Black Sea and study its annual variability. The annual average amount of available potential energy in the active 0–300-m layer is equal to 13.5×1014 J, its maximum amount attained in March is 25×1014 J, and its minimum amount attained in August is 8×1014 J. In the autumn–winter period, the density of available potential energy in the 0–50-m layer increases and its maximum is attained in December–January (30 J/m3). Beginning with January, the density of available potential energy increases in the layer of the main pycnocline (50–150 m). Its maximum is attained in March (50 J/m3) and then this quantity decreases till August (down to 14 J/m3). In August, we study the interdecadal variability of the available potential energy in the 50–300-m layer by using the data accumulated for decadal periods with five-year shifts in 1956–1995. The maximum variability is observed at a depth of 100 m. The maximum average (over the sea) amounts of available potential energy were observed in 1961–1970 and 1986–1995. The minimum amounts were recorded in 1976–1985.  相似文献   

17.
The food choice of juvenile flounder (Platichthys flesus) and turbot (Scophthalmus maximus) was studied in the northern Baltic Sea during the years 1988, 1989, 1994 and 1995. The diet included organisms from 30 species/taxa in flounder (n = 306) and 10 species/taxa in turbot (n = 41). Flounder 45 mm mainly consumed meiofauna (dominating taxon: Harpacticoida, Copepoda) and larger fish (46–101 mm) consumed macrofauna (dominating taxa: Oligochaeta, Amphipoda and Chironomidae). In terms of biomass, macrofauna dominated for all sizes of flounders, and meiofauna was important only for the smallest fish. A strong seasonal variation could be detected in the diet. In spring, macrofauna dominated for all size classes of fish (only fish > 30 mm were caught in spring), while in summer and autumn meiofauna dominated the diets for fish 45 mm in size. Juvenile turbot (22–88 mm) consumed macrofauna and small fish. Turbot 30 mm consumed mainly amphipods, while > 30 mm turbot consumed mysid shrimps, amphipods and fish.The ontogenetic shift from meio- to macrofauna-sized prey in flounders occurs at a larger fish size in the northern Baltic Sea than reported in other areas, possibly depending on the increased relative importance of meiofauna in the northern Baltic. The seasonal variation in the diet could be due to seasonally changing abundances in the zoobenthos, or for the small fish (1-group, spring), to switching from meio- to macrofauna in order to optimize their energy gain. The 0-group flounders consumed meiofauna for a long period, possibly due to a learning-process or simply due to easy availability of meiofauna. Turbot has a much larger mouth gap than flounders, thus allowing them to consume macrofauna from the beginning of their benthic life.  相似文献   

18.
The North Atlantic margins are archetypally passive, yet they have experienced post-rift vertical movements of up to kilometre scale. The Cenozoic history of such movements along the NW European margin, from Ireland to mid-Norway, is examined by integrating published analyses of uplift and subsidence with higher resolution tectono-stratigraphic indicators of relative movements (including results from the STRATAGEM project). Three episodes of epeirogenic movement are identified, in the early, mid- and late Cenozoic, distinct from at least one phase of compressive tectonism. Two forms of epeirogenic movement are recognised, referred to as tilting (coeval subsidence and uplift, rotations <1° over distances of 100s of Kilometres) and sagging (strongly differential subsidence, rotations up to 4° over distances <100 km). Each epeirogenic episode involved relatively rapid (<10 Ma) km-scale tectonic movements that drove major changes in patterns of sedimentation to find expression in regional unconformity-bounded stratigraphic units. Early Cenozoic tilting (late Paleocene to early Eocene, c. 60–50 Ma) caused the basinward progradation of shelf-slope wedges from elongate uplifts along the inner continental margin and from offshore highs. Mid-Cenozoic sagging (late Eocene to early Oligocene, c. 35–25 Ma) ended wedge progradation and caused the onset of contourite deposition in deep-water basins. Late Cenozoic tilting (early Pliocene to present, <4±0.5 Ma) again caused the basinward progradation of shelf-slope wedges, from uplifts along the inner margin (including broad dome-like features) and from offshore highs. The early, mid- and late Cenozoic epeirogenic episodes coincided with Atlantic plate reorganisations, but the observed km-scale tectonic movements are too large to be accounted for as flexural deflections due to intra-plate stress variations. Mantle–lithosphere interactions are implied, but the succession of epeirogenic episodes, of differing form, are difficult to reconcile with the various syn-to post-rift mechanisms of permanent and/or transient movements proposed in the hypothetical context of a plume beneath Iceland. The epeirogenic movements can be explained as dynamic topographic responses to changing forms of small-scale convective flow in the upper mantle: tilting as coeval upwelling and downwelling above an edge-driven convection cell, sagging as a loss of dynamic support above a former upwelling. The inferred Cenozoic succession of epeirogenic tilting, sagging and tilting is proposed to record the episodic evolution of upper mantle convection during ocean opening, a process that may also be the underlying cause of plate reorganisations. The postulated episodes of flow reorganisation in the NE Atlantic region have testable implications for epeirogenic movements along the adjacent oceanic spreading ridge and conjugate continental margin, as well as on other Atlantic-type ‘passive’ margins.  相似文献   

19.
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20.
As part of a joint workshop organised by the International Council for the Exploration of the Sea (ICES) and the Intergovernmental Oceanographic Commission (IOC) on biological effects monitoring techniques. dad (Limanda limanda) were examined from six spaced stations along a 200 km transect extending from near the mouth of the Elbe River out to the Dogger Bank in the North Sea. Based on historical data, differences in contaminant concentrations in sediments exist along the transect (lipophilic organic xenobiotics at the inshore sites and heavy metals offshore over the Dogger Bank). The most contaminated site sampled was the most inshore, the least contaminated was approximately midway along the transect, with contamination building up again over the farthest point along the transect, over the Dogger Bank. Multiple organs and tissues were examined for full pathology from each fish sampled. Only liver data are presented here (11–20 livers per station). The most significant lesions were considered to be well-developed foci of cellular alteration, high mitotic activity and high neutral lipid accumulation in livers from dab sampled from the most inshore site examined. Livers from the least-contaminated station showed minimal evidence of such changes. Foci of cellular alteration and neutral lipid accumulation were also seen in dab liver sampled from fish from the Dogger Bank site. Thus far, the hepatic changes seen correlate well with the most contaminated sites along the transect. The value of comprehensively examining the histopathology of an organ of toxicological significance, such as the liver in a European species of flatfish, is demonstrated.  相似文献   

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