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1.
The Lower Cambrian Chengjiang Lagerstatte provides a good window to explore the origin and radiation of early bilaterians. Here we describe a netted sclerite-bearing worm Tabelliscolex hexagonus gen. et sp. nov., and tentatively assign it to palaeoscolecidans of priapulids. The cuticle of the animal is covered with two kinds of platy sclerites which are constructed by hexagonally arranged tubercles. Similar structures of the sclerites can be seen on some Cambrian palaeoscolecidans and lobopods, so, this new species is critical for understanding the relationships between lobopods and palaeoscolecidans.  相似文献   

2.
Abstract The Lower Cambrian Chengjiang Lagerstätte provides a good window to explore the origin and radiation of early bilaterians. Here we describe a netted sclerite‐bearing worm Tabelliscolex hexagonus gen. et sp. nov., and tentatively assign it to palaeoscolecidans of priapulids. The cuticle of the animal is covered with two kinds of platy sclerites which are constructed by hexagonally arranged tubercles. Similar structures of the sclerites can be seen on some Cambrian palaeoscolecidans and lobopods, so, this new species is critical for understanding the relationships between lobopods and palaeoscolecidans.  相似文献   

3.
Tylotites petiolaris吻部构造的发现   总被引:1,自引:0,他引:1  
早寒武世澄江动物群提供了探索早期翻吻动物辐射的一个绝佳的窗口。澄江动物群翻吻动物包括8科10属10种,其中罗惠麟等(1999)依据仅有的一块不完整的标本建立Tylotites petiolaris,并将其归入叶足动物。2002年西北大学早期生命研究所野外工作组在尖山剖面采集到两块完整的具有吻部的标本,本文籍此对该属进行了进一步完善和补充,特别是吻部和躯干表面的特征,从而说明T.petiolaris为翻吻动物。因为与澄江动物群中其他翻吻动物形态有较大差异,据此建立瘤节虫科,新科Tylotitidae fam.nov.。描述如下:虫体较长,呈圆管状,长可达100mm,宽2~4mm。身体基本可以分吻部和躯干部两部分。吻部与Cricocosmia特别相似,为典型的三段式构造,从下到上可依次分为Ⅰ、Ⅱ、Ⅲ段,Ⅰ段与躯干前端连接,直径同于躯干,表面有10列纵向的略为弯曲的小刺(图版Ⅰ,图1a,1b;插图1)。最前端吻刺最为粗壮,向后的吻刺逐渐变小。Ⅱ段位于Ⅰ段之前,向前略微收缩,光滑无刺。Ⅲ段位于吻部顶端,可进一步分为A、B两部分:A段直径后端大于Ⅱ段,表面密布斜列排列的小刺,从后向前,小刺变大;B段光滑无饰,直径小于A段。Ⅲ段可强烈自由弯曲(图版Ⅰ,图1a,1d)。颈部长约5mm,表面光滑无饰,直径略小于躯干。躯干表面具56~58个环脊,52排粗刺,最前面第4~6个环脊表面无刺,环脊之间为环沟。躯干由前向后直径逐渐增加,在中后部增至最大,然后虫体直径基本上一致。躯干刺成行成列排列。环状分布的躯干刺随着环脊与环沟落差的变化而改变,到躯干后部逐渐减小。每一个环脊之上有12个粗刺,腹刺长度小于背刺。有些躯干刺的轴部显示较强的黄铁矿化,表明其可能为中空结构(图版Ⅰ,图1c),可能有体腔伸入,躯干末端具有一对尾刺,躯干整体弯曲成6字形(图版Ⅰ,图1a,1b)。肠管从吻部一直延伸到躯干末端。肠道位于一个空闻的体腔内,Ⅰ段表面宽约1mm的黑色区域可能为胃,肛门位于躯干末端的腹面(图版Ⅰ,图1b)。T.petiolaris很可能表栖生活于海底底面上。因为体表长刺具有很强的防御作用,腹刺在运动过程中可以抓牢沉积物,所以,很可能通过背腹波动在海底底面作蠕形运动,也不排除蜷曲一解蜷的运动方式。吻部的主要功能可能为捕食。几条T.petiolaris个体相互盘绕(图版Ⅰ,图2),很可能与其两性交配有关。  相似文献   

4.
赵新伟  曾伟 《中国地质》2016,(3):921-935
层序地层学是地层学研究中的第三次革命,其强调年代地层框架内的沉积相关系和地层结构。河北路通沟剖面寒武系地层连续,层序界面清楚而独特,沉积现象丰富,是进行华北地台寒武系层序地层划分的典型地点之一。该剖面岩相类型以泥晶灰岩、鮞粒灰岩及白云岩为主,发育潮坪相、颗粒滩相、缓坡相及陆棚相等沉积相。在新的年代地层框架下,依据沉积相的垂向叠加样式及其反映的长周期旋回特征,将路通沟剖面寒武系划分为9个三级层序,以第三统徐庄组组成的三级层序(TST+CS+HST)为界,整个沉积序列从混积潮坪相沉积变为缓坡型碳酸盐岩沉积,层序也从以"TST+HST"为主的沉积序列转变为以"CS+HST"为特征的淹没不整合型层序序列。芙蓉统中众多的叠层石生物丘暗示:寒武系末期贫乏骨骼的风暴海洋中微生物碳酸盐岩随后生动物辐射也在增加的特殊沉积现象。基于沉积相叠置方式及旋回特征对路通沟剖面寒武系的层序地层研究,将为探索华北地台东北部广泛分布的寒武系地层的层序划分提供一个典型的实例,为今后大范围的追索和对比提供更多的思考途径。  相似文献   

5.
Cambrian Fossil Embryos from Western Hunan,South China   总被引:1,自引:1,他引:0  
The exquisitely preserved fossil embryos of Markuelia recovered from the limestones of the Middle Cambrian Haoqiao Formation and Upper Cambrian Bitiao formation in western Hunan, South China are described and illustrated in detail for the first time.A new species Markuelia elegans sp.nov.is established based mainly on embryos from the Upper Cambrian.A few of animal's resting eggs,which are comparable with those of the Ediacaran Doushantuo Formation,have been also found in the Upper Cambrian of western Hu...  相似文献   

6.
Exceptionally preserved fossils,such as those from Cambrian Burgess Shale-type fossil-Lagerstatten are critical because of their unique contributions to knowledge of the phylogenetic radiation and palaeoecological expansion of metazoans during the Cambrian explosion.Critically,these deposits provide information that is usually unobtainable from shelly and skeletonized fossils alone.The Guanshan Biota(Cambrian Series 2,Stage 4)in the Yunnan Province of South China,has produced abundant and diverse,exquisitely preserved fossils that often retain soft tissues and organs.To date,most fossils from the Guanshan Biota have been collected from localities such as Gaoloufang and Gangtoucun,which have become inaccessible due to new urban expansions and constructions of residential buildings.Here we present the first report of soft bodied fossils from a new section at Kanfuqing,close to the Wulongqing village in Malong County,approximately 3 km east of the Wulongqing Formation stratotype section.Fossils retain soft morphology,and include brachiopods with delicate marginal setae,priapulids with well-preserved sclerites and vetulicolians with entire sections of body.In addition,this fauna includes rare occurrences of trilobites preserved with soft tissues replicated as pyrite pseudomorphs after weathering.This discovery represents an important palaeogeographical extension of soft-bodied fossils of the Guanshan fauna to the east of the Xiaojiang Fault(related to Tsinning tectonic movements ca.700 Ma).The fauna from the new Kanfuqing section is similar to that reported from the Wulongqing Formation west of the Xiaojiang Fault,and thus has significant implications for early Cambrian palaeogeography,faunal successions and palaeoenvironments of eastern Yunnan.  相似文献   

7.
库鲁克塔格地区保存了相对完整的寒武纪-奥陶纪沉积地层,其沉积特征及演化规律的研究,对塔东地区寒武系-奥陶系油气勘探具有重要意义。通过库鲁克塔格地区详细的野外地质调查,结合前人相关研究成果,对该区寒武系-奥陶系沉积体系、沉积模式及演化规律的研究表明,寒武纪时期,库鲁克塔格地区经历了一次大的海侵-海退旋回,南、北两区沉积具有相似性。寒武纪早期的快速海侵导致南、北两区均发育陆棚相-深水盆地相沉积;寒武纪晚期,在逐渐海退的背景下,南、北两区开始出现沉积分异。奥陶纪,经历了新一轮大的海侵-海退旋回,南北两区沉积差异显著。北区从早奥陶世到晚奥陶世,发育台地边缘斜坡相-广海陆棚相-缓斜坡相-台地边缘礁滩相-开阔台地相相序,构成整体向上变浅的碳酸盐岩沉积层序;而南区发育深水盆地相-陆棚斜坡相-浊流盆地相-碎屑陆棚相相序,形成一套巨厚的深水复理石建造。库鲁克塔格地区寒武纪时期发育缓坡型碳酸盐岩台地,因台地不断向南构筑以及断裂活动,导致奥陶纪晚期台地边缘快速变陡,并在经历斜坡相快速堆积填平补齐之后,重新演变为缓坡型碳酸盐岩台地。晚奥陶世,由于周缘构造活动影响,却尔却克山-雅尔当山一带下沉,逐渐向远端变陡缓坡型碳酸盐岩台地演化。  相似文献   

8.
The radiation of macroscopic animals in the Early Cambrian, commonly be called as ‘Cambrian explosion’, had a close relationship with Earth oxygenation. These macrofossils are widely preserved within the Early Cambrian in the northern and western Yangtze Craton. In order to show the casual relationship between animal evolution and Earth redox environment, Mo content and isotopic composition analyses have been conducted on dolomites of the Tianzhushan Formation and the black shales of the Shuijingtuo Formation from the lowermost Cambrian profile in the eastern Huangling dome, northern Yangtze Craton. The calcitic dolostones from the Tianzhushan Formation display lower concentrations of Mo contents (0.055–0.666 ppm) than that of the black silty shales from the Shuijingtuo Formation (6.0–172 ppm). This magnitude difference is most likely related to the different lithologies due to Mo enrichment in shales. The Mo isotope values (δ98/95Mo) of the calcitic dolostones for the Tianzhushan Formation vary from −0.73‰ to 0.41‰. In comparison, the black sandy shales from the Shuijingtuo Formation exhibit δ98/95Mo of −1.04–1.84‰, including 5 samples from the earliest Shuijingtuo Formation displaying systematically higher δ98/95Mo values (0.09–1.84‰) than those of the other 5 samples from upper layers (δ98/95Mo = −1.04 to −0.19‰). These geochemical data suggest that the redox environment of the early Cambrian is fluctuating, and the peak oxygenation of the ocean had already reached a level similar to modern environments. According to the Mo concentrations and δ98/95Mo data, the sedimentary succession can be divided into five intervals, which are closely correlated to the episodic evolution of early life in the Cambrian. Together with the available published data from other profiles in the Yangtze Craton, our new data reveal that the higher oxidation environment had a positive effect on the evolution of early life.  相似文献   

9.
寒武纪年代地层的研究现状和研究方向   总被引:9,自引:6,他引:3  
全球寒武系的底、顶界层型已被确定 ,但寒武系内的统和阶一级的全球标准划分尚待进行。介绍了国际地层委员会寒武系分会有关建阶的意见和推荐的 13个分阶的层位 ;同时介绍了劳亚大陆新近提出的 4统 6阶的年代地层划分方案。回顾了我国的建阶过程 ,指出我国现有的阶一级单位实质上仍然是岩石地层单位 ,具有建立在单位层型上、底界被粗略界定因而时间意义不明确、对比性差、与岩石地层单位重名和不利于竞争国际标准阶和GSSPs等一系列缺点。在湘西、黔东地区新建的 5个阶可取代我国现有的中—晚寒武世年代地层系统。建议暂将我国寒武系划分为 3统和 9个阶 ,即 :下寒武统梅树村阶、筇竹寺阶、沧浪铺阶和龙王庙阶 ;中寒武统台江阶和王村阶 ;上寒武统酉水阶、瓦儿岗阶和桃源阶。  相似文献   

10.
刺胞动物是华南寒武纪早期海洋生态系统的重要组成部分,针对这类化石的研究有助于深入了解寒武纪多幕式爆发事件。本文主要根据寒武纪早期扬子板块陕南宽川铺生物群、湖北岩家河生物群以及澄江生物群产出的刺胞动物,探讨刺胞动物在寒武纪早期的演化框架,取得了以下初步认识。截至目前,所发现的寒武纪早期刺胞动物绝大多数都属于水母超纲,珊瑚纲的化石记录在寒武系幸运阶、第二阶保存较为罕见。其中寒武系幸运阶刺胞动物的化石记录全部属于底栖类型,体型小,且具有多种类型的身体辐射对称形式,绝大多数类型可以确定属于直接发育。第二阶刺胞动物仍以底栖固着类型为主,体形增大,仅见两、四辐射对称形式;而寒武系第三阶刺胞动物仍有底栖固着类型,但游泳的水母开始出现,体型增大明显,代表水母冠群的出现,以及世代交替的复杂生命周期的正式建立。从幸运阶到第三阶刺胞动物体型明显增大,可能与海水氧气含量增加有关。在寒武纪大爆发的背景下,刺胞动物的分异度和丰度在寒武系幸运阶就已经达到顶峰,然后在第二阶、第三阶开始衰减,这种变化可能与两侧对称动物辐射演化有关。华南刺胞动物的化石记录表明,寒武纪大爆发并非纯粹的一幕式爆发,其中充斥着一系列生物群的替代甚至灭绝事件。  相似文献   

11.
A carbon isotope stratigraphy from the lowest Cambrian strata in the Adelaide fold belt of South Australia is interpreted in terms of the geological context of the sequence: major transgression, developing circulation patterns and upwelling resulting from an adjacent opening ocean, and biomass increases stemming from the Cambrian radiation event. There is also evidence from the textures of ooids and cements in late Precambrian-Cambrian strata for subtle variations in seawater chemistry across the boundary and these may well have been instrumental in the development of calcareous skeletons in the early metazoans.  相似文献   

12.
埃迪卡拉纪—寒武纪转换时期小壳化石的突然出现是世人瞩目的重要演化事件之一,它见证了动物的爆发式快速演化过程,标志着以两侧对称动物为特征的寒武纪生态系统的开始。湖北三峡地区寒武系纽芬兰统岩家河组处于幸运阶—第二阶这一关键层段,是研究上述问题的理想地区之一。岩家河组小壳化石研究最近取得的进展,为寒武系纽芬兰统幸运阶/第二阶界线的标定,和寒武纪早期软体动物和刺胞动物的演化及个体发育提供了重要的化石依据,本文对其进行了总结。全球界线层型剖面和点位(GSSP)寒武系第二阶底界划分标志的Watsonella crosbyiAldanella attleborensis在岩家河组第5段底部的同时出现,意味着岩家河组第5段已归属寒武系第二阶。这两类软体动物化石均广泛分布于华南、蒙古、西伯利亚和北美等大陆碳酸盐岩相寒武系第二阶地层中,它们均是定义寒武系第二阶底界的有力竞争者。另外,W.crosbyi壳顶下方一对肌肉附着结构的发现提供了重要的软组织结构信息,证实了W.crosbyi为内腹型壳,归属为未扭曲的软体动物太阳女神螺类。新发现六方锥石类Septuconularia yanjiaheensis具侧向压扁、两辐射对称的、14个等大锥面的围鞘,是迄今所描述的六方锥石类中锥面数最多的类型。根据横肋疏密程度,围鞘由下向上可分为三部分,可能代表胚胎期、幼年期和成年期不同的生长阶段。Septuconularia显然由寒武系幸运阶Hexaconularia通过扇面扩增演变而来,证明Hexaconularia可能是ArthrochitesSeptuconularia之间的中间过渡类型。新发现的橄榄蛋类Octapyrgites elongatus与幸运阶OlivooidesQuadrapyrgites十分相似,围鞘由方形的壳顶区和波纹状倒宝塔形的远顶区组成。然而Octapyrgites只有8个口折叶,有别于具12个口折叶的Quadrapyrgites。通过与幸运期水母类多种对称方式对比,发现在寒武系第二阶橄榄蛋类数量减少和五辐射对称骨状壳类缺失,表明在幸运期/第二期转换时期,底栖固着型的刺胞动物水母类多样性发生了明显衰落,甚至部分绝灭,而同期的两侧对称动物则迅速辐射演化。  相似文献   

13.
通过对塔东地区古城4井上寒武统和中、上奥陶统碳酸盐岩围岩及充填物的碳、氧、锶同位素地球化学和流体包裹体成分的对比研究表明,充填于中、上奥陶统和上寒武统的流体分属于2个不同来源的流体体系。充填于中、上奥陶统灰岩裂缝中方解石脉的w(87Sr)/w(86Sr)介于0.7084~0.7088,它与早奥陶世海水的w(87Sr)/w(86Sr)相近;流体体系为CH4-H2O体系;充填物与围岩间具有明显的碳、氧同位素差异,表明上部流体体系中的流体来自于奥陶系地层本身。上寒武统白云岩裂缝中方解石脉的w(87Sr)/w(86Sr)为0.7138,明显地高于同时代海水的锶同位素值;流体体系为CO2-H2O体系;下部流体体系中的流体为外来富锶流体。上、下流体体系间互不连通暗示着上寒武统地层具有相对较好的保存条件。  相似文献   

14.
Early Cambrian radiation of brachiopods: A perspective from South China   总被引:6,自引:4,他引:2  
Brachiopods, a group of benthic suspension-feeding marine invertebrates, made their first appearance in the Lower Cambrian. In the Yangtze Platform (South China), well-exposed Lower Cambrian stratigraphic succession represents shallow to deeper water environments. Strata from eastern Yunnan, southern Shaanxi and the Yangtze gorges areas of western Hubei Province, deposited in muddy-siltstone and carbonate lithofacies, contained an abundant variety of brachiopods, including all the representatives of the subphylum Linguliformea and the calcareous-shelled genera of Kutorgina and Nisusia from the subphylum Rhynchonelliformea. Thus the fossil assemblage bears witness to the first major phase of evolutionary radiation of brachiopods during the ‘Cambrian explosion’ interval of metazoans. Brachiopods from the celebrated Chengjiang fauna have exquisitely preserved soft-tissues, which reveal the body plans and evolutionary acquisition of morphological novelties of the early stocks, and also provide a good opportunity for testing the analogies with the stem groups from the extant representatives. These fossils have corroborated the view that brachiopods developed complex organization of tissues, and achieved considerable evolutionary success already by the onset of ‘Cambrian Explosion’. Thus it is not improbable that a large part of this radiation occurred within, or only just before early Cambrian time. Studies of Chengjiang brachiopods suggest that attachment by a pedicle to the substrate was probably the most common relationship of Cambrian brachiopods with the substrate where they inhabited.  相似文献   

15.
李才  吴彦旺  王明  杨韩涛 《地质通报》2010,29(12):1733-1736
在申扎地区发现了较为完整的寒武纪地层和奥陶系与寒武系之间的"泛非运动"角度不整合。这是迄今为止青藏高原南部发现的唯一确切的寒武纪地层,不但进一步丰富和健全了青藏高原的地层系统,而且也是在青藏高原和周边发现的唯一的下奥陶统与寒武系"泛非运动"角度不整合界面。通过对申扎"泛非运动"界面的研究,可以准确界定"泛非运动"的时限及其动力学过程,也是研究"泛非运动"之前冈瓦纳大陆北缘沉积作用不可替代的标本,对青藏高原构造演化研究具有重要、深远的意义。  相似文献   

16.
文章描述了云南寒武纪早期澄江生物群小细丝海绵属一新种--螺旋小细丝海绵(Leptomitella spiralis sp. nov.)。新种 具有横向骨针束围绕海绵体呈螺旋状排列的典型特征,明显区别于属内其他种;此外,在外部形态和骨针类型上与属内其他种 也存在较大差异。新种的发现增加了Leptomitella种间形态结构的多样性,也为海绵动物的早期演化及寒武纪海绵动物辐射演 化研究提供新的化石资料。  相似文献   

17.
文章描述了云南寒武纪早期澄江生物群小细丝海绵属一新种——螺旋小细丝海绵(Leptomitella spiralis sp. nov.)。新种 具有横向骨针束围绕海绵体呈螺旋状排列的典型特征,明显区别于属内其他种;此外,在外部形态和骨针类型上与属内其他种 也存在较大差异。新种的发现增加了Leptomitella种间形态结构的多样性,也为海绵动物的早期演化及寒武纪海绵动物辐射演 化研究提供新的化石资料。  相似文献   

18.
《China Geology》2021,4(2):205-214
During the Late Neoproterozoic and Early Cambrian, a series of paleogeographic and paleoecological events occurred associated with deglaciations and the evolution of life. The appearance of fossils representing diverse phyla, novel body plans and complex ecologies in Ediacaran and Cambrian sedimentary successions has sparked diverse hypotheses about potential drivers for the radiation of early animals during this interval. Recently, new macroscopic fossils of carbonaceous compressions with unique features have been found in Anning, Yunnan, China. The fossils’ bodies are ribbon-shaped and bilaterally symmetric, with dense longitudinal features and transversal features. The fossils occur 0.68 m below a bentonite interlayer which has been dated 535.2 ± 1.7 Ma by Ri-xiang Zhu and his team in 2009. The relatively simply morphology of these fossils, coupled with a lack of preserved internal structures challenge efforts taxonomically identify the precursor organism and definitively ally it to a living group. However, the symmetry and unusual features of the body are analogous to members of Platyzoa. The presence of ribbon-shaped fossils in the Zhongyicun Member in Anning indicates that these organisms were at least a locally significant component of Cambrian seafloor ecosystems, and may hold important implications for our understanding of the early evolution of Bilateria.  相似文献   

19.
Five species, Lipopora lissa Jell and Jell, 1976, Lipopora daseia Jell and Jell, 1976, Tretocylichne perplexa Engelbretsen, 1993 from Australia, Cambroctoconus orientalis Park, Woo, Lee, Lee, Lee, Han and Chough, 2011 from China, and Cambroctoconus kyrgyzstanicus Peel, 2014 from Kyrgyzstan, belonging to the Cambrian stem-group cnidarians have been documented in the fossil record. Cambroctoconus coreaensis sp. nov., interpreted here as a stem-group cnidarian, from the Seokgaejae section in the Daegi Formation, Taebaek Group (Cambrian Series 3), Taebaeksan Basin, central-eastern Korean Peninsula, has a slender cup-shaped skeleton. A cladistic analysis produced 21 most parsimonious trees, which invariably placed the six stem-group cnidarians below the crown-group, but their relationships within the stem-group are unresolved. Nine out of the 21 trees suggest a monophyletic relationship for the Cambrian stem-group cnidarians, whereas in six other trees a monophyly of Cambroctoconus and Tretocylichne appeared as the sister-group to the crown-group cnidarians with Lipopora at the most basal branch. This result may reflect the fact that crown-group cnidarians evolved in the Precambrian, and suggests that the diversity of stem-group cnidarians was a result of an independent radiation in the Cambrian.  相似文献   

20.
寒武纪生命扩张及澄江动物群的意义   总被引:1,自引:0,他引:1  
生命在地球上的出现及其演化有较长的历史。地球的物理、化学条件是早期生命出现的外因。早寒武世西南地区地质、气候及富含营养的浅海水域是澄江动物群在后生动物演化的关键时期出现的外因。“寒武纪生物的扩张”应有恰当的翻译,同时对现代生物学的“适应辐射”应有清楚的中文解释。从新近发现看,早、中寒武世的澄江动物群及布吉斯页岩动物群与埃迪卡拉动物群具有一些联系,埃迪卡拉动物群并没有在寒武纪时完全绝灭。  相似文献   

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