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1.
New species of a gecko of the genus Euleptes is described here—E. klembarai. The material comes from the middle Miocene (Astaracian, MN 6) of Slovakia, more precisely from the well-known locality called Zapfe`s fissure fillings (Devínska Nová Ves, Bratislava). The fossil material consists of isolated left maxilla, right dentary, right pterygoid and cervical and dorsal vertebrae. The currently known fossil record suggests that isolation of environment of the Zapfe`s fissure site, created a refugium for the genus Euleptes in Central Europe (today, this taxon still inhabits southern part of Europe and North Africa—E. europea), probably resulting from the island geography of this area during the middle Miocene. The isolation of this territory might have facilitated allopatric speciation.  相似文献   

2.
Fossil anguine lizard specimens from several Turkish localities are described in this paper. The material comes from ten different localities, spanning a large geographic area consisting of both parts of the European Turkey and Anatolia, and ranging in age from the Oligocene to the Late Miocene. In certain cases, the generic determination was possible and, accordingly, members of Ophisaurus and Anguis were identified and described in detail. The specimens of Anguis, found in different, Middle and Late Miocene localities from Anatolia, represent two of only a few fossil occurrences of this taxon. Moreover, the material reported herein represents the oldest occurrences of anguine lizards, not only from Turkey, but from southeastern Europe and the Eastern Mediterranean basin as a whole. These rare records provide important information about the dispersal routes of anguines from Europe to Asia and significantly enhance our understanding of their biogeography.  相似文献   

3.
The first unequivocal remains of medium to large-sized mustelids from the middle Miocene Halamagai and Kekemaideng formations have been found in the Ulungur River area, Xinjiang, northwestern China. These new fossils are referred to the hypercarnivorous mustelid Hoplictis Ginsburg, 1961 and denote the first record of the genus in East Asia.We define Hoplictis baihu n. sp., for the mustelid from Tieersihabahe(Halamagai Fm.), which represents the smallest species of the genus. This primitive form is closer to H. florancei and H. noueli than to H. anatolicus and later, larger and more derived Hoplictis spp., from Europe and North America. A large toothless mandible from Duolebulejin(Kekemaiden Fm.) is assigned to Hoplictis cf. helbingi, and it presumably might represent the first record of H. helbingi outside Western Europe. The systematic position of Hoplicitis in relation to Ischyrictis with which it is similar is clarified. The occurrence in East Asia of two species of Hoplictis greatly expands the known distribution and diversity of the genus, and supports a Palaearctic Neogene dispersal event of carnivorans between Europe and Asia during the late Shanwangian–early Tunggurian equivalent to MN5–6 in Europe, and indication of another dispersal event from Europe to North America,through Northwest China during the late Tunggurian, equivalent to MN7–8 in Europe.  相似文献   

4.
本文记述了采自甘肃广河石那奴中中新世地层的哺乳动物化石6目10科12属16种类(以小化石为主),其中新种两个:Alloptox guangheensis sp. nov., Pseudaelurus guangheensis sp. nov.。通过对动物群性质的研究及与其它地区的详细对比,认为下化石层的时代应为中中新世早期,上化石层应为中中新世晚期。二者分别与山旺期、通古尔期大体相当。 广河地区中中新世早期车头沟组和中中新世晚期咸水河组的建立,表示该地区存在着中中新世两个时期的不同环境沉积。  相似文献   

5.
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia.  相似文献   

6.
《Cretaceous Research》2012,33(6):781-785
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia.  相似文献   

7.
The Agenian is the earliest Neogene European Land Mammal Age. It encompasses the mammalian zones MN1 (23.03–22.7 Ma) and MN2 (22.7–20.0 Ma) and roughly coincides with the Aquitanian standard age. Agenian mammalian assemblages from Western Europe encompass a mixture of rhinocerotid taxa of Oligocene affinities and of Miocene newcomers, mostly recorded in France, Germany, Switzerland, and to a lesser extent, Spain. Rhinocerotidae are documented by seven species referred to five genera (Pleuroceros pleuroceros, Protaceratherium minutum, Plesiaceratherium aquitanicum, Mesaceratherium paulhiacense, Diaceratherium lemanense, D. asphaltense, and D. aginense), further attesting to a low suprageneric diversity. Their systematics, morphology, ecology, stratigraphical and geographical ranges are detailed in the present article. Occurrences and geographical ranges of all seven rhinocerotid species are illustrated on palaeogeographical maps of the circum-Mediterranean region at 23 Ma (MN1) and 21 Ma (MN2). The richest Agenian localities (Paulhiac, MN1; Laugnac, MN2) record a specific diversity similar to that of Orleanian rhinocerotid assemblages, with up to five/six associated species. All Agenian rhinocerotid species from Western Europe are endemic to the concerned region, which is consistent with the complete geographic isolation of Western Europe by earliest Miocene times. However, all five genera are documented by twin species in coeval localities of South and Central Asia, which implies (1) vicariant speciation events by latest Oligocene times and (2) the existence of intermittent pathways for terrestrial megamammals such as rhinocerotids during the concerned interval.  相似文献   

8.
《Geodinamica Acta》2013,26(4):151-165
Two magnetostratigraphic profiles (450 samples) have been carried out to constrain the age of synorogenic formations in the southern foreland of the High Atlas of Morocco. The Amekchoud profile covers the Aït Ouglif and Aït Kandoula alluvial formations that form the bulk of the Ouarzazate basin fill, indicating an age between the upper Langhian and the Messinian (Miocene). Data obtained in the previously unexplored Hadida formation profile covers the oldest terms of the foreland basin succession, but the low quality of the magnetic record only allows proposing a tentative age between the middle Lutetian and an undetermined middle to late Eocene. The correlation of the Amekchoud profile is based on the recognition of the long C5n chron (Tortonian) in the middle part of the section studied and a new vertebrate locality of upper Tortonian age found in the upper part. These results indicate a discontinuous record of foreland basin development in the southern Atlas domain from mid Eocene to late Miocene times, punctuated by an intermediate large hiatus of 20-25 ma (late Eocene to mid Miocene). Thrusting in the Sub-Atlas frontal thrust belt began before the Aït Ouglif and Kandoula formations, probably during the Oligocene, and extends up to recent times. The alternation of periods of deposition with others of no sedimentary record, which does not coincide with specific tectonic events, results probably from the interference of orogenic deformation and the mantle-related thermal uplift events that have been described for the Moroccan Atlas.  相似文献   

9.
A fossil-bearing locality near Padhri village, Dhok Pathan, 55 km away from the tehsil Dina, Jhelum District, in the Potwar Plateau, Middle Siwaliks, Punjab, northern Pakistan, is significantly rich in mammalian fossils. This site has provided an abundant mammalian fossil fauna of Late Miocene age from the Dhok Pathan Formation (Fm.). The recovered material belongs to four families: Equidae (horses), Rhinocerotidae (rhinos), Bovidae (cows), and Suidae (pigs). We discovered a new skull of hipparionine Hipparion theobaldi from this locality along with 22 specimens from the associated assemblage of fossil mammals. The recovered material includes seven other species: the aceratheriine Chilotherium intermedium, boselaphines Tragoportax punjabicus, Selenoportax vexillarius, Pachyportax latidens, the antelope Gazella lydekkeri and suinine Propotamochoerus hysudricus. The specimens are isolated teeth, fragments of maxilla, mandibles and horn cores. The Dhok Pathan Fm. is generally composed of claystone, siltstone and sandstone beds and, based on the mammalian fauna, the Padhri fossil locality is dated as Late Miocene. Thi99s formation was deposited in a subtropical paleoenvironment and the predominance of fossil bovids indicates extremely moist conditions with small but frequent standing water bodies.  相似文献   

10.
The biostratigraphy and diversity patterns of terrestrial, hoofed mammals help to understand the transition between the Palaeogene and the Neogene in Western Europe. Three phases are highlighted: (1) the beginning of the Arvernian (Late Oligocene, MP25-27) was characterised by a “stable” faunal composition including the last occurrences of taxa inherited from the Grande Coupure and of newly emerged ones; (2) the latest Arvernian (Late Oligocene, MP28-30) and the Agenian (Early Miocene, MN1-2) saw gradual immigrations leading to progressive replacement of the Arvernian, hoofed mammals towards the establishment of the “classical” Agenian fauna; (3) the beginning of the Orleanian (Early Miocene, MN3-4) coincided with the African-Eurasian faunal interchanges of the Proboscidean Datum Events and led to complete renewal of the Agenian taxa and total disappearance of the last Oligocene survivors. Faunal balances, poly-cohorts and particularly cluster analyses emphasise these three periods and define a temporally well-framed Oligocene–Miocene transition between MP28 and MN2. This transition started in MP28 with a major immigration event, linked to the arrival in Europe of new ungulate taxa, notably a stem group of “Eupecora” and the small anthracothere Microbunodon. Due to its high significance in the reorganisation of European, hoofed-mammal communities, we propose to name it the Microbunodon Event. This first step was followed by a phase of extinctions (MP29-30) and later by a phase of regional speciation and diversification (MN1-2). The Oligocene–Miocene faunal transition ended right before the two-phased turnover linked to the Proboscidean Datum Events (MN3-4). Locomotion types of rhinocerotids and ruminants provide new data on the evolution of environments during the Oligocene–Miocene transition and help understand the factors controlling these different phases. Indeed, it appears that the faunal turnovers were primarily directed by migrations, whereas the Agenian transitional phase mainly witnessed speciations.  相似文献   

11.
The first record of the superfamily Trigonioidoidea (Order: Unionoida) from the Lower Cretaceous of Teruel, Spain is described and compared to known taxa from Europe and Asia. This is a new record of the Trigonioidoidea from Europe, following material described from the Barremian of the Isle of Wight (southern England) and Aptian-Barremian from Iberian Range (Spain). The new material, recorded from the Albian of the Escucha Formation (“Barriada” Member) in Utrillas (Teruel, northeastern of Spain), comprises four specimens. Analysis of the main morphological characters including anterior musculature, dentition, and ornamentation indicates that this material constitutes a new genus and species, Iberanaia iberica. The palaeoenvironment in which the bivalves lived corresponds to a fluvial-palustral setting connected to a deltaic system.  相似文献   

12.
A substantial part of the European record of hadrosauroid ornithopod dinosaurs comes from the Maastrichtian (Upper Cretaceous) strata of the south-central Pyrenees (northern Catalunya and Aragón, northeastern Spain). Yet, our understanding on the taxonomy and phylogenetic systematics of these animals in this region is hindered by the lack of complete specimens, paucity of diagnostic material, and poor preservation of the fossils. The Basturs Poble site (Lleida Province, northern Catalunya) has yielded over 900 bones attributed to a hadrosauroid monodominant population. The dentary is the most common, and one of the better-preserved elements found in this locality. Here, we evaluate the utility of dentary morphology as an indicator of hadrosauroid taxonomic diversity. We do so via principal component analysis, cluster analysis, and mean comparison tests of various morphometric data from the available sample of dentaries collected so far in the Basturs Poble locality and other Maastrichtian sites in northeastern Spain. Three different dentary morphotypes are recognized: two different hadrosauroids, one more derived than the other, and one lambeosaurine type. One of the hadrosauroid morphotypes corresponds to relatively small individuals that may represent insular dwarf species. Concerning Basturs Poble locality, our results suggest the coexistence of hadrosauroid and lambeosaurine dinosaurs during the late Maastrichtian on the Ibero-Armorican Island.  相似文献   

13.
Analysis of geological sections, paleogeography, and paleomagnetic data is used to reveal succession of the middle to late Sarmatian mammal localities of the Eastern Paratethys and their correlation with the continental stratigraphic scale of Western Europe. Until recently, the late Sarmatian localities were correlated with MN10 and even MN11 zones. As is proved, all the known late Sarmatian localities should be correlated with the upper half of Zone MN9. The terminal late Sarmatian faunas only, which are correlative with the lowermost Chron C4Ar3r and older than 9.6 Ma, can be referred to Zone MN10. According to essential changes in taxonomic composition of faunas from continental localities around the Eastern Paratethys, which are recorded in the mid-late Sarmatian, Zone MN9 can be divided in two subzones MN9a and MN9b separated by boundary at ~10.5 Ma. The refined correlation can be used to establish difference between faunas of the Southeastern, Central, and West European paleozoogeographic provinces and to assess diachronism in dispersal of mammals. In the Southeastern province, many forms characteristic of the Turolian in Central and Western Europe first appeared as early as in the mid-Vallesian. The results obtained indicate that faunal criteria used to define boundaries of MN zones in Western Europe are of a regional importance being inapplicable to the entire North Eurasia and even to Europe as a whole that is unfortunately ignored by many paleontologists. Criteria of distinction should be worked out for each paleozoogeographic province. As geochronological levels of faunal changes are identical throughout the northern Palearctic, they suggest impact of global factors despite variable manifestation in different regions.  相似文献   

14.
The fossil record of small-sized theropods in south-western Europe is scarce and fragmentary. In absence of more direct evidence, the oological record arises as an alternative source of information to infer biodiversity. In this regard, the controversial dinosaur eggshells from the Les Labadous locality in southern France are herein re-evaluated in the light of new material and current parataxonomic information. New data and analyses reveal a new eggshell type, Montanoolithus labadousensis oosp. nov., characterized by a unique combination of characters. Additionally, the phylogenetic analysis reinforces the placement of the new oospecies as the sister taxon of the North American Montanoolithus strongorum, refusing the previous attribution of the Les Labadous remains to Elongatoolithidae. The combination of microstructural and phylogenetic results suggests that Montanoolithus labadousensis was likely produced by a dromaeosaurine dromaeosaurid, a taxonomic attribution that is consistent with the biodiversity of small-sized theropods at the latest Cretaceous of south-western Europe.  相似文献   

15.
16.
The Fichtelgebirge and Steinwald areas are situated at the western end of the Cenozoic Eger graben. Repeated Neogene fault activity and polyphase morphogenesis created a complex morphotectonic structure. Based on morphotectonic criteria three stages of tectonic activity can be determined: (a) late Oligocene to early Miocene; (b) middle-late Miocene; and (c) late Pliocene to early Pleistocene. Due to the Neogene tectonic movements, the pre-basaltic (preOligocene to Miocene) landscape has been differentially uplifted/downthrown and intensely modified by post-basaltic surface planation (HF). Broad paleovalleys (AT) of late Miocene to early Pliocene age and a system of Pliocene and Quaternary fluvial terraces and pediments (BT and ET) have been incised into the widespread HF relief elements. Due to repeated faulting and erosion, the stratigraphic record of Neogene sediments within the studied area is fragmentary.  相似文献   

17.
This paper presents one of the richest and most complete vertebrate faunas of the late Middle Miocene (~12 Ma) of Central Europe. Up to now, sixty-two vertebrate taxa, comprising all major groups (fishes, amphibians, reptiles, birds, mammals), have been recorded. Based on sedimentological and palaeobiological evidences, this Fossillagerstätte is assumed to originate from a floodplain paleosol formed on top of a braided river sequence. The fauna points to a highly structured, somewhat vegetated landscape with a wide array of habitats (e.g., fluvial channels, sporadically moist floodplains, short-lived ponds, savannah-like open areas and screes). It was preserved due to a rapid drowning and the switch to a freshwater lake environment. Palaeoclimatological data, derived from pedogenic features as well as from biota, indicate an overall semi-arid, subtropical climate with distinct seasonality (mean annual precipitation 486 ± 252 mm, mean annual temperature ~15°C). This underlines the late Middle/early Late Miocene dry-spell in Central Europe. From taphonomical point of view, the irregularly distributed but roughly associated larger vertebrate remains refer to an in situ accumulation of the bone bed. Splintered bones, gnawing marks as well as rhizoconcretions and root corrosion structures record some pre- and post-burial modification of the taphocoenose. However, the findings of pellet remains argue for a very fast burial and thus to a low degree of time-averaging. For this reason, the fossil fauna reflects the original vertebrate community rather well and is a cornerstone for the understanding of late Middle Miocene terrestrial ecosystems in this region. Certainly, Gratkorn will be one of the key faunas for a high-resolution continental biostratigraphy and the comprehension of Europe’s faunal interchanges near the Middle/Late Miocene transition.  相似文献   

18.
A new species, Tsuga nanfengensis sp. nov. (Pinaceae), is described on the basis of lignified fossil wood from the late Miocene of the Xianfeng Basin, central Yunnan, southwestern China. Detailed observation of the fossil wood specimens show the following characteristics: distinct growth rings, absence of resin canals, uniseriate bordered pits in the radial wall of tracheids, ray tracheids and piceoid and cupressoid cross-field pits. These features indicate similarities to the wood of extant Tsuga canadensis, T. chinensis, and T. dumosa. According to the fossil record, Tsuga was present in Xundian County during the Miocene. Today Tsuga is drought intolerant, preferring wet conditions with no extant species growing naturally in Xundian County. The presence of Tsuga in the Miocene of Xundian County indicates a humid climate consistent with previous palaeoclimatic reconstructions showing a wetter and probably shorter dry season in the Miocene, relative to the present day. Therefore, the change in the local climate such as increasing aridity through the Miocene might explain the local extinction of Tsuga from central Yunnan.  相似文献   

19.
20.
The continental climatic evolution of Anatolia has been reconstructed quantitatively for the last 45 million years using the coexistence approach. Although there were some regional effects, the Anatolian Cenozoic continental climate record correlated with the European climatic condition and the global oxygen isotope record from marine environments. From middle Eocene to late Miocene, continental warming in Anatolia was pronounced for inferred winter temperature and mean annual temperature as in Europe. Generally, the palaeoclimatic property of Anatolia resembles the European climatic changing and marine temperature changing based on the oxygen isotope record; however, climatic values of the terrestrial area in Anatolia are higher from Lutetian to Aquitanian and these values are lower than European values from Aquitanian to Tortonian. Correspondingly, Cenozoic climatic cooling in Anatolia is directly associated with an increase of seasonality, palaeogeographic position and terrestrial condition. Furthermore, mean annual precipitation values of Anatolia remained relatively stable during the Eocene–Oligocene; however, these values indicated changing throughout middle–late Miocene. Moreover, in this study, decline of abundance and variables for the mangrove and back mangrove palaeocommunities during the last 45 million years is recorded because of the decreasing of humidity, temperature and increasing of terrestrial condition.  相似文献   

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