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1.
A well resolved and highly accurate direct numerical simulation (DNS) solver has been developed to understand the implication of hydrodynamics to sediment transport. In the first part of the study we focus on steady flow over two-dimensional and three-dimensional ripples at two Reynolds numbers Reτ = 180 and 400 (defined by channel half-height and wall-friction velocity) in a channel geometry. The DNS scheme is based on a fourth-order vertical velocity and second-order vertical vorticity formulation, which resolves the difficulties in pressure boundary condition encountered when solving the Navier–Stokes equations. The complex boundary introduced due to the ripples has been imposed in the Cartesian domain using an elegant immersed boundary method. Detailed hydrodynamic analysis has revealed turbulence statistics (in particular, the higher order) and henceforth, the flow structures are sensitive — whether the ripples are two-dimensional or three-dimensional. The importance of fluctuating component of the bottom stress in addition to its mean component; and its significance to sediment transport and ripple migration speed have been investigated.  相似文献   

2.
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3.
A numerical model is developed to predict the onset of local scour below offshore pipelines in steady currents and waves. The scour is assumed to start when the pressure gradient underneath the pipeline exceeds the floatation gradient of the sediments. In this model, the water flow field above the bed is determined by solving the two-dimensional (2-D) Reynolds-averaged Navier–Stokes equations with a k-ω turbulence closure. The seepage flow below the seabed is calculated by solving the Darcy's law (Laplace's equation) with known pressure distribution along the common boundaries of the flow domains-seabed. The numerical method used for both the turbulent flow around the pipeline and Darcy's flow in the seabed is a fractional finite element method. The average pressure gradient along the buried pipe surface is employed in the evaluation of onset condition with a calibration coefficient. The numerical model is validated against experimental data available in literature. A unified onset condition for steady currents and waves is proposed. Influences of flow parameters, including water depth, embedment depth, boundary layer thickness, Reynolds number (Re) and Keuleagan–Carpenter (KC) number, on the pressure drop coefficient over the pipeline are studied systematically.  相似文献   

4.
This paper presents an investigation of the roughness effects in the turbulent boundary layer for asymmetric waves by using the baseline (BSL) k–ω model. This model is validated by a set of the experimental data with different wave non-linearity index, Ni (namely, Ni = 0.67, Ni = 0.60 and Ni = 0.58). It is further used to simulate asymmetric wave velocity flows with several values of the roughness parameter (am/ks) which increase gradually, namely from am/ks = 35 to am/ks = 963. The effect of the roughness tends to increase the turbulent kinetic energy and to decrease the mean velocity distribution in the inner boundary layer, whereas in the outer boundary layer, the roughness alters the turbulent kinetic energy and the mean velocity distribution is relatively unaffected. A new simple calculation method of bottom shear stress based on incorporating velocity and acceleration terms is proposed and applied into the calculation of the rate of bed-load transport induced by asymmetric waves. And further, the effect of bed roughness on the bottom shear stress and bed-load sediment transport under asymmetric waves is examined with the turbulent model, the newly proposed method, and the existing calculation method. It is found that the higher roughness elements increase the magnitude of bottom shear stress along a wave cycle and consequently, the potential net sediment transport rate. Moreover, the wave non-linearity also shows a big impact on the bottom shear stress and the net sediment transport.  相似文献   

5.
A boundary layer flow under spilling breakers in a laboratory surf zone with a smooth bottom is investigated using a high resolution particle image velocimetry (PIV) technique. By cross-correlating the images, oscillatory velocity profiles within a viscous boundary layer of O(1) mm in thickness are resolved over ten points. Using PIV measurements taken for an earlier study and the present study, flow properties in the wave bottom boundary layer (WBBL) over the laboratory surf zone are obtained, including the mean velocities, turbulence intensity, Reynolds stresses, and intermittency of coherent events. The data are then used to estimate the boundary layer thickness, phase variation, and bottom shear stress. It is found that while the time averaged mass transport inside the WBBL is onshore in the outer surf zone, it changes to offshore in the inner surf zone. The zero Eulerian mass transport occurs at h/hb ≈ 0.92 in the outer surf zone. The maximum overshoot of the streamwise velocity and boundary layer thickness are not constant across the surf zone. The bottom shear stress is mainly contributed by the viscous stress through mean velocity gradient while the Reynolds stress is small and negligible. The turbulence level is higher in the inner surf zone than that in the outer surf zone, although only a slight increase of turbulent intensity is observed inside the WBBL from the outer surf zone to the inner surf zone. The variation of phase inside and outside the WBBL was examined through the spatial velocity distribution. It is found the phase lead is not constant and its value is significantly smaller than previous thought. By analyzing instantaneous velocity and vorticity fields, a remarkable number of intermittent turbulent eddies are observed to penetrate into the WBBL in the inner surf zone. The size of the observed large eddies is about 0.11 to 0.16 times the local water depth. Its energy spectra follow the − 5/3 slope in the inertial subrange and decay exponentially in the dissipation subrange.  相似文献   

6.
The conditions for energy flux, momentum flux and the resulting streaming velocity are analysed for standing waves formed in front of a fully reflecting wall. The exchange of energy between the outer wave motion and the near bed oscillatory boundary layer is considered, determining the horizontal energy flux inside and outside the boundary layer. The momentum balance, the mean shear stress and the resulting time averaged streaming velocities are determined. For a laminar bed boundary layer the analysis of the wave drift gives results similar to the original work of Longuet–Higgins from 1953. The work is extended to turbulent bed boundary layers by application of a numerical model. The similarities and differences between laminar and turbulent flow conditions are discussed, and quantitative results for the magnitude of the mean shear stress and drift velocity are presented. Full two-dimensional simulations of standing waves have also been made by application of a general purpose Navier–Stokes solver. The results agree well with those obtained by the boundary layer analysis. Wave reflection from a plane sloping wall is also investigated by using the same numerical model and by physical laboratory experiments. The phase shift of the reflected wave train is compared with theoretical and empirical models.  相似文献   

7.
Sand transport in Lido and Chioggia inlets was measured using modified Helley–Smith sand traps equipped with 60-micron nets. The traps had an efficiency of about 4% only but provided enough material for analysis. Very fine sand (0.07 < d < 0.11 mm) only was collected in the traps. Transport of sand was greatest in the bottom 10% of the water column and followed a Rouse profile. Sand extended to a height of about 4 m above the bed during peak flows corresponding to the estimated thickness of the boundary layer; and observed in synoptic ADCP profiles. The sand in the benthic boundary layer was largely inorganic (>95%); above this layer, organic content varied widely and was greatest near the surface. The movability number Ws/U∗Ws/U∗ showed a linear relationship to dimensionless grain diameter (D*): (Ws/U∗)=(D∗/10)(Ws/U∗)=(D∗/10); D* < 10. Sand concentration in suspension was simulated by a mean Rouse parameter of −2.01 ± 0.66 (Lido inlet) and −0.82 ± 0.27 (Chioggia inlet). The β parameter ( Hill et al., 1988) was correlated with D* and movability number in the form: β=2.07−2.03D∗+59(Ws/U∗)2β=2.07−2.03D∗+59(Ws/U∗)2 (r2 = 0.42). Von Karman's constant was back-calculated from a Law of the Wall relationship as a test on the accuracy of U* estimates; a mean value of 0.37 ± 0.1 (compared to the accepted value of 0.41) suggest U* was accurate to within 10%. The constant of proportionality (γ = 3.54 × 10−4) between reference concentration (Ca) and normalized excess bed shear stress was in line with the published literature.  相似文献   

8.
The hydrodynamic properties and the capability to measure sediment-water solute fluxes, at assumed steady state conditions, were compared for three radically different benthic chamber designs: the “Microcosm”, the “Mississippi” and the “Göteborg” chambers. The hydrodynamic properties were characterized by mounting a PVC bottom in each chamber and measuring mixing time, diffusive boundary layer thickness (DBL thickness) shear velocity (u∗), and total pressure created by the water mixing. The Microcosm had the most even distribution of DBL thickness and u∗, but the highest differential pressure at high water mixing rates. The Mississippi chamber had low differential pressures at high u∗. The Göteborg chamber was in between the two others regarding these properties. DBL thickness and u∗ were found to correlate according to the following empirical formula: DBL=76.18(u∗)−0.933. Multiple flux incubations with replicates of each of the chamber types were carried out on homogenized, macrofauna-free sediments in four tanks. The degree of homogeneity was determined by calculating solute fluxes (of oxygen, silicate, phosphate and ammonium) from porewater profiles and by sampling for porosity, organic carbon and meiofauna. All these results, except meiofauna, indicated that there were no significant horizontal variations within the sediment in any of the parallel incubation experiments. The statistical evaluations also suggested that the occasional variations in meiofauna abundance did not have any influence on the measured solute fluxes. Forty-three microelectrode profiles of oxygen in the DBL and porewater were evaluated with four different procedures to calculate diffusive fluxes. The procedure presented by Berg, Risgaard-Petersen and Rysgaard, 1989 [Limnol. Oceanogr. 43, 1500] was found to be superior because of its ability to fit measured profiles accurately, and because it takes into consideration vertical zonation with different oxygen consumption rates in the sediment. During the flux incubations, the mixing in the chambers was replicated ranging from slow mixing to just noticeable sediment resuspension. In the “hydrodynamic characterizations” these mixing rates corresponded to average DBL thickness from 120 to 550 μm, to u∗ from 0.12 to 0.68 cm/s, and to differential pressures from 0-3 Pa. Although not directly transferable, since the incubations were done on a “real” sediment with a rougher surface while in the characterizations a PVC plate simulated the sediments surface, these data give ideas about the prevailing hydrodynamic condition in the chambers during the incubations. The variations in water mixing did not generate statistically significant differences between the chamber types for any of the measured fluxes of oxygen or nutrients. Consequently it can be concluded that, for these non-permeable sediments and so long as appropriate water mixing (within the ranges given above) is maintained, the type of stirring mechanism and chamber design used were not critical for the magnitude of the measured fluxes. The average measured oxygen flux was 11.2 ± 2.7 (from 40 incubations), while the diffusive flux calculated (from 43 profiles using the Berg et al., 1989 [Limnol. Oceanogr. 43, 1500] procedure) was 11.1 ± 3.0 mmol m−2 day−1. This strongly suggests that accurate oxygen flux measurements were obtained with the three types of benthic chambers used and that the oxygen uptake is diffusive.  相似文献   

9.
High Reynolds number flows around a circular cylinder close to a flat seabed have been computed using a two-dimensional standard high Reynolds number k–ε turbulence model. The effects of gap to diameter ratio, Reynolds number and flat seabed roughness for a given boundary layer thickness of the inlet flow upstream of the cylinder have been investigated. Hydrodynamic quantities and the resulting bedload transport have been predicted, and the vortex shedding mechanisms have been investigated. Predictions of hydrodynamic quantities around a cylinder located far away from the bed (so that the effect of the bed is negligible) are in satisfactory agreement with published experimental data and numerical results obtained for the flow around an isolated cylinder. Results for lower Reynolds number flows have also been computed for comparison with the high Reynolds number flow results. Overall it appears that the present approach is suitable for design purposes at high Reynolds numbers which are present near the seabed in the real ocean.  相似文献   

10.
Turbidity limits gas exchange in a large macrotidal estuary   总被引：1，自引：0，他引：1  
In estuaries, the gas transfer velocity (k) is driven by a combination of two major physical drivers, wind and water current. The k values for CO2 in the macrotidal Gironde Estuary were obtained from 159 simultaneous pCO2 and floating chamber flux measurements. Values of k increased with wind speed and were significantly greater when water currents and wind were in opposing directions. At low wind speeds (<1 m s−1), k increased with water current velocities (0–1.5 m s−1) following an exponential trend. The latter was a good proxy for the Y-intercept in a generic equation for k versus wind speed in estuaries. We also found that, in this turbid estuary, k was significantly lower at high turbidity. The presence of suspended material in great concentrations (TSS > 0.2 g L−1) had a significant role in attenuating turbulence and therefore gas exchange. This result has important consequences for modeling water oxygenation in estuarine turbidity maxima. For seven low turbidity estuaries previously described in the literature, the slope of the linear regression between k and wind speed correlates very well with the estuary surface area due to a fetch effect. In the Gironde Estuary, this slope follows the same trend at low turbidity (TSS < 0.2 g L−1), but is on average significantly lower than in other large estuaries and decreases linearly with the TSS concentration. A new generic equation for estuaries is proposed that gives k as a function of water current velocity, wind speed, estuarine surface area and TSS concentration.  相似文献   

11.
Turbulent flow fields under spilling breaking waves are measured by particle image velocimetry and analyzed using the wavelet techniques in a laboratory surf zone. The turbulent vortical structures and corresponding length scales in the flow are detected through the eduction of the most excited mode with local intermittency measure that is found to correlate with the passage of the structure. Distributions and evolution of the educed vortical structures are presented and discussed. Packets of vortical structures with high intermittency is observed to stretch downward below the initially low-intermittency trough level, indicating these structures play a crucial role in turbulent mixing below the trough level. It is found that the probability density functions of the intermittent energy of the educed structures, vorticity and swirl strength display an exponential decay. Ensemble-averaged length scales of the educed vortical structures are found to be about 0.1 to 0.2 times the local water depth, close to the turbulent mixing length reported in the surf zone. The Kolmogorov microscale is evaluated and the turbulent mixing length is estimated using the k − ε relation and mixing length hypothesis. The k − ε relation may overestimate the mixing length scale for energetic descending eddies.  相似文献   

12.
A simple relationship has been developed between the wall coordinate y+ and Kolmogorov's length scale using direct numerical simulation (DNS) data for a steady boundary layer. This relationship is then utilized to modify two popular versions of low Reynolds number k–ε model. The modified models are used to analyse a transitional oscillatory boundary layer. A detailed comparison has been made by virtue of velocity profile, turbulent kinetic energy, Reynolds stress and wall shear stress with the available DNS data. It is observed that the low Reynolds number models used in the present study can predict the boundary layer properties in an excellent manner.  相似文献   

13.
Picoplankton distribution at the boundary zone of the southern Adriatic in May 2009 on a 75 km long shelf-continental slope transect was assessed by combining epifluorescence microscopy, flow cytometry and high-performance liquid chromatography data with hydrographic observations. The picoplankton distribution was greatly influenced by the hydrographic conditions prevailing in the southern Adriatic because of the influence of the Levantine Intermediate Water (LIW) and East Adriatic Current (EAC) forcing. Heterotrophic bacteria numerically dominated the picoplankton community through the entire transect with no significant accumulation. By contrast, picophytoplankton accumulated in the 50–75 m layer, forming a pronounced deep chlorophyll maximum. Synechococcus dominated the photosynthetic picoplankton, whereas picoeukaryotes were the least abundant. The intrusion of warm LIW observed in the layer between 100 and 350 m was followed by Prochlorococcus and Synechococcus peaks (10 × 103 cells mL−1 and 90 × 103 cells mL−1, respectively), as well as by the appearance of two Synechococcus ecotypes. Most picoeukaryotes were observed at the offshore stations, where geostrophic current calculation revealed the strongest EAC influence. A strong EAC spread over the central and eastern basin created a barrier for Prochlorococcus, whereas the picoeukaryote maxima coincided with the core of the EAC, suggesting its persistence to hydrological instabilities.  相似文献   

14.
Large eddy simulations of the flow around a circular cylinder at high Reynolds numbers are reported. Five Reynolds numbers were chosen, such that the drag crisis was captured. A total of 18 cases were computed to investigate the effect of gridding strategy, turbulence modelling, numerical schemes and domain width on the results. It was found that unstructured grids provide better resolution of key flow features, when a ‘reasonable’ grid size is to be maintained.When using coarse grids for large eddy simulation, the effect of turbulence models and numerical schemes becomes more pronounced. The dynamic mixed Smagorinsky model was found to be superior to the Smagorinsky model, since the model coefficient is allowed to dynamically adjust based on the local flow and grid size. A blended upwind-central convection scheme was also found to provide the best accuracy, since a fully central scheme exhibits artificial wiggles, due to dispersion errors, which pollute the solution.Mean drag, fluctuating lift Strouhal number and base pressure are compared to experiments and empirical estimates for Reynolds numbers ranging from 6.31 × 104 to 5.06 × 105. In terms of the drag coefficient, the drag crisis is well captured by the present simulations, although the other integral quantities (rms lift and Strouhal number) show larger discrepancies. For the lowest Reynolds number, the drag is seen to be more sensitive to the domain width than the spanwise grid spacing, while at the higher Reynolds numbers the grid resolution plays a more important role, due to the larger extent of the turbulent boundary layer.  相似文献   

15.
Phytoplankton composition and biomass across the southern Indian Ocean   总被引：2，自引：0，他引：2  
Phytoplankton composition and biomass was investigated across the southern Indian Ocean. Phytoplankton composition was determined from pigment analysis with subsequent calculations of group contributions to total chlorophyll a (Chl a) using CHEMTAX and, in addition, by examination in the microscope. The different plankton communities detected reflected the different water masses along a transect from Cape Town, South Africa, to Broome, Australia. The first station was influenced by the Agulhas Current with a very deep mixed surface layer. Based on pigment analysis this station was dominated by haptophytes, pelagophytes, cyanobacteria, and prasinophytes. Sub-Antarctic waters of the Southern Ocean were encountered at the next station, where new nutrients were intruded to the surface layer and the total Chl a concentration reached high concentrations of 1.7 ??g Chl a L−1 with increased proportions of diatoms and dinoflagellates. The third station was also influenced by Southern Ocean waters, but located in a transition area on the boundary to subtropical water. Prochlorophytes appeared in the samples and Chl a was low, i.e., 0.3 ??g L−1 in the surface with prevalence of haptophytes, pelagophytes, and cyanobacteria. The next two stations were located in the subtropical gyre with little mixing and general oligotrophic conditions where prochlorophytes, haptophytes and pelagophytes dominated. The last two stations were located in tropical waters influenced by down-welling of the Leeuwin Current and particularly prochlorophytes dominated at these two stations, but also pelagophytes, haptophytes and cyanobacteria were abundant. Haptophytes Type 6 (sensuZapata et al., 2004), most likely Emiliania huxleyi, and pelagophytes were the dominating eucaryotes in the southern Indian Ocean. Prochlorophytes dominated in the subtrophic and oligotrophic eastern Indian Ocean where Chl a was low, i.e., 0.043-0.086 ??g total Chl a L−1 in the surface, and up to 0.4 ??g Chl a L−1 at deep Chl a maximum. From the pigment analyses it was found that the dinoflagellates of unknown trophy enumerated in the microscope at the oligotrophic stations were possibly heterotrophic or mixotrophic. Presence of zeaxanthin containing heterotrophic bacteria may have increased the abundance of cyanobacteria determined by CHEMTAX.  相似文献   

16.
Steady streaming around a circular cylinder in an oscillatory flow   总被引：1，自引：0，他引：1  
Steady streaming induced by an oscillatory flow around a circular cylinder is investigated using a numerical method. Two-dimensional Reynolds-averaged Navier-Stokes equations are solved using a finite element method with a k-ω turbulent model closure. The range of the Keulegan-Carpenter (KC) number investigated is between 2 and 40, which is substantially higher than those reported in literature related to steady streaming to date. A constant value of Stokes number (β) of 196 is chosen in this study. The steady streaming structures and velocity distribution are analysed in detail. It is found that the characteristics of steady streaming are strongly related to the vortex shedding flow regimes.  相似文献   

17.
The chlorophyll a specific absorption coefficient of phytoplankton, aφ∗(λ) is an important parameter to determine for primary production models and for the estimation of phytoplankton physiological condition. Knowledge of this parameter at high latitudes where nutrient rich cold water submitted to low incident light is a common environment is almost nonexistent. To address this issue, we investigated the light absorption properties of phytoplankton as a function of irradiance, temperature, and nutrients using a large data set in the southern Beaufort Sea during the open water to ice cover transition period. The aφ∗(λ) tended to increase from autumn when open water still existed to early winter when sea ice cover was formed, resulting from a biological selection of smaller-size phytoplankton more efficient to absorb light. There was no significant correlation between aφ∗(λ) and irradiance or temperature for both seasons. However, aφ∗(λ) showed a significant positive correlation with NO3 + NO2. Implications of the results for phytoplankton community adaptation to changing light levels are discussed.  相似文献   

18.
Wave boundary layer over a stone-covered bed   总被引：1，自引：0，他引：1  
This paper summarizes the results of an experimental investigation on wave boundary layers over a bed with large roughness, simulating stone/rock/armour block cover on the sea bottom. The roughness elements used in the experiments were stones the size of 1.4cm and 3.85cm in one group of experiments and regular ping-pong balls the size 3.6cm in the other. The orbital-motion-amplitude-to-roughness ratio at the bed was rather small, in the range a/ks = 0.6–3. The mean and turbulence properties of the boundary-layer flow were measured. Various configurations of the roughness elements were used in the ping-pong ball experiments to study the influence of packing pattern, packing density, number of layers and surface roughness of the roughness elements. The results show that the friction factor seems to be not extremely sensitive to these factors. The results also show that the friction factor for small values of the parameter a/ks does not seem to tend to a constant value as a/ks → 0 (contrary to the suggestion made by some previous investigators). The present friction-factor data indicates that the friction factor constantly increases with decreasing a/ks. An empirical expression is given for the friction factor for small values of a/ks. The results further show that the phase lead of the bed friction velocity over the surface elevation does not seem to change radically with a/ks, and found to be in the range 12°–23°. Furthermore the results show that the boundary-layer turbulence also is not extremely sensitive to the packing pattern, the packing density, the number of layers and the surface roughness of the roughness elements. There exists a steady streaming near the bed in the direction of wave propagation, in agreement with the existing work. The present data indicate that the steady streaming is markedly smaller in the case where the ping-pong balls are aligned at 45° to the wave direction than in the case with 90° alignment. Likewise, it is found that the steady streaming is relatively smaller in the case of the one-layer ping-pong-ball roughness than in the case of the two-layer situation.  相似文献   

19.
The annual total and organic mercury bioaccumulation pattern of Scrobicularia plana and Hediste diversicolor was assessed to evaluate the potential mercury transfer from contaminated sediments to estuarine food webs. S. plana was found to accumulate more total and organic mercury than H. diversicolor, up to 0.79 mg kg−1 and 0.15 mg kg−1 (wet weight) respectively, with a maximum annual uptake of 0.21 mg kg−1 y−1, while for methylmercury the annual accumulation was similar between species and never exceeded 0.045 mg kg−1 y−1. The higher organic mercury fraction in H. diversicolor is related to the omnivorous diet of this species. Both species increase methylmercury exposure by burrowing activities and uptake in anoxic, methylmercury rich sediment layers. Integration with the annual biological production of each species revealed mercury incorporation rates that reached 28 μg m−2 y−1, and to extract as much as 11.5 g Hg y−1 (of which 95% associated with S. plana) in the 0.4 km2 of the most contaminated area, that can be transferred to higher trophic levels. S. plana is therefore an essential vector in the mercury biomagnification processes, through uptake from contaminated sediments and, by predation, to transfer it to economically important and exploited estuarine species.  相似文献   

20.
The role of copepod grazing on the ecosystem dynamics in the Oyashio region, western subarctic Pacific was investigated during six cruises from June 2001 to June 2002. In situ grazing rates of the copepod community (CGR) were measured by the gut fluorescence method in respect to developmental stages of dominant species. In terms of biomass, more than 80% of the copepod community was dominated by six large calanoid species (Neocalanus cristatus, Neocalanus flemingeri, Neocalanus plumchrus, Eucalanus bungii, Metridia pacifica and Metridia okhotensis) throughout the year. Resulting from the observed pattern of the interzonal migrating copepods, the CGR in the Oyashio region was divided into three phases, i.e. spring (bloom), summer (post-bloom) and autumn-winter phase. During the spring bloom, late copepodites of the interzonal migrating species, N. cristatus, N. flemingeri and E. bungii appeared in the surface layer (0-50 m) to consume the production of the bloom, resulting in a high grazing rate of the copepod community (7.9 mg Chl m−2 d−1), though its impact on phytoplankton community was low due to the high primary productivity. During the post-bloom period, although the copepod community which was dominated by N. cristatus, N. plumchrus, M. pacifica and newly recruited E. bungii still maintained a high biomass, the CGR was generally lower (1.8-2.6 mg Chl m−2 d−1 for June and August 2001), probably due to the lower availability of phytoplankton. Nevertheless, the highest CGR was also observed during this period (10.5 mg Chl m−2 d−1 in June 2002). The high CGR on autotrophic carbon accounted for 69% of the primary production, suggesting that the copepod community in the Oyashio region potentially terminates the phytoplankton bloom. Abundant occurrence of young E. bungii, which is a characteristic phenomenon in the Oyashio region, was largely responsible for the high grazing pressure in June 2002 suggesting that success of reproduction, growth, and survival in E. bungii during the spring bloom is an important factor in controlling phytoplankton abundance during the post-bloom season. During autumn and winter, CGR was the lowest in the year (0.29-0.38 mg Chl. m−2 d−1) due to the disappearance of the interzonal migrating copepods from the surface layer. Diel migrant M. pacifica was the most important grazer during this period. The annual ingestion of the copepod community is estimated as 37.7 gC m−2 on autotrophic carbon (converted using C:Chl ratio of 30) or 137.9 gC m−2 on suspended particles (using C:Chl ratio of in situ value, 58-191), accounting for 13% and 46% of annual primary production, respectively. This study confirms that copepod grazing is an important pathway in carbon flow in the Oyashio region and in particular their role in the phytoplankton dynamics is significant for the termination of the spring bloom.  相似文献   

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