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1.
The dietary regimes of 15 ungulate species from the middle Pleistocene levels of the hominid-bearing locality of Elandsfontein, South Africa, are investigated using the mesowear technique. Previous studies, using taxonomic analogy, classified twelve of the studied species as grazers (Redunca arundinum, Hippotragus gigas, Hippotragus leucophaeus, Antidorcas recki, Homoiceras antiquus, Damaliscus aff. lunatus, Connochaetes gnou laticornutus, Rabaticerus arambourgi, Damaliscus niro, Damaliscus sp. nov., an unnamed “spiral horn” antelope and Equus capensis), one as a mixed feeder (Taurotragus oryx) and two as browsers (Tragelaphus strepsiceros and Raphicerus melanotis). Although results from mesowear analysis sustain previous dietary classifications in the majority of cases, five species were reclassified. Three species previously classified as grazers, were reclassified as mixed feeders (H. gigas, D. aff. lunatus and R. arambourgi), one previously classified as a grazer, was reclassified as a browser (the “spiral horn” antelope), and one previously classified as a mixed feeder, was reclassified as a browser (T. oryx). While current results broadly support previous reconstructions of the Elandsfontein middle Pleistocene environment as one which included a substantial C3 grassy component, the reclassifications suggest that trees, broad-leaved bush and fynbos were probably more prominent than what was previously thought.  相似文献   

2.
During the late Pleistocene and early Holocene, Bison was widely dispersed across North America and occupied most regions not covered by ice sheets. A dietary study on Bison paleopopulations from Alaska, New Mexico, Florida, and Texas was performed using two methods that relate dental wear patterns to diet, mesowear analysis and microwear analysis. These data were compared to a mixed sample of extant Bison from the North American central plains, extant wood Bison from Alberta (Canada) and a variety of other modern ungulates. Mesowear relates macroscopic molar facet shape to levels of dietary abrasion. The mesowear signature observed on fossil Bison differs significantly from the hyper-abrasive grazing diet of extant Bison. Tooth microwear examines wear on the surface of enamel at a microscopic scale. The microwear signal of fossil samples resembles to modern Bison, but the fossil samples show a greater diversity of features, suggesting that fossil Bison populations regularly consumed food items that are texturally inconsistent with the short-grass diet typical of modern plains Bison. Mesowear and microwear signals of fossil Bison samples most closely resemble a variety of typical mixed feeding ungulates, all with diets that are substantially less abrasive than what is typical for modern plains Bison. Furthermore, statistical tests suggest significant differences between the microwear signatures of the fossil samples, thus revealing geographic variability in Pleistocene Bison diets. This study reveals that fossils are of value in developing an understanding of the dietary breadth and ecological versatility of species that, in recent times, are rare, endangered, and occupy only a small remnant of their former ranges.  相似文献   

3.
The Cape grysbok is endemic to southern Africa's Cape Floral Region where it selectively browses various species of dicotyledonous vegetation. Fossil evidence indicates that the grysbok persisted under glacial and interglacial conditions throughout the late Quaternary and inhabited a range of environments. This study employs mesowear analysis to reconstruct grysbok diets over time and in response to changing environments at Nelson Bay Cave, Die Kelders Cave 1, Klasies River Mouth, and Swartklip 1. Results indicate that the amount of grasses (monocots) versus leafy vegetation (dicots) included in the diet fluctuated over time and largely in agreement with changes in faunal community structure. The case for dietary flexibility is particularly clear at Nelson Bay Cave, where there is a significant trend from mixed feeding towards increased browsing from the late Pleistocene (~ 18,500 14C yr BP) through the Holocene. Dietary shifts at Nelson Bay Cave are consistent with the hypothesis that declining grassland productivity is responsible for the terminal Pleistocene extinction of several large ungulates in southern Africa. Furthermore, the short-term dietary shifts demonstrated here (100s to 1000s of years) provide an important caution against relying on taxonomic uniformitarianism when reconstructing the dietary preferences of fossil ungulates, both extant and extinct.  相似文献   

4.
The geographical distribution of species affects their dietary traits relative to resources available in different latitudes. Dietary traits of Rangifer tarandus, a species with a wide geographical distribution, were investigated using tooth mesowear and microwear methods in eight extant populations from Canada. The data show a latitudinal shift corresponding to a vegetational gradient from the taiga to the tundra, i.e. an increase of lichen consumption from the low to the high latitudes. This pattern is also evidenced in the Pleistocene fossil record of Europe where R. tarandus populations from low latitude localities show a lower consumption of lichen than at higher latitudes.  相似文献   

5.
The paleodiet of Eucladoceros ctenoides (Nesti 1841), has been investigated on two sub-species from the Villafranchian fossiliferous sites of Saint-Vallier (Drôme, France), E. ctenoides vireti Heintz 1970, and Senèze (Haute-Loire, France), E. ctenoides senezensis Depéret 1910, using skull and mandible morphological features as well as the meso- and microwears methods. The validity of skull and mandible characters used to infer diet has been tested on extant cervid species: browsers (roe deer, moose and white-tailed deer) and mixed feeders at different level (Père David’s deer, fallow deer, chital, sambar, red deer and elk). The obtained results show that the Eucladoceros specimens from Saint-Vallier and Senèze shared paleodiet and ecologic niche. Cranial morphology and tooth wear are more similar to those characterising browsers than mixed feeders. Cranial morphology can be considered as a useful tool in inferring palaeodiets, in particular when tooth samples are too scanty to apply of micro- and/or mesowear analytic methods.
Manuscrit reçu le 17 septembre 2003 Révision acceptée le 16 février 16, 2005  相似文献   

6.
7.
New Mesozoic Mesopsychidae(Mecoptera)from Northeastern China   总被引:2,自引:2,他引:0  
<正>The Mesozoic family Mesopsychidae Tillyard,1917 presently consists of seven described genera and 13 species from the mid-Triassic to the Early Cretaceous of Australia,South Africa and Eurasia.In the present paper one new genus and three new species of fossil mesopsychids are described that add significant distributional and stratigraphic extensions to the family.This finding documents the first formal record of fossil Mesopsychidae in China.Both Lichnomesopsyche gloriae gen.et sp.nov.and L.daohugouensis gen.et sp.nov. were found from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,whereas Vitimopsyche kozlovi sp. nov.,of mid Early Cretaceous age,was collected from the Jehol Biota of northern Hebei Province.This new,well-preserved material from China reveals complete and previously unknown body features,including head, antennae,mouthparts,legs and abdomen.The delicate and long proboscides of these new taxa indicate that they were feeding on externally exposed,nutrient-rich fluids of gymnospermous ovulate fructifications,and incapable of piercing surface epidermis-attributable principally to the absence of stylets.These proboscides originated, perhaps multiply,among basal Mecoptera and are functionally and structurally convergent with equivalent mouthparts borne by fossil and extant Diptera,Lepidoptera,Neuroptera and Coleoptera.  相似文献   

8.
The so-called Hundsheim rhinoceros, Stephanorhinus hundsheimensis, was a very common faunal element of the Early to early Middle Pleistocene period in the western Palaearctic. In this study, individuals from two different central European populations of the Hundsheim rhinoceros were analysed in order to determine whether their local dietary signals could reflect differing food availability between the two populations, and whether such information could provide a better understanding of the ecological role of S. hundsheimensis within corresponding faunal assemblages, and of its principal subsistence strategy in the western Palaearctic. The mesowear traits observed in the studied S. hundsheimensis populations have been interpreted as representing biome-specific signals, indicating grassland vegetation at the site of Süßenborn, and dense to open forests at Voigtstedt (both localities in Germany). The analyses performed on the fossil rhino material demonstrate the most pronounced dietary variability ever established for a single herbivorous ungulate species by mesowear studies. This variability ranges from an attrition dominated grazing regime, to a one of predominantly browsing, and characterises S. hundsheimensis as the most ecologically tolerant rhinoceros of the Palaearctic Plio-Pleistocene. Although such dietary flexibility proved an effective enough subsistence strategy over a period of 600–900 ka (1.4/1.2–0.6/0.5 Myr) in the western Palaearctic, the situation changed dramatically after 0.6 Myr BP, when the new species of rhinoceroses, Stephanorhinus hemitoechus and Stephanorhinus kirchbergensis, appeared and started to compete for both the grass and the browse. For the generalist S. hundsheimensis, this bilateral interference was detrimental to its success in all of its habitats. The successful competition of specialised forms of rhinoceroses, which might have originated as a result of the development of 100 ka periodicity in the global climatic record, is proposed as the main reason for the extinction of S. hundsheimensis during the early Middle Pleistocene.  相似文献   

9.
Palaeolithic people commonly hunted Equus hydruntinus, an extinct species of equid whose cursorial body proportions suggest an adaptation to semi-arid conditions. Despite the frequency with which it is encountered in fossil deposits, only partial cranial remains have been reported until now. As a result, the systematic affiliation of the species remains a subject of controversy. Two nearly complete E. hydruntinus crania are presented here for the first time. These skulls show that E. hydruntinus is a distinct species, more closely related to the hemiones (Asiatic asses) than to any other equid. This suggests that the social organisation of E. hydruntinus followed one of two known equid sociotypes: resource defense territoriality.  相似文献   

10.
A partial skeleton of straight-tusked elephant (Palaeoloxodon antiquus) excavated at Selsey beach (Sussex, UK) in 1961 is one of very few from Marine Isotope Stage 7 (MIS 7), ca. 240–200 ka, in Europe. Dental morphometrics, including X-ray micro-computed tomography, confirms the species identification. Age determination through dental wear and skeletal morphology, sex determination through pelvic anatomy, and body-size measurements, together identify the individual as a female of approximately 30 years old with a live shoulder height of ca. 3 m and a body mass of ca. 5 t. Dental mesowear indicates mixed feeding with a significant proportion of grass, indicating selective feeding in a predominately forested environment. Although taphonomic conclusions are limited by the nature of the excavation, identification of 107 recovered skeletal elements indicates predominant preservation of the right side of the animal, suggesting death or burial on the right side in a shallow, slow-moving freshwater environment, with possible removal of other elements by scavenging or decay. Combining the Selsey data with 29 comparative skeletons from across Europe, we demonstrate the validity of pelvic proportions for determining sex and provide a graphical representation of variation and growth rate in males and females of this species.  相似文献   

11.
The 121 local faunas of large mammals from Late Pleistocene sites of the South (56–51°N), Middle (59–56°N) and North (64–59°N) Urals have been studied. All local faunas were combined into eight chronological groups on the basis of radiocarbon dates and the evolutionary level of rodents present in them. On the basis of species composition analysis of the faunas, three chronological complexes have been distinguished: Mikulino, Early–Middle Valdai and Late Valdai. The first is characterized by the presence of Hystrix vinogradovi and Ursus thibetanus; the second, by the presence of a large form of horse (Equus (E.) cf. latipes), Crocuta crocuta, Ursus spelaeus and U. savini; the third, by the presence of a small horse (E. uralensis) and absence of U. spelaeus, U. savini and C. crocuta. The latter two complexes were represented by three geographical variants: southern (South Urals), northern (North Urals) and transitional (Middle Urals). Differences between theriocomplexes are related to changes in morphology and areas and extinctions of a series of species. The existence of chronological theriocomplexes and their geographical variants was determined by chronological and geographical change in structure of paleophytocoenoses. It should be noted that the role of human in changes of chronological complexes and species extinctions in the Late Pleistocene has not been demonstrated in the Urals. In the Urals U. savini probably became extinct at the end of the Middle Valdai, C. crocuta at the beginning of the last glacial maximum (LGM), U. spelaeus at the end of the LGM, Coelodonta antiquitatis at the beginning of the Preboreal and Megaloceros giganteus at the middle of the Atlantic.  相似文献   

12.
Mesowear and microwear on enamel from 763 teeth of middle and late Pleistocene ungulates were analysed to infer the potential of dental wear analysis of faunal remains as a paleoenvironmental and paleoclimatic proxy in relation to climatic changes and diversity of vegetation available in the environment. Fossil localities including levels belonging to two glacial and two interglacial stages were selected in Germany, France, and Spain. At a temporal scale, results indicate that the dietary diversity in ungulates is higher during interglacial phases (MIS 5 and 3) than during pleniglacial phases (MIS 8 and 4). Dietary diversity is concluded to be related to climate-driven vegetation changes which during interglacials lead to increased variety of potential food items available to ungulates. At the geographical scale, during interglacials, changes in diet composition are evident along geographical gradients. The corresponding dietary gradients are proposed to be related to climate and vegetation gradients reflecting more arid climates in the Mediterranean area compared to North-Western Europe. Species consistently represented at all localities investigated are Cervus elaphus (Cervidae, Artiodactyla) and Equus ferus (Equidae, Perissodactyla). C. elaphus populations are found to consistently have less abrasive diets than E. ferus populations but dietary traits of both species varied largely, revealing a significant plasticity in the feeding adaptation of both species. Those traits are concluded to be related to differences in vegetation structure at each locality and complement the evidence that ungulates have broader dietary habits than what is usually assumed.  相似文献   

13.
The Mission Paléoanthropologique Franco-Tchadienne (MPFT) found a new species of Orycteropodidae (Mammalia, Tubulidentata) in the Kollé fossiliferous sector, northern Chad. After Orycteropus abundulafus [Journal of Vertebrate Paleontology 20 (1) (2000) 205–209; Lehmann, T., Vignaud, P., Likius A., Brunet M., in press. A new Orycteropodidae (Mammalia, Tubulidentata) in the Mio-Pliocene of Northern Chad. Zool. J. Linnean Soc.], this specimen is the second complete skeleton of fossil aardvark found in the Djurab desert. It is the first complete representative of an Orycteropus species found in the Pliocene of Africa. In regard to the Miocene fossil aardvarks, this new taxon, Orycteropus djourabensis nov. sp., shows more affinities with the extant O. afer. The main differences are the larger teeth and the shorter hand in the fossil form. Kossom Bougoudi and Kollé represent a chronological series that gives a unique opportunity for studying the evolution of the African Tubulidentata around the Mio-Pliocene boundary (5.5-4 My). The new species is distinct from the older Chadian Orycteropodid from KB and it embodies the taxonomic turnover that took place within the order Tubulidentata around this boundary in Africa. Moreover, this new species is the oldest known Orycteropus species that clearly belongs to the modern forms including the extant aardvark.  相似文献   

14.
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia.  相似文献   

15.

Geological mapping of fault systems on the Gazelle Peninsula, eastern New Britain arc, combined with a reinterpretation of existing sea floor data indicate that faults previously thought to be a possible location of the boundary between the North and South Bismarck Plates, do not appear to be directly related to the plate boundary spreading centres and transform faults in the 3.5 Ma Manus Basin. Structure on the Gazelle Peninsula is dominated by the Mediva Fault (new name) and the Wide Bay Fault System, both north‐northwest trending, deep‐seated features. The Mediva Fault, an element of the Baining Mountain Horst and Graben Zone, is an extensional structure which has focused Middle Miocene intrusive activity, controlled Mio‐Pliocene sedimentation in the central Gazelle Peninsula, and displaced Quaternary volcanic deposits. The Wide Bay Fault System has been active since at least the Late Oligocene. One hundred kilometres of sinistral strike‐slip motion is likely on this fault since at least the late Middle Miocene, moving the Gazelle Peninsula in a north‐northwest direction with respect to the remainder of New Britain. The nature and timing of movements along these two major structures indicate that some other major tectonic process has operated (and presently continues) in this region of the New Britain arc to create these structures.  相似文献   

16.
《Quaternary Science Reviews》2007,26(3-4):415-435
The Orce fossil quarries, in the Baza Basin of southeastern Spain, are a rich source of Early Pleistocene Palaeolithic tools and vertebrate remains. Geologic fieldwork during the last decade has placed these fossiliferous strata within the context of a thick Neogene continental sequence. Detailed lithostratigraphic and magnetostratigraphic results indicate that at least the upper 60 m of this sequence are of Early Pleistocene age. The quarried strata (Venta Micena, Barranco León and Fuentenueva-3) are from a narrow time span (<100 ka) starting before 1.3 Ma. A new, lower excavation at ∼1.5 Ma (Fuentenueva-1 quarry) has a distinctly older fauna. These Orce strata provide a high resolution, Early Pleistocene record of grassland fauna that shows the end of Mammal Neogene fauna (MN17) in the Fuentenueva-1 site (with Gazella borbonica, Equus stenonis) and the beginning of more characteristic Pleistocene fauna in the Venta Micena site (with Hippopotamus antiquus, Equus granatensis, Homo sp.). Thus far, no evidence for human occupation has been found within the earlier Fuentenueva-1 quarry, although many of the same terminal MN17 species have been found with hominids on the other side of Europe at the Dmanisi site (earliest Pleistocene) in the Republic of Georgia.  相似文献   

17.
This paper addresses the use of fossil species of the horse Equus as stratigraphic markers in the Pliocene and Pleistocene. Current research shows that the species recently so used do not succeed one another in orderly fashion, but occur and co-occur at ages other than presupposed by the stratigraphic schemes (usually earlier). Horse species should be identified on their morphology, not on their stratigraphic age. ‘Subspecies’ of fossil horses do not conform to subspecies in zoology, as they lack a geographic range. They thus lack taxonomic status and stratigraphic significance and should be avoided.  相似文献   

18.
19.
Palaeolithic people commonly hunted Equus hydruntinus, an extinct species of equid whose cursorial body proportions suggest an adaptation to semi-arid conditions. Despite the frequency with which it is encountered in fossil deposits, only partial cranial remains have been reported until now. As a result, the systematic affiliation of the species remains a subject of controversy. Two nearly complete E. hydruntinus crania are presented here for the first time. These skulls show that E. hydruntinus is a distinct species, more closely related to the hemiones (Asiatic asses) than to any other equid. This suggests that the social organisation of E. hydruntinus followed one of two known equid sociotypes: resource defense territoriality.  相似文献   

20.
We report the first record of anilioids from southern South America, a region where they do not live today. The fossils come from the Allen Formation (Late Campanian–Early Maastrichtian) at Bajo Trapalcó and Bajo de Santa Rosa localities, Río Negro province, Argentina. The remains consist of several vertebrae, most of which belong to the mid–posterior precloacal region of the column. Comparisons to other extant and extinct anilioid taxa indicate that these remains represent a new taxon, Australophis anilioides gen. et sp. nov. Australophis is morphologically closer to Palaeocene Hoffstetterella from Brazil and extant South American Anilius than to any other snake. These taxa might be closely related, in which case they would represent a lineage distinct from that including Cylindrophis and uropeltids. The two lineages must have diverged by the Late Cretaceous.  相似文献   

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