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1.
We present a database of late-Quaternary plant macrofossil records for northern Eurasia (from 23° to 180°E and 46° to 76°N) comprising 281 localities, over 2300 samples and over 13,000 individual records. Samples are individually radiocarbon dated or are assigned ages via age models fitted to sequences of calibrated radiocarbon dates within a section. Tree species characteristic of modern northern forests (e.g. Picea, Larix, tree-Betula) are recorded at least intermittently from prior to the last glacial maximum (LGM), through the LGM and Lateglacial, to the Holocene, and some records locate trees close to the limits of the Scandinavian ice sheet, supporting the hypothesis that some taxa persisted in northern refugia during the last glacial cycle. Northern trees show differing spatio-temporal patterns across Siberia: deciduous trees were widespread in the Lateglacial, with individuals occurring across much of their contemporary ranges, while evergreen conifers expanded northwards to their range limits in the Holocene.  相似文献   

2.
The mandible of a polar bear (Ursus maritimus Phipps) found in about 1920 at Kjul Å, North Jutland, and described by Nordmann & Degerbol in 1930. has been l4C dated to 11.100 ± 160 B.P. It is so far the only find of polar bear in Denmark. Comparison with recent 14C datings of Swedish and Norwegian polar bears shows that the Danish specimen was a member of a southern Scandinavian Late Weichselian population. The contemporaneous Zirphaeu sea deposits can be regarded as the boreal-arctic shallow water equivalent of the arctic Upper Saxicava sand deposits from northern Jutland. The polar bear mandible, however, was deposited on land, as was the metacarpal bone of a brown bear ( Ursus arctos ) from the nearby Nr. Lyngby locality of Allerød age. The overall picture of the Late Weichselian mammal fauna in Denmark shows a mixed composition of different ecotypes. Their sympatric occurrence points at a unique environment not comparable to any now existing, and probably related to the very low latitude of the Weichselian ice sheet.  相似文献   

3.
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4.
Norwegian caves contain stratigraphical information regarding the historical faunal composition valuable in the assessment of postglacial colonization and phylogeographical structure. In some of these limestone caves, brown bear ( Ursus arctos L.) remains have been excavated and radiometrically dated. We present osteology (95 traits defined) from 29 brown bears found within 22 Norwegian caves above 658N. Our data span the time interval 6210–420 14C yr BP. The sex of individuals was based on dimorphic canines, while age was determined by cementum analysis and body size estimated from regressions between morphological traits compared to a contemporary reference collection. Five females and 14 males were recognized, while the remaining bears could not be sexed because of their small size, low age or lack of canines. The ages of 26 bears (tooth cementum age and estimates) ranged from juveniles (<0.5 year) to adults (23 years), the majority being old animals. Both sexes have used these caves for over-wintering lairs, as seen in several caves, where denning is also suggested based on the observations of adult females and cubs. There were no signs of predator or human transported bear remains, suggesting natural mortality. The age of the bears suggests that young or old bears may have died from insufficient nutrient storage, environmental harshness, or may have drowned in spring or autumnal flooding. We conclude that Norwegian brown bears have used these caves sporadically for thousands of years, which suggests that caves are not a preferred site for the brown bear during hibernation or denning.  相似文献   

5.
Carbon dioxide records from polar ice cores and marine ocean sediments indicate that the last glacial maximum (LGM) atmosphere CO2 content was 80–90 ppm lower than the mid-Holocene. This represents a transfer of over 160 GtC into the atmosphere since the LGM. Palaeovegetation studies suggest that up to 1350 GtC was transferred from the oceans to the terrestrial biosphere at the end of the last glacial. Evidence from carbon isotopes in deep sea sediments, however, indicates a smaller shift of between 400 and 700 GtC. To understand the functioning of the carbon cycle this apparent discrepancy needs to be resolved. Thus, older data have been reassessed, new data provided and the potential errors of both methods estimated. New estimates of the expansion of terrestrial biomass between the LGM and mid-Holocene are 700 GtC ± > 300 GtC, using the ocean carbon isotope-based method, compared with of 1100 GtC ± > 500 GtC using the palaeovegetation estimate. If these estimates of the carbon shift to the terrestrial biosphere are equilibrated with the dissolved carbon in the oceans, and the CaCO3 compensation of the ocean is taken into account, then the glacial atmospheric CO2 would have been between 50 (± 30) ppm and 95 (± 50) ppm higher. The glacial atmosphere therefore should have had a CO2 partial pressure of between 330 and 375 μatm. Hence, a rise of between 130 and 175 μatm in atmospheric CO2, rather than 80 μatm, at the end of the last glacial must be accounted for.  相似文献   

6.
Oxygen isotope stage 3 (OIS 3), encompassing the long middle section of the last glacial interval, has been the focus of an intensive high-resolution climate modeling effort for Europe. These model simulations produce substantially colder climates than modern simulations; however, the temperatures appear warmer than many proxy indicators suggest. In order to evaluate the importance of the model boundary conditions, comparable simulations are completed for the last glacial maximum (LGM). The LGM simulation produces a much colder European continent than OIS 3, despite similarities in the specification of sea-surface temperatures (SSTs). Ice-sheet dimension is evidently a key factor in explaining the difference in European climates over the past 40,000 yr. However, underestimates in specified OIS 3 ice sheets cannot be invoked to explain the discrepancies, since data strongly indicate small ice-sheet extents at that time; this leaves errors in specified OIS 3 SSTs as the most likely cause.  相似文献   

7.
Geomorphic evidence of former glaciation in the high Drakensberg of southern Africa has proven controversial, with conflicting glacial and non‐glacial interpretations suggested for many landforms. This paper presents new geomorphological, sedimentological and micromorphological data, and glacier mass‐balance modelling for a site in the Leqooa Valley, eastern Lesotho, preserving what are considered to be moraines of a former niche glacier that existed during the Last Glacial Maximum (LGM). The geomorphology and macro‐sedimentology of the deposits display characteristics of both active and passive transport by glacial processes. However, micromorphological analyses indicate a more complex history of glacial deposition and subsequent reworking by mass movement processes. The application of a glacier reconstruction technique to determine whether this site could have supported a glacier indicates a reconstructed glacier equilibrium line altitude (ELA) of 3136 m a.s.l. and palaeoglacier mass balance characteristics comparable with modern analogues, reflecting viable, if marginal glaciation. Radiocarbon dates obtained from organic sediment within the moraines indicate that these are of LGM age. The reconstructed palaeoclimatic conditions during the LGM suggest that snow accumulation in the Drakensberg was significantly higher than considered by other studies, and has substantial relevance for tuning regional climate models for southern Africa during the last glacial cycle. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

8.
New 10Be dates for glacial landforms in the Fuentes Carrionas area (Cantabrian Mountains, nothern Spain) are presented. Mapped and dated landforms in Fuentes Carrionas made possible a palaeoglacier reconstruction for four glacial stages. Results were compared to other nearby palaeoenvironmental proxies, so a final approximation on the mean annual temperature and annual precipitation that caused the four glacial advance stages is proposed. Glaciers reached their maximum extension at 36 ka, in a cold and dry environment. A second advance stage took place between 18.5 and 19.5 ka, during the Last Glacial Maximum (LGM), when glaciers advanced in a wet environment, with positive rainfall anomalies. A third glacial advance was dated during the Oldest Dryas, in which climate shifted to extremely cold and dry conditions. Finally, a last stage has been identified and proposed to the Younger Dryas, in which precipitation anomalies are negligible. Our results confirm some of the previously made palaeoglacial and palaeoenvironmental inferences for the Iberian Peninsula, as well as provide valuable and accurate anomalies, which are useful for climate modelling.  相似文献   

9.
Paleoecological (pollen, phytolith, and wood) analyses of sediments, radiocarbon dated 33,000 to 26,000 yr B.P., from two sites in Ecuadorian Amazonia provide data that suggest a cooling of ca. 7.5°C below present in equatorial lowlands from 33,000 to 30,000 yr B.P. A period of warning followed in which novel species assemblages, a blend of montane and lowland floral components, persisted for at least 4000 years. These data of forest community change, from sites lying within the postulated glacial rain forest Napo refugium, provide the strongest paleoecological refutation of the refugial hypothesis yet obtained. The large temperature depression at ca. 30,000 yr B.P. allows the possibility that if maximum cooling at the equator was synchronous with the last glacial maximum (LGM) of the northern hemisphere, freezing temperatures would have been experienced in parts of lowland Amazonia between 25,000 and 18,000 B.P.  相似文献   

10.
《地学前缘(英文版)》2020,11(3):807-820
The fungus Ophiocordyceps sinensis is endemic to the vast region of the Qinghai-Tibetan plateau(QTP).The unique and complex geographical environmental conditions have led to the "sky island" distribution structure of O.sinensis.Due to limited and unbalanced sample collections,the previous data on O.sinensis regarding its genetic diversity and spatial structure have been deemed insufficient.In this study,we analyzed the diversity and phylogeographic structures of O.sinensis using internally transcribed spacer region(ITS) and 5-locus datasets by a large-scale sampling.A total of 111 haplotypes of ITS sequences were identified from 948 samples data of the fungus O.sinensis,with representing high genetic diversity,and 8 phylogenetic clades were recognized in O.sinensis.Both the southeastern Tibet and the northwestern Yunnan were the centers of genetic diversity and genetic differentiation of the fungus,and they were inferred as the glacial refugia in the Quaternary.Three distribution patterns were identified to correspond to the 8 clades,including but not limited to the coexistence of widely and specific local distributive structures.It also revealed that the differentiation pattern of O.sinensis did not fit for the isolation-by-distance model.The differentiation into the 8 clades occurred between 1.56 Myr and6.62 Myr.The ancestor of O.sinensis most likely originated in the late Miocene(6.62 Myr) in the northwestern Yunnan,and the Scene A-C of the Qinghai-Tibetan movements may have played an important role in the differentiation of O.sinensis during the late Miocene-Pliocene periods.Our current results provide a much clearer and detailed understanding of the genetic diversity and geographical spatial distribution of the endemic alpine fungus O.sinensis.It also revealed that the geochronology resulting from paleogeology could be cross-examined with biomolecular clock at a finer scale.  相似文献   

11.
Comparisons of historical and modern assemblages of mammals can yield important insights into patterns and processes of environmental change. Here, we present the first analyses of small mammal assemblages present in northern Patagonia during the last glacial maximum (LGM). Using remains obtained from owl pellets excavated from an archeological cave site (Arroyo Corral I, levels VII–V, carbon dates of 22,400–21,530 cal yr BP), we generate estimates of the minimum number of individuals for all species detected; these estimates, in turn are used to determine relative species abundances. Comparisons of these data with similar analyses of small mammal remains obtained from a second archeological site (ACoII, levels IV–V, carbon dates of 10,010–9220 cal yr BP) as well as from modern owl pellets reveal pronounced changes in relative species abundance since the LGM. In particular, Euneomys chinchilloides and Ctenomys sociabilis – the predominant species during the LGM – declined markedly, suggesting a change from open, bare habitat punctuated by patches of wet meadows and shrubs to the more densely vegetated mosaic of ecotone habitats found in this region today. These data provide important new insights into the environmental changes that have occurred in northern Patagonia over the last 20,000 years.  相似文献   

12.
The Namur area in Belgium is useful to study brown (Ursus arctos) and cave bears (Ursus spelaeus) as the assemblage contains little temporal and no geographical variation. Here, we aim to assess ontogenetic allometry within cave bears, as well as ecomorphological differences between adult brown bears (n = 9), adult cave bears (n = 5) and juvenile cave bears (n = 3). Landmarks for 3D digitization of the mandible were chosen based on the taphonomical damage of the specimens. Extant brown bears and extinct Pleistocene brown and cave bears were digitized with a Microscribe G2. Generalized Procrustes superimposition was performed on the coordinates. Allometry was studied using regression analysis. Principal component analysis (PCA) was conducted to assess ecomorphological differences between the groups. 61% of the shape variance within juvenile and adult cave bears was predicted by size (n = 8, p < 0.01). The juvenile cave bears have relatively deep horizontal rami. In adult cave bears, the horizontal ramus is much narrower dorsoventrally. Juvenile cave bears have a small masseteric fossa and a short coronoid process, whereas both are larger, relative to mandible size, in adult cave bears. This made juvenile cave bears likely less effective masticators than fully grown cave bears. In the PCA, principal component (PC) 1 accounts for 45.0% of the total variance and PC2 accounts for 27.6%. Fossil U. arctos from Namur fall within the 95% confidence interval of modern North American U. arctos on both PCs, but are more similar to cave bears than the average extant brown bear. From the similarity of fossil and modern brown bears, it can be deduced that the diet of fossil brown bears was probably also within the range of their modern North American conspecifics, although they might have been more efficient at masticating plant matter.  相似文献   

13.
Equilibrium-line-altitude (ELA) reconstructions using the toe-to-headwall-altitude ratio method for paleoglaciers in the Cordilleras Blanca and Oriental, northern Peruvian Andes (7–10°S; 77°20'–77°35'W), indicate that ELAs during the last glacial maximum (LGM; marine isotope stage 2) were c . 4300 m in the Cordillera Blanca, c . 3900–3600 m on the west side of the Cordillera Oriental, and c . 3200 m on the east (Amazon Basin) side of the Cordillera Oriental. Comparison with estimated modern ELAs and glaciation thresholds indicate that ELA depression ranged from c . 700 m in the Cordillera Blanca to c . 1200 m on the east side of the Cordillera Oriental. This augments data from many mountain ranges in middle- and low-latitude regions that indicate that ELAs during the LGM were depressed by c . 1000 m. Published palynological evidence for drier conditions during the LGM in the tropical Andes suggests that ELA depression of this amount involved a temperature reduction (> 5–6°C) that greatly exceeded the tropical sea-surface temperature depression estimates of CLIMAP (< 2°C). The west to east increase in ELA depression during the LGM indicates that the steep modern precipitation gradients may have been even steeper during the LGM.  相似文献   

14.
In recent years it has been demonstrated that the formation of long-timescale river terrace sequences, which are generally found in areas beyond the extent of most if not all of the Middle and Late Pleistocene ice sheets, has invariably been a response to uplift during the Late Cenozoic and especially the Quaternary. Climatic fluctuation at a Milankovitch timescale has driven the alternations of aggradation and incision recorded in such terraces. It has been widely observed, however, that fluvial terraces also occur in areas glaciated during the Last Glacial Maximum (LGM), which coincides with marine oxygen isotope stage (MIS) 2. This paper, in seeking to compare records from inside and outside of the LGM ice limit, concentrates on a single English river system, that of the Humber. The Humber estuary is shared by the largely Pennine-derived drainage of the Yorkshire Ouse, to the north and entirely within the MIS 2 glacial limit, and, to the south, the Trent, which is almost wholly outside the LGM limit. Thus the Trent has a terrace sequence extending back to the Middle Pleistocene, whereas in the component rivers of the Ouse system, records begin with the melting of the last glacial ice. Importantly, there is considerable difference in the disposition of the post-LGM fluvial deposits in these two subsystems. In the Ouse system there are modest terrace staircases, commencing with full glacial deposits that stand up to 30 m above the modern floodplain. In the Trent, in contrast, last glacial gravels form the foundation of the modern floodplain, with Holocene sediments emplaced directly above them. Thus there is little or no post-LGM incision in the Trent, whereas in the Ouse several incision events are recorded, continuing into the later stages of the Holocene. Wider comparison reveals that the Ouse system is an exemplar for other sequences within the MIS 2 limit, whereas systems beyond this glaciation typically have last glacial sediments beneath their modern floodplains and show little evidence of Holocene incision. The various possible explanations of these differences are discussed, with emphasis placed on glacio-isostatic uplift of areas glaciated during MIS 2 as the main reason for the significant post-glacial incision that typifies valleys in such regions. A new approach to modelling glacio-isostatic adjustment is outlined, from which it is concluded that lower-crustal flow plays a significant role in this process in regions of relatively hot and dynamic crust, like northern England, in addition to the mantle flow that is considered in conventional analyses of glacio-isostasy. Lower-crustal flow has a significant effect due to the combination of the small spatial scale of the glaciated region of northern England and the high mobility of the lower-crustal layer beneath it, due to the heating effect of the widespread Palaeozoic granite in the area.  相似文献   

15.
This study presents new ages for the northwest section of the Laurentide Ice Sheet (LIS) glacial chronology from material recovered from two retrogressive thaw slumps exposed in the Richardson Mountains, Northwest Territories, Canada. One study site, located at the maximum glacial limit of the LIS in the Richardson Mountains, had calcite concretions recovered from aufeis buried by glacial till that were dated by U/Th disequilibrium to 18,500 cal yr BP. The second site, located on the Peel Plateau to the east yielded a fossil horse (Equus) mandible that was radiocarbon dated to ca. 19,700 cal yr BP. These ages indicate that the Peel Plateau on the eastern flanks of the Richardson Mountains was glaciated only after 18,500 cal yr BP, which is later than previous models for the global last glacial maximum (LGM). As the LIS retreated the Peel Plateau around 15,000 cal yr BP, following the age of the Tutsieta phase, we conclude that the presence of the northwestern margin of the LIS at its maximum limit was a very short event in the western Canadian Arctic.  相似文献   

16.
Although glacial landscapes have previously been used for the reconstruction of late Quaternary glaciations in the Central Andes, only few data exist for the Eastern Cordillera in Bolivia. Here, we present results from detailed morphostratigraphic mapping and new data of surface exposure dating (SED), optically stimulated luminescence (OSL), and radiocarbon dating (14C) from the Huara Loma Valley, Cordillera de Cochabamba (Bolivia). Discrepancies between individual dating methods could be addressed within the context of a solid geomorphic framework. We identified two major glaciations. The older is not well constrained by the available data, whereas the younger glaciation is subdivided into at least four major glacial stages. Regarding the latter, a first advance dated to ~ 29-25 ka occurred roughly contemporaneous with the onset of the global last glacial maximum (LGM) and was followed by a less extensive (re-)advance around 20-18 ka. The local last glacial maximum (LLGM) in the Huara Loma Valley took place during the humid lateglacial ~ 17-16 ka, followed by several smaller readvances until ~ 10-11 ka, and complete deglaciation at the end of the Early Holocene.  相似文献   

17.
We present a well‐dated, high‐resolution and continuous sediment record spanning the last c. 24 000 years from lake Bolshoye Shchuchye located in the Polar Ural Mountains, Arctic Russia. This is the first continuous sediment succession reaching back into the Last Glacial Maximum (LGM) ever retrieved from this region. We reconstruct the glacial and climate history in the area since the LGM based on sedimentological and geochemical analysis of a 24‐m‐long sediment core. A robust chronology was established using a combination of AMS 14C‐dating, the position of the Vedde Ash and varve counting. The varved part of the sediment core spans across the LGM from 24 to 18.7 cal. ka BP. We conclude that the lake basin remained ice‐free throughout the LGM, but that mountain glaciers were present in the lake catchment. A decrease in both glacial varve preservation and sedimentation rate suggests that these glaciers started to retreat c. 18.7 cal. ka BP and had disappeared from the catchment by 14.35 cal. ka BP. There are no indications of glacier regrowth during the Younger Dryas. We infer a distinct climatic amelioration following the onset of the Holocene and an Early to Middle Holocene thermal optimum between 10–5 cal. ka BP. Our results provide a long‐awaited continuous and high‐resolution record of past climate that supplements the existing, more fragmentary data from moraines and exposed strata along river banks and coastal cliffs around the Russian Arctic.  相似文献   

18.
Using data from glacial geomorphology, tephra–soil stratigraphy and mineralogy, palynology, and radiocarbon dating, a sequence of glacial and bioclimatic stades and interstades has been identified for the last ca. 50000 yr in the Ruiz-Tolima massif, Cordillera Central, Colombia. Six Pleistocene cold stades separated by warmer interstades occurred: before 48000, between 48000 and 33000, between 28000 and 21000, from ≥16000 to ca. 14000, ca. 13000–12400, and ca. 11000–10000 yr BP. Although these radiocarbon ages are minimum-limiting ages obtained from tephra layers on top of tills, the tills are not significantly older because most are bracketed by dated tephra sets in measured stratigraphic sections. Two minor moraine stages likely reflect glacier standstill during cold intervals ca. 7400 yr BP and slightly earlier. Finally, glaciers readvanced between the seventeenth and nineteenth centuries. In contrast to the ice-clad volcanoes of the massif, ca. 34 km2 in area above an altitude of ca. 4800 m, the ice cover expanded to 1200 km2 during the Last Glacial Maximum (LGM) and was still 800 km2 during Late-glacial time (LGT). Glacier reconstructions based on the moraines suggest depression of the equilibrium line altitude (ELA) by ca. 1100 m during the LGM and 500–600 m during LGT relative to the modern ELA, which lies at ca. 5100 m in the Cordillera Central. Glaciers in this region apparently reached their greatest extent when the climate was cold and wet, e.g. during stades corresponding to Oxygen Isotope Stage 3; glaciers were still expanding during the LGM ca. 28000–21000 yr BP, but they shrank considerably after 21000 yr BP because of greatly reduced precipitation. © 1997 John Wiley & Sons, Ltd.  相似文献   

19.
Cave bears (Ursus spelaeus) are an iconic component of the European late Quaternary Ice Age megafauna. Recent demographic analyses based on cave bear mtDNA sequences and refined radiocarbon dating indicate that cave bear population size and genetic diversity started to decline some 50 kilo years ago (kya). Hence, neither the coldest phase of the last glaciation (started some 24 kya), nor the colonization of Europe by Palaeolithic hunters (started some 45 kya) coincides with the beginning of population decline. Here, we reconstructed cave bear climatic niche evolution through time. Then, we performed spatially explicit population viability analyses to assess cave bear demographics through time in response to climatic changes, human effects on bear survival and their combination. We found that climate change was responsible for a 10‐fold decrease in cave bear population size after 40 kya. However, climate change on its own could not explain U. spelaeus extinction at 24 kya. Additional negative effects consistent with human population expansion are required to explain both U. spelaeus' retreat from eastern Europe since 40 kya and its final extinction.  相似文献   

20.
Identifying differential population structure within metacommunities is key toward describing the mechanisms that maintain biodiversity in natural systems. At both local and regional scales on the North American Atlantic coast, we assessed phylogeographic and genetic diversity patterns of six common salt marsh invertebrates using equivalent sampling schemes and sequence data from the same mitochondrial locus. In general, our results suggest little genetic structure across four previously sampled biogeographic regions and a slight increase in genetic diversity from northern to southern areas; however, two of the species (Geukensia demissa and Uca pugilator) exhibited significant differentiation between the northernmost populations and other regions, consistent with a number of previous studies. Although the minimal genetic structure recovered in this community is consistent with expectations based on the larval life history of the species examined, confirmation of this result suggests that latitudinal shifts in ecological interactions in salt marsh systems are environmentally driven, rather than due to heritable adaptation.  相似文献   

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